1.5 systems offer increased flexibility for call recording configurations.
For each potential destination, it is now possible to fully control the call recording behaviour.
You can now select individual inbound routes and outbound routes. Further, call recording could be switched off for some destinations, and then re-enabled on other destinations.
It may be easier to think of the "No" setting as "I would rather not record this portion of the call, unless I have been told to record this call earlier in the call". In this case, the "Yes" setting would mean "I'd like to record this portion of the call, unless I have been explicitly told not to record the call, earlier in the call flow."
"Force" would then mean "Start or resume recording now" and "Never" would mean "stop recording the call now".
"Don't care" then means "Don't change the call recording status now".
Further details on call recording can be found in the documentation provided by FreePBX.