Herbs, annual to perennial, (shrubs, lianes); cyclopeptides, glycoflavones, anthraquinones, anthocyanins +, betalains 0; true tracheids, fibres +; (wood rayless); sieve element plastid with polygonal central crystalloid; pericyclic fibres +; stomata often diacytic; (cuticle waxes as rodlets); lamina vernation conduplicate or ± flat, stipules +, ± scarious; flowers 4-5-merous; hypanthium +; P = K, quincuncial, + C; common A-C primordia [?level]; A 10; outer secondary parietal cell of anther wall dividing, tapetal cells 2-nucleate; (pollen 6(+) pantoporate); (nectary +, surrounding or abaxial to [esp. antesepalous] A on receptacle); G esp. [3, 5], when 5 alternate with/opposite to P/K, when 3 odd member adaxial, placentation ± axile, styles impressed [distribution?]; ovules with funicle longish to short, parietal tissue 3-10(-30) cells across, (in radial rows), (nucellar cap 2 cells across), (nucellar epidermis dividing along sides of ovules), obturator +, placental, hairy; fruit an utricle/achene/nutlet; (endotesta thickened; endotegmen ± thickened), ?tegmen bar thickenings; post-fertilization embryo sac with (chalazal) haustorium/diverticulum during early development, (suspensor massive), embryogeny solanad[!]; x = 9, nuclear genome [1 C] (0.092-)1.009(-11.079) pg; protein bodies in nuclei; mitochondrial coxII.i3 intron 0; sporophytic self-incompatibility system present.