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Lactarius glutininitens Har.Takah., Mycoscience 42: 355-360, 2001　

 傘の表面は著しい粘液に被われ、 乳液は半透明で変色性はありません。

 肉眼的特徴: 傘は径 30-80 mm, 最初饅頭形で縁部は内側に巻き, のち平開し, 中央部がく

ぼんでやや漏斗型になり, 平坦, 平滑, 同心円状の環紋を欠き, 表面は粘液に被われ, 最初淡

灰褐色のち淡灰色. 肉の厚さは 7 mm以下, 汚白色～淡クリ－ム色, 特別な臭いはなく, 後

口が僅かに辛い. 乳液は半透明で変色性を欠く. 柄は 30-70×5-16 mm mm, ほぼ上下同大, 

中心生, 時にやや偏圧され, 中空, 平滑, 乾性, 類白色～汚白色または淡灰色, 根本は根状

菌糸束に接続する. ヒダは直性～やや垂生し, 柄に到達するヒダは 36-42, 幅 6 mm以下, 淡

クリ－ム色; 縁部は平坦, 縁取りを欠く. 胞子紋はクリ－ム色.

  顕微鏡的特徴: 担子胞子は 8.5-11×7-8.5μm [Q = length/breadth: 1.21-1.30], 類球形～

広楕円形, 表面は高さ 2 mm の孤立した突起と帯状の隆起がある. 担子器は 20-30×6-8μ

m, こん棒形, 2-4 胞子性. ヒダ縁部の*マクロシスチジアは 22-30×5-7.5μm, 群生し, 類こん

棒形～類紡錘形, 頂部は尖り, 時に微突形, 平坦, 無色, 薄壁. ヒダ側面のマクロシスチジアは

35-90×7-10 μm, 子実層から突出し, 槍先形または類こん棒形～類紡錘形, 頂部は尖り, 時

に微突形, 平坦, 無色, 薄壁. 子実層托実質は平列型, 菌糸は傘実質に似るが, 無色. 傘表皮

の菌糸は上向性, 幅 5-10 μm, 円柱形, 無色, 粘質物に埋もれる.  傘と柄の実質は異質な細

胞 (球形細胞と糸状菌糸) が混在した構造 (Heteromerous Trama) をなし, 無色.  柄シスチジ

アはない. 全ての組織において菌糸はクランプを欠く.

  供試標本: KPM-NC0007430 (基準標本), Izumino-mori, Yamato-shi, Kanagawa-ken, 13 

Oct. 2000, H.Takahashi; KPM-NC0007431, the same place, 14 Oct. 2000, H.Takahashi.

  コメント: 傘の表面は淡灰色で著しい粘性を表し, 乳液は半透明で変色性はない. 神奈川県

大和市泉の森のシラカシとコナラを中心とする林内地上に発生. 標本は神奈川県立生命の

星・地球博物館の標本庫(KPM)に登録, 収蔵されている.

  * マクロシスチジア (大のう体, Macrocystidia): 排泄機能を有する偽シスチジアの一型で,

スルフォバニリンにより濃青色に, またはスルフォベンズアルデヒドにより黒色に染まり, 帯状～

粒状の内容物を持つ. ハラタケ目ではベニタケ科のみに存在する.

Lactarius glutininitens Har.Takah., Mycoscience 42: 355-360, 2001　　 　Figs. 1, 2

  Pileus 30-80 mm in diam, at first convex with involute margin, expanding to plane with 

depressed center, smooth, glabrous, azonate, glutinous, pale grayish brown when young, then 

pale grayish. Flesh up to 7 mm, beige to pale cream; odor none, taste after a while slightly 

acrid. Latex watery, unchanging. Stipe 30-70×5-16 mm, subequal, central, terete or 

compressed, hollow, glabrous, dry, whitish to beige or pale grayish; base with a few mycelial 

cords. Lamellae adnate to subdecurrent, 36-42 reach the stipe, up to 6 mm broad, pale 

cream; edges even, concolorous. 

  Spore print cream. Basidiospores 8.5-11×7-8.5 μm [Q = length/breadth: 1.21-1.30], 

subglobose to broadly elliptic, with an ornamentation of isolated warts and ridges up to 2 um 

high. Basidia 20-30×6-8 μm, clavate, two- to four-spored. Basidioles clavate. 

Cheilomacrocystidia 22-30×5-7.5 μm, abundant, subclavate to subfusiform, with acute 

apex, sometimes mucronate, smooth, colorless, thin-walled. Pleuromacrocystidia 35-90×7-

10 μm, projecting, lanceolate or subclavate to subfusiform, with acute apex, sometimes 

mucronate, smooth, colorless, thin-walled. Hymenophoral trama regular; element hyphae 5-

15 μm wide, cylindric. Pileipellis an ixotrichoderm consisting of ascending, cylindric, hyaline 

hyphae 5-10 μm wide. Trama of pileus and stipe heteromerous, colorless, composed of 

massed sphaerocysts (15-40 μm wide) intermixed with relatively few cylindric elements (4-

12 μm wide). Stipitipellis a cutis of interwoven, repent hyphae 2-15 μm wide, cylindric, 

often branched, colorless; caulocystidia none. All tissues without clamp connections. 

  Known distribution: Japan (Kanagawa). Habitat: Solitary to scattered, basidiomata usually 

produced on leaf litter in broad-leaved forest dominated by Quercus myrsinaefolia and 

Quercus serrata, September to October, common. 

  Specimens examined: KPM-NC0007430 (holotype), Izumino-mori, Yamato-shi, Kanagawa-

ken, 13 Oct. 2000, H.Takahashi; KPM-NC0007431, the same place, 14 Oct. 2000, H.

Takahashi. 

  Japanese name: Numeri-hatsu (first collected and named by Mr. Minoru Aoki). 

  Notes: This species is characterized by its pale grayish, strongly glutinous pileus, the 

watery latex without discoloration, the relatively large basidiospores with ornamentation of 

isolated warts and ridges, the ixotrichodermial elements of pileipellis, and its habitat in 

Quercus forests. These characteristics suggest that this species belongs in the section 

Triviales Hesler & A.H.Sm. of the subgenus Russularia (Fr.) Kauffman (Hesler and Smith, 

1979). 

    Lactarius glutininitens appears to be closely related to the following two taxa with an 

azonate, slimy and pale colored pileus: Lactarius trivialis (Fr.: Fr.) Fr. from Europe (Basso, 

1999; Heilmann-Clausen et al., 1998) and North America (Hesler and Smith, 1979) and 

European Lactarius utilis (Weinm.) Fr. (Basso, 1999; Heilmann-Clausen et al., 1998). These 

species differ in having larger basidiomata (pileus 40-150 mm in diam: Heilmann-Clausen et 

al., 1998), a viscid stipe, acrid latex with different coloration, and association with Picea and 

Betula. Lactarius glutininitens is also similar to European Lactarius albocarneus Britzelm. 

(Basso, 1999; Heilmann- Clausen et al., 1998), which differs in forming a cream to buff pileus, 

a viscid stipe, a white, acrid to bitter latex, and association with Abies.