ケアシハイイロタケ

Mycena lanuginosa Har. Takah., Mycoscience (2007) 48: 342-357.　

最初暗青紫褐色を帯び、のち次第に灰色を呈するクヌギタケ属の仲間で、しばしば空中伸長型の菌糸束を伴います。

  肉眼的特徴: 傘は径 7-11 mm, 釣り鐘形で平開せず, 明瞭な放射状の溝線を表 し, やや吸

水性, 湿時半透明の条線を表し, 乾性, 平滑, 最初白色粉状, まもなく平滑になり, 中央部は灰

褐色～紫褐色, 周縁部に向かって退色する.　肉は傘の中央部において厚さ 0.5 mm 以下, 白

色, 特別な味や臭いはない. 柄は 30-60×0.8-1.3 mm, 円柱形, 中心生, やせ型, 中空, 乾性, 

上部は白色粉状, 下部は長毛に被われ, 傘よりやや淡色, 根本は白色の剛毛に被われる. ヒ

ダは上生し, 疎 (柄に到達するヒダは 12-18), 幅 1.5 mm以下, 類白色, 縁取りを欠く. 

  顕微鏡的特徴: 担子胞子は 10-12×5.5-6.5 μm, 楕円形, 表面は平坦, 無色, アミロイド, 薄

壁. 担子器は 35-42×7-9 μm, こん棒形, 4胞子性. 縁シスチジアは 40-80×5-15 μm, 群生

し, 紡錘形, しばしば頂部が微突形になり, 無色, 平坦, 薄壁. 側シスチジア .63-102×8-15 μ

m, 紡錘形, 無色, 平坦, 薄壁. 子実層托実質の菌糸は偽アミロイド. 傘の上表皮層の菌糸は匍

匐性で, 幅 2-6μm, 平行に配列し, 円柱形, 疣状～短指状突起に被われ, 帯褐色, 薄壁. 上表

皮下層の菌糸は無色または帯褐色, 実質の大部分は偽柔組織状 [pseudoparenchymatous: 

著しく肥大し, 複数の隔壁を持った短形の菌糸細胞] で, 菌糸は平列 し，偽アミロイド. 柄表皮

の菌糸は平坦または疣状～短指状突起に被われ, 無色または帯褐色. 傘と柄の表皮組織に

クランプが存在する.

  供試標本: KPM-NC0008702 (holotype), Izumino-mori, Yamato-shi, Kanagawa pref., 27 

Apr. 2000, coll. H.Takahashi; KPM-NC0008729, the same place, 25 Nov. 1999 coll. H. 

Takahashi; the same place, 16 Mar.2000, coll. H. Takahashi; the same place, 27 Nov. 1999, 

coll. H.Takahashi; Mt.Takao, Hachioji, Tokyo, 30 Oct. 1999, coll. H. Takahashi.  

  コメント: 傘は放射状の溝線を表し, 灰褐色～紫褐色で, 柄は下半部を中心に長毛に被わ

れ, 縁- 及び側シスチジアは平坦な紡錘形, 傘と柄の表皮組織はクランプを有する多分枝菌

糸からなる.  

  春から晩秋の頃, 神奈川県大和市泉の森及び東京都高尾山の広葉樹林内の落ち葉また

は落枝上に発生. 標本は神奈川県立生命の星・地球博物館 (KPM) の標本庫に登録, 収蔵さ

れている.

  本種は Maasの分類概念 (Maas 1980, 1988a) によるアクニオイタケ節 (section 

Fragilipedes) に属し, 節内において本種はオランダから報告された Mycena pilosella Maas 

Geest. (Courtecuisse and Duhem 1994; Maas 1988b; Robich 2003)及びヨーロッパに広く分布

する Mycena zephirus (Fr.: Fr.) P. Kumm. (Courtecuisse and Duhem 1994; Dahncke 1993; 

Kuhner 1938; Lisiewska 1987; Maas 1988c; Moser and Julich 1999; Robich 2003)に近縁と思

われるが,  Mycena pilosella は直角に立ち上がった長い円柱形の柄シスチジアを持ち, 側シス

チジアを欠く点で異なり, またMycena zephirus は類白色の傘, 平滑な柄, 大根臭, 長楕円形

の担子胞子, 頂部が分岐した縁シスチジアを有する.

Mycena lanuginosa Har. Takah., Mycoscience (2007) 48: 342-357.   

　 Etymology: From Latin, lanuginosa = lanugineous, referring to the pilose stipe. 

  Pileus 7-11 mm in diam, conico-convex to campanulate, distinctly radially sulcate-striate 

almost to the disc, subhygrophanous, dry, minutely white pruinose at first, soon becoming 

glabrous, dark brown (8F5-6 to 9F5-7) at the center, reddish brown (8E4-7 to 9E4-7) 

farther outward, much paler again to almost whitish at the margin . Flesh up to 0.5 mm, 

white; odor and taste not distinctive. Stipe 30-60×0.8-1.3 mm, cylindrical, central, slender, 

terete, hollow, dry, white pruinose above, pilose toward the base, greyish brown (8E3) to 

reddish brown (8E4) at the apex, reddish brown (8E4-7) downward; base covered with long, 

fairly coarse, whitish fibrils. Lamellae adnexed, distant (12-18 reach the stipe), up to 1.5 mm 

broad, thin, whitish; edges concolorous. 

  Basidiospores (n = 69 spores of 6 basidiocarps) 10-12×5.5-6.5 μm, Q (length/breadth) 

= 1.8, short ellipsoid to ellipsoid, smooth, colorless, amyloid, thin-walled. Basidia 35-42×7-9 

μm, clavate, four-spored, with basal clamps. Basidioles clavate. Cheilocystidia 40-80×5-15 

μm, abundant, forming a sterile lamella edge, fusiform, sometimes mucronate, smooth, 

colorless, thin-walled. Pleurocystidia 63-102×8-15 μm, abundant, fusiform, smooth, 

colorless, thin-walled. Hymenophoral trama regular; element hyphae 3-25 μm wide, 

cylindrical, often inflated, walls thin, smooth, hyaline, dextrinoid. Pileipellis a cutis of parallel, 

repent hyphae 2-6 μm wide, cylindrical, smooth or covered with scattered, warty or finger-

like diverticulae, walls thin, brownish; underlying hyphae parallel, hyaline or brownish, 

dextrinoid, with short and inflated cells up to 35 μm wide. Stipitipellis a cutis of parallel, 

repent hyphae 3-6 μm wide, cylindrical, smooth or covered with scattered, warty or finger-

like diverticulae, colorless or brownish, inamyloid or dextrinoid, thin-walled; terminal cells 

(caulocystidia) 5-9 μm wide, cylindrical, smooth or covered with scattered, warty or finger-

like diverticulae, colorless, thin-walled. Stipe trama composed of longitudinally running, 

cylindrical hyphae 6-20 μm wide, smooth, hyaline, dextrinoid. Clamp connections present in 

the cortical layer of pileus and stipe, and at the basal septa of basidia. 

  Known distribution: Japan (Kanagawa, Tokyo).

　 Habitat: Solitary or scattered, on dead leaves and twigs in lowland forests dominated by 

Quercus myrsinaefolia Blume and Q. serrata Thunb.ex Murray., from March to November. 

  Specimens examined: KPM-NC0008702 (holotype), Izumino-mori, Yamato-shi, Kanagawa 

pref., 27 Apr. 2000, coll. H.Takahashi; KPM-NC0008729, the same place, 25 Nov. 1999 coll. H. 

Takahashi; the same place, 16 Mar.2000, coll. H. Takahashi; the same place, 27 Nov. 1999, 

coll. H.Takahashi; Mt.Takao, Hachioji, Tokyo, 30 Oct. 1999, coll. H. Takahashi. 

  Japanese name: Keashi-haiirotake.

　 Notes: This species is distinct in its sulcate-striate, grayish brown to violet brown pileus, 

the pilose stipe, the amyloid basidiospores, the smooth, fusiform cheilocystidia and 

pleurocystidia, and the diverticulate elements in the cortical layer of pileus and stipe. These 

characteristics suggest placement of this taxon in the section Fragilipedes (Fr.) Quel., as 

defined by Maas Geesteranus (Maas Geesteranus 1980, 1988a). Within the section, M.

lanuginosa seems to be closely allied with M. pilosella Maas Geest., originally described from 

Netherlands (Maas Geesteranus 1988b; Courtecuisse and Duhem 1994; Robich 2003), and 

European M. zephirus (Weinm.) Quel. (Kuhner 1938; Lisiewska 1987; Maas Geesteranus 

1988c; Dahncke 1993; Courtecuisse and Duhem 1994; Moser and Julich 1999; Robich 2003). 

Mycena pilosella differs in having densely diverticulate elements of the pileipellis, long, 

cylindrical caulocystidia which diverge at right angle, and lacking pleurocystidia. Mycena

zephirus is distinct in forming a whitish pileus, a glabrescent stipe, raphanoid odor, ellipsoid-

cylindrical basidiospores, and apically branched cheilocystidia.  

References

Breitenbach J, Kranzlin F (1991) Fungi of Switzerland 3. Boletes and agarics, 1st part. 

  Edition Mycologia, Lucerne

Corner EJH (1994) Agarics in Malesia II. Mycenoid. Beih Nova Hedwigia109:165-271

Courtecuisse R, Duhem B (1994) Les Champignons de France. Eclectis,Paris

Dahncke RM (1993) 1200 Pilze in Farbfotos. AT Verlag, Stuttgart

Kornerup A, Wanscher JH (1978) Methuen handbook of colour, 3rd edn. Methuen, London

Kuhner R (1938) Le genre Mycena. Paul Lechevalier, Paris

Lisiewska M (1987) Flora Polska, Tom XVII. Polska Akademia Nauk, Warszawa

Maas Geesteranus RA (1980) Studies in Mycenas 15. A tentative subdivision of the genus 

  Mycena in the northern Hemisphere. Persoonia 11:93.120

Maas Geesteranus RA (1981) Studies in Mycenas 26. The Mycenas described by P.A. 

  Karsten. Proc K Ned Akad Wet C 84:221.231

Maas Geesteranus RA (1984) Conspectus of the Mycenas of the Northern Hemisphere 3. 

  Section Filipedes. Proc K Ned Akad Wet C 87:413.447

Maas Geesteranus RA (1985) Conspectus of the Mycenas of the Northern Hemisphere 4. 

  Section Mycena. Proc K Ned Akad Wet C 88:339.369

Maas Geesteranus RA (1988a) Conspectus of the Mycenas of the Northern Hemisphere 9.   

  Section Fragilipedes, species A.G. Proc K Ned Akad Wet C 91:43.83

Maas Geesteranus RA (1988b) Conspectus of the Mycenas of the Northern Hemisphere 9. 

  Section Fragilipedes, species I.R. Proc K Ned Akad Wet C 91:129.159

Maas Geesteranus RA (1988c) Conspectus of the Mycenas of the Northern Hemisphere 9. 

  Section Fragilipedes, species S.Z. Proc K Ned Akad Wet C 91:283.314

Maas Geesteranus RA (1989) Conspectus of the Mycenas of the Northern Hemisphere 13. 

  Section Calamophilae and Calodontes. Proc K Ned Akad Wet C 92:477.504

Maas Geesteranus RA (1990) Conspectus of the Mycenas of the Northern Hemisphere 14. 

  Section Adonideae, Aciculae, and Oregonenses. Proc K Ned Akad Wet C 93:163.186

Maas Geesteranus RA (1991) Conspectus of the Mycenas of the Northern Hemisphere 15. 

  Sections Hiemales and Exornatae. Proc K Ned Akad Wet. 94:81.102

Maas Geesteranus RA, de Meijer AAR (1997) Mycenae Paranaenses. Royal Netherlands 

  Academy of Arts and Sciences, Amsterdam

Moser M, Julich W (1993) Colour atlas of Basidiomycetes, Lief. Mycena 18. Fischer Verlag, 

  Berlin

Robich G (2003) Mycena d’Europa. AMB, Trento

Smith AH (1947) North American species of Mycena. Univ Mich Stud Sci Ser 17:1.521