オリーブアシナガタケ
Mycena intersecta Har. Takah., Mycoscience (2007) 48: 342-357.
アクニオイタケ節の仲間で、傘と柄はオリーブ緑色～オリーブ褐色を帯び、ヒダは連絡脈を表します。 欧州産 Mycena viridimarginata (キアシクヌギタケ)に子実体の色が類似しています。
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A Basidiospores. B Basidia and basidioles. C Cheilocystidia.
D Elements of the pileipellis. E Caulocystidia. Bars 10 μm.
肉眼的特徴: 傘は径 8-12 mm, 釣り鐘形で平開せず, 湿時半透明の条線を表し, やや吸水
性, 乾性, 平滑, 最初白色粉状, のち平滑になり, 全体にオリーブ褐色～黄褐色. 肉は傘の中
央部において厚さ 0.7 mm 以下, 白色, 特別な味や臭いはない. 柄は 50-80×0.7-1.2 mm, 円
柱形, 中心生, やせ型, 中空, 乾性, 最初白色粉状, のち平滑になり, 傘よりやや淡色, 根本は
白色の剛毛に被われる. ヒダは直性～やや垂生し, 疎 (柄に到達するヒダは 16-19), 幅 1.5
mm以下, 類白色, 連絡脈を有し, 縁取りを欠く.
顕微鏡的特徴: 担子胞子は 7.5-8.5×5-6 μm, 短楕円形, 表面は平坦, 無色, 非アミロイド
～弱アミロイド, 薄壁. 担子器は 22-31×5.5-7 μm, こん棒形, 1-2胞子性. 縁シスチジアは 27-
40×3-6 μm, 群生し, 不規則な円柱形, 無色, 平坦, 薄壁. 側シスチジアを欠く. 子実層托実質
の菌糸は弱偽アミロイド. 傘の上表皮層の菌糸は匍匐性で, 幅 2-6μm, 平行に配列し, 円柱
形, 疣状～短指状突起に密に被われ, 無色, 薄壁, 非アミロイド. 上表皮下層の菌糸はオリー
ブ褐色, 実質の大部分は偽柔組織状 [pseudoparenchymatous: 著しく肥大し, 複数の隔壁を持
った短形の菌糸細胞] で, 菌糸は平列 し，弱偽アミロイド. 柄表皮の菌糸は平坦または断続的
に疣状～短指状突起に被われ, オリーブ褐色, 菌糸の末端部は外側に屈曲して柄シスチジア
を形成し, 非アミロイド, 薄壁; 柄シスチジアは 13-29×3-7 μm, 広こん棒形から不規則な形状
になり, しばしば1-3個の瘤状分岐物を持つ. 菌糸はクランプを欠く.
供試標本: KPM-NC0008728 (holotype), Izumino-mori, Yamato-shi, Kanagawa pref., 25
Nov. 1999, coll. H.Takahashi; the same place, 27 Nov. 1999, coll. H.Takahashi; the same place,
21 Nov.2000, coll. H. Takahashi.
コメント: 傘と柄はオリーブ褐色で, ヒダは連絡脈を持ち, 縁シスチジアは平坦, 不規則な円柱
形で, 側シスチジアを欠き, 傘と柄の表皮組織は多分枝菌糸からなり, 柄の表皮は広こん棒形
から不規則な形状の柄シスチジアを形成し, 実質は弱偽アミロイドに染まり, 菌糸はクランプを
欠く. 晩秋の頃 (11月), 神奈川県大和市泉の森のシラカシ, コナラを中心とする広葉樹林内の
落ち葉上に発生. 標本は神奈川県立生命の星・地球博物館 (KPM) の標本庫に登録, 収蔵さ
れている.
本種は Maasの分類概念 (Maas 1980, 1988a) によるアクニオイタケ節 (section
Fragilipedes) に属し, 欧州産 Mycena viridimarginata P.Karst. (Breitenbach and Kranzlin
1991; Courtecuisse and Duhem 1994; Kuhner 1938; Lisiewska 1987; Maas 1981, 1988c;
Moser and Julich 1999; Robich 2003) に近縁と思われるが, M. viridimarginata は, 緑色に縁取
られたヒダを持ち, 縁シスチジアは微突形で, 菌糸はクランプを有する.
Mycena intersecta Har. Takah., Mycoscience (2007) 48: 342-357.
Etymology: From Latin, intersecta = intersected, referring to the intervenose lamellae.
Pileus 8-12 mm in diam, conico-convex to campanulate, faintly translucent-striate when
moist, subhygrophanous, dry, minutely white pruinose at first, soon glabrescent, at first olive-
brown (4E5-8) to yellowish brown (5E5-7) overall, then somewhat paler from the margin.
Flesh up to 0.7 mm, white; odor and taste not distinctive. Stipe 50-80×0.7-1.2 mm,
cylindrical, slender, terete, hollow, dry, pale olive-brown (4E3-4) at the apex, olive-brown
(4E4-6) downward, at first entirely white pruinose, glabrescent in age; base white strigose.
Lamellae adnate to subdecurrent, distant (16-19 reach the stipe), up to 1.5 mm broad, thin,
somewhat intervenose, whitish; edges concolorous.
Basidiospores (n = 73 spores of 5 basidiocarps) 7.5-8.5×5-6 μm, Q (length/breadth) =
1.4-1.5, short ellipsoid, smooth, colorless, inamyloid to weakly amyloid, thin-walled. Basidia 22
-31×5.5-7 μm, clavate, mostly two-spored. Basidioles clavate. Cheilocystidia 27-40×3-6
μm, abundant, forming a sterile lamella edge, irregularly cylindrical to strangulated, smooth,
colorless, thin-walled. Pleurocystidia absent. Hymenophoral trama regular; element hyphae 5
-18 μm wide, cylindrical, walls thin, smooth, colorless, weakly dextrinoid. Pileipellis a cutis
of parallel, repent hyphae 2-6 μm wide, cylindrical, densely covered with warty or finger-
like diverticulae, walls thin, colorless, inamyloid; underlying hyphae parallel, olive-brown,
weakly dextrinoid, with short and inflated cells up to 22 μm wide. Stipitipellis a cutis of
parallel, repent hyphae 2-4 μm wide, cylindrical, smooth to sparsely diverticulate, curving
outwards to form clavate or irregularly shaped caulocystidia, olive-brown, inamyloid, thin-
walled; terminal cells (caulocystidia) 13-29×3-7 μm, broadly clavate to irregularly shaped,
often with one to three knob-like excrescences, ascendant. Stipe trama composed of
longitudinally running, cylindrical hyphae 5-12 μm wide, smooth, colorless, weakly dextrinoid.
Clamp connections absent in all tissues.
Known distribution: Japan (Kanagawa).
Habitat: Solitary or scattered, on dead leaves in lowland forests dominated by Quercus
myrsinaefolia Blume and Q. serrata Thunb.ex Murray.
Specimens examined: KPM-NC0008728 (holotype), Izumino-mori, Yamato-shi, Kanagawa
pref., 25 Nov. 1999, coll. H.Takahashi; the same place, 27 Nov. 1999, coll. H.Takahashi; the
same place, 21 Nov.2000, coll. H. Takahashi.
Japanese name: Oriibu-ashinagatake.
Notes: The important combination of features delimiting this species is its glabrescent,
olive-brown pileus and stipe, the somewhat intervenose lamellae, the smooth, irregularly
cylindrical to strangulated cheilocystidia, the absence of pleurocystidia, the diverticulate
pileipellis elements, the broadly clavate to irregularly shaped caulocystidia, the weakly
dextrinoid trama, and the absence of clamp connections. These characteristics place this
species in the section Fragilipedes (Fr.) Quel., as defined by Maas Geesteranus (Maas
Geesteranus 1980, 1988a).
Mycena viridimarginata P.Karst., reported from Europe (Kuhner 1938; Maas Geesteranus
1981, 1988c; Lisiewska 1987; Breitenbach and Kranzlin 1991; Courtecuisse and Duhem 1994;
Moser and Julich 1999; Robich 2003), seems to be closely allied to M. intersecta. The former
species, however, differs in having greenish marginate lamellae, mucronate cheilocystidia,
and clamp connections.
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