Chapter 23b

Puccinia

TFTaDMSSoCAR P. acetosae On Rumex acetosa Tay (Fife Drumcarro Hill, M.Stewart, 6 vii 1966; St Andrews? J. Mcdonald vii 1925; Burghead, U. Duncan, vi 1943; Edzell, Angus U. Duncan, viii 1944; Ballinluig, vii 1915; Ballindalloch, ix 1937), Solway (New Galloway, Mossdale, ix 1931), Forth (Dunbar, M. Wilson, vii 1930; Cove, ix 1930), Moray (Forres, viii 1933; Loch Duntelchaig, J.A.Macdonald, 27 viii 1936; Beauly, vi 1929; Kincraig, M.Wilson, viii 1944), Argyll (Oban Glen Lonan, J.A.Macdonald, vii 1925). Also II Shetland Papa Stour R.Watling, 9 viii 1989, Orkney Waulkmill Mainland, R.Watling, 14 viii 1989, Eigg, Edinburgh Univ.Bio.Soc., vii/viii 1949, and Mull 11 vi 1970.On R. acetosella II, Argyll (Taynuilt, viii 1934), Dee (Crathes Castle, D.M.Henderson, 4 viii 1962).

Also known from Lewis, Harris, North & South Uists, Vatersay, Tiree, Coll, Raasay, Skye, Rhum, Eigg, Mull, Colonsay, Lismore, Islay, Arran, Gigha, Kintyre Orkney and Shetland. Also from Forth (Roslin Woods, vii 1821 prob. Greville) Recorded as Uredo bifrons from all but Sutherland but additionally on Polygonum aviculare in Stevenson! See Uromyces polygoni-avilculare below.

TFTaDMSSoCAR P. adoxae On Adoxa moschatellina Tay (II St Andrews, Kenly Den, J.A.Macdonald, 18 iv 1936), Forth (Stirligshire, Ballindalloch, A.Sauten & G.Hadley, 1 v 1971; East Lothian, Gifford Wester Wood, D. M. Henderson, 22 v 1955; III Roslin, vi 1936), Solway (Kirkcudbrightshire, River Cluden between New Bridge & West Cluden, 2 v 1997, O.M.Stewart), Clyde (Glasgow Bishopton, J.R.Lae, 1 v 1948; Chatelherault & Falls of Clyde) and Moray (Inverness Beauly, U.Duncan, 18 iv 1984). In Stevenson recorded from Tay (Glamis Den) and introduced with short description. Edinburgh, W.Greene, 1821; poss. Greville Herb.

TFTaDMSSoCAR P. aegopodii On Aegopodium podograria Tay (III Angus, B.L.Burtt, 11 v 1971; Glen Fincastle, D.M.Henderson, 13 vii 1969; Blairgowrie, 20 vi 1947; Arbroath Waulkmills, U.Duncan, vi 1944; Fife, River Eden near Dairsie Bridge, A.Angus, 17 v 1965; Fife Kingsbarns, A.Angus, 3 vii 1966; Eden Estuary, M.Stewart, 22 v 1966; St.Andrews, Lade Braes, J.A.Macdonald, 6 v 1936), Forth (Edinburgh, Duddingston, M.Noble, iv 1938; Gogarburn, 30 iv 1940; Midlothian Crichton, 6 vi 1924; Fife Largo, 16 vi 1954; Balerno, v 1936; East Craigs High Field, 29 iv 1950; Roslin, 1 vi 1947, 9 v 1952 & 14 iv 1961, D.M.Henderson), Moray (Abernethy,v 1914). Also Mull near Glenforsa hotel, M.C.Clark, 16 vi 1971). Known from Orkney. In Stevenson recorded from Tweed, Forth, Tay, Moray and Clyde.

TFTaDMSSoCAR P. albescens On Adoxa moschatellina Tay (OI Perthshire Killiecrankie, J.Anthony, 10 vi 1958; Perthshire near Innerpefary, D.M.Henderson, 8 v 1966; Blairgowrie, D.M.Henderson, 18 vii 1954), Forth (RBGdn., Edinb.? no further data; East Lothian, Wester Wood, D.M.Henderson, 22 ii 1955; Lothian, Loanhead Burdiehouse, D.M.Henderson, 20 iv 1953; Chirnside Bluestone, Ford, J.Bowden, 2 v 1935), Tweed (Roxburgh, M.Wilson, iv 1941) and Solway (OI & III, Largs, J.R.Leca, 19 iv 1948), Single record from Kintyre. As Aecidium albescens in Stevenson from Tay, Solway and Argyle.

TFTaDMSSoCAR P. albulensis On Veronica alpina Tay (Caenlochan, 1874 & D.M.Henderson, 8 vii 1953) and Argyll (III Glen Nevis, Aonach Mor summit ridge, coll. J.Muscot, 27 viii 1984, A.P.Bennell). As Puccinia veronicanum Tay (Glen Clova, Corrie above Craig Maud, M. Wilson, 13 vii 1909). In Stevenson recorded fror Tweed, Tay, Moray & Clyde all on V. alpina.

TFTaDMSSoCAR P. alii On Allium cepa var. aggregatum Forth (Blinkbonny, Blackhall, Edinb, R.Watling, 18 vii & 27 vii 1989). On A. cyaneum Forth (Afton Terrace, Edinb., D.M.Henderson, 26 viii 1986). On A. porrum, Tay (Fife St. Andrews Bassa Guard, J.Mackie, 21 ix 1975). On A. sativum As P. porrii Solway (Kirkcudbright, D.S.Anderson)On A. schoenoprasum Forth (III, East Lothian Haddington, D.M.Henderson, 28 viii 1987; II & III, Midlothian, G.B.Wallace, 7 vi & I 13 vii 1968; Flotterstone, D.M.Henderson, 12 vii 1968); also Orkney, T. Shearer viii 1930 and at Firth, Watling, 1999. As Puccinia porri on Allium porrum Dee (Aberdeenshire, 2 vii 1948), On A. scorodoprasum as P. porrii Tay (Kingsburn Nursery St Andrews, x 1946), Forth (Logie, Bridge of Allan, vii 1956; Midlothian 10 xi 1959, no further data; Dalkeith, 1 1928; Eskbank x 1923; Musselburgh, 14 11 1928; Pathhead Ford, 18 111 1924; Ferry Rd,1 vii 1979 & 13 viii 1978 and Blinkbonny, Blackhall 31 x 1983 both Edinb.; Spittal Fm., Longniddry, 26 vi 1968), Clyde (Irvine, Ayr, 23 viii 1983), Solway (Dunragit, Glen Luce,no further details) and Dee (nr. Peterhead, x 1969).

On Allium vineale as P. porrii Tay (Arbroath Seaton, U.Duncan, viii 1944; Fife, vii 1940; Angus Arbroath Carlingheugh Bay, U.Duncan, viii 1944) and Kintyre. On Allium spp. Tay (Glen Fincastle, D.M. Henderson, 13 vii 1969) and Solway (III Wigtownshire Sandhead, D.M.Henderson, 18 vii 1958). Recorded from Arran. Very common on chives in the North East and on leek in the south East and S.Central areas of Scotland (Foister, 1958). As Uromyces alliorum recorded from Tay (Dun) and as Uredo alliorum  ‘Scotland, M.C.Cooke in Stevenson and as P. allii with short description in Stevenson Add.4 added for Dee (Nigg, Aberdeen, W.Forest) and as U. alliorum also for Dee.

TFTaDMSSoCAR P.angelicae On Angelica sylvestris, III/II Scotland, D.M.Henderson, 14 ix 1963 no further data; Ross (Totaig, M.Wilson, viii 1934) and Tay (Ballenluig, M.Wilson, vii 1915; Dunkeld, M.Wilson, viii 1932). Record from Barra ex Campbell, 1936.

TFTaDMSSoCAR P.annularis On Teucrium scorodonia Tay (Dunkeld The Hermitage, B.Coppins, 27 ix 1975; Dunkeld, D.M.Henderson, 5 ix 1953 & viii 1977; Ballenluig M.Wilson, ix 1921; Black Wd., Rannoch, D.M.Henderson, 18 ix 1960; Faskally, D.M.Henderson, 6 viii 1957; Angus, Edzell, U.Duncan), Dee (Dinnet Oakwood, D.M.Henderson, viii 1963; III Aberdeen, D.M.Henderson, 6 ix 1975), Forth (Penicuik House, R.B.Johnstone, ix 1934), Tweed (Peebles, N.L.Alcock & M.Wilson March 1920s; III Dryburgh Sir Walter Scott's View A.& F.M.Bennell, 5 viii 1978), Ross (Wester Ross, D.M.Henderson, 2 ix 1952; Strome, D.M.Henderson, 22 ix 1960; Ross-shire, D.M.Henderson, 11 ix 1963), Argyll (Benmore Gdn., D.M.Henderson, 15 ix 1959; Argyllshire, Balvicar Island, D.M.Henderson, 3 viii 1984; Loch Seil, D.M.Henderson, 3/8/1984), Solway (Moniaive, M.Wilson, ix 1931) and Moray (III, Loch Duntelchaig, J.A.Macdonald, 29 viii 1935; Aviemore, M.Wilson, ix 1927). Also Mull 6 ix 1968, Dunvegan south end of castle estate mixed woodland coastal, Skye, AP & FM Bennell, 21 x 1980. Records from Raasay, Skye, Mull, Colonsay, Islay, Arran, Gigha and Kintyre. Recorded in Stevenson as P. scorodoniae from all Watersheds except Solway, Argyle and Sutherland; also from Argyle in Add 4.

TFTaDMSSoCAR P.antirrhini On Antirrhinum majus Tay (Fife St.Andrews, A.Angus, 26 v 1965), Forth (Blnkbonny, Blackhall, Edinb., R.Watling, viii 1981, 9 viii 1982 & 10 ix 1984; East Craigs, C.E.Foister, ix 1939; Orchard Brae, Edinb., 24 viii 1976). On A. glutinosum Tay (St.Andrews Bot. Gdn., J.A.Macdonald, 21 ix 1955).

On Antirrhinum spp., Clyde (Auchincruive, 29 iv 1944), Forth (II Edinburgh, D.M.Henderson, 14 ix 1969; East Lothian, Whitekirk, Miss Broadfoot) and Tay (Fife St.Andrews East Scores, x 1937).

TFTaDMSSoCAR P. apii On Apium cultivars; recorded from Tay (Balnamoon, Rev. Anderson) in Stevenson.

TFTaDMSSoCAR P. arenariae On Arenaria sp. Tay (Blairgowrie Cluny, 23 vi 1951). On Moehringia trinervia Forth (Stirlingshire, P.H.Davis, 7 vi 1960). On Sagina normaniana Dee (Glen Clova, Feula Burn, U.Duncan, vi 1940). As P. saginae on Sagina x normaniana (as procumbens?) Tay (Glen Clova, vi 1940, U. Duncan). On S. procumbens Tay (Pitcardly Hill, Forfar, x 1944, U. Duncan). On Sagina sp. Dee (Angus Forfar, U.Duncan, x 1944), Argyll (no further data D.M.Henderson, 3 viii 1984) and Lewis, North Uist, Skye and Islay. On Silene dioica Tay (Perthshire, Kippen, D.M.Henderson, 12 ix 1959), Forth (on Melandrium pratense? East Lothian Tantallon, D.M.Henderson, 29 viii 1965; Dalmeny, D.M.Henderson, 16 xi 1968; Midlothian Dalkieth Estate, D.M.Henderson, 13 x 1956), Clyde (Chatelherault; Lanarkshire Hamilton Cadzow Park, D.M.Henderson, 14 ix 1959), Tay (Arbroath Redhead, U.Duncan, v 1944, ix 1944 & x 1944 and ix 1943; Fife Parkmill, A.Angus, 19 x 1986; St.Andrews Lade Braes Walk, A.Angus, 30 x 1965; Perthshire Ballinluig, vii 1915 & as P. lychnidearum, road near Balmutto, 11 x 1936 host as Lychnis diurna) and ‘Scotland’, D.M.Henderson,16 vii 1962 (no further data). Also records from Skye, Mull, Colonsay, Arran, Kintyre, Orkney and Shetland. On Stellaria graminea Moray (Forres, 16 x 1880), Dee (III Aberdeenshire, D.M.Henderson, 9 ix 1975) and Tay (Fife Kittock's Den, A.Angus, 22 viii 1974). On S. media Clyde (Lanarkshire Hamilton Cadzow Park, D.M.Henderson, 14 ix 1959), Tay (Dunkeld House, D.M.Henderson, 6 ix 1953), Solway (Thornhill, G.F.Scott-Elliot, ix 1942), Forth (Dalmeny, D.M.Henderson, 6 x 1968) and Orkney. On S. nemorum III Clyde (Hamilton, J.A.Macdonald, 14 ix 1954). As Puccinia lychnidearum on Stellaria nemorum Forth (East Lothian Tantallon Castle, R.D., ix 1939; Midlothian Roslin, x 1935), Solway (Ayrshire, Miss.M.Campbell). On S. uliginosa  Ross (Gairloch, viii 1934). On Stellaria sp. Tay (Perthshire Dunkeld, D.M.Henderson, 23 viii 1953), Clyde ( III Hamilton, D.M.Henderson, 14 ix 1959). Collection on Claytonia sibirica from Clyde (Arran); recently re-found at Forth (Thruipmuir Balerno, Edinb. 2008) apparently requiores further investigation. On Dianthus cultivated Forth (Midlothian, A.W.Boyd, late 1956) and noted by Foister (1958) on Dianthus barbatus (Sweet William).

TFTaDMSSoCAR P. asparagi First recorded in 1824 around Ediburgh but not seen since (Foister, 1958). On dead stems of Asparagus officinalis, ‘Autumn Edinburgh’ Greville, Fl. Edin. but no material located in E.

TFTaDMSSoCAR P. behensis On Silene dioica Kergord, Shetland. As P. lychnidearum Solway (Ayr, N.L.Alcock, ix 1926), Forth (Roslin, 7 28 viii 1943). As P. silenes on Lychnis dioica Tay (West dunes, St Andrews, 6 ix 1936).

Noted from Tweed, Tay and Moray in Stevenson.

TFTaDMSSoCAR P. betonicae On Stachys officinalis. Indicated as occurring in Scotland but noted as rare, Henderson, 2000. Recorded in Stevenson from Forth; Greville record in Fl. Edin. but no material traced in E

TFTaDMSSoCAR P. bistortae, III, On Polygonum bistorta Argyll (Argyll, M.Wilson,1943; Argyle Loch Eck, M.Wilson,1943), Solway (Dumfries, Langholm Lodge, D.M.Henderson, 10 ix 1951). On Pesicaria viviparia Tay (Loch Moraig, Blair Atholl, c360m., 17 vii 2009). By the distribution the records in Stevenson as P. polygonorum belong here; known from all but Sutherland, Solway, Argylle & Ross. As Puccinia polygoni-vivipari III on Polygonum viviparum from Tay (Ben Lawers, R.Watling, 3 ix 1983; Glen Tilt, R.W.G. Dennis, 26 vii 1942; Blair Atholl, D.M.Henderson, 8 vii 1968) and Moray (Inverness-shire Ben Alder, R.Watling, 7 viii 1980; Boat of Garten, M.Noble, vi 1949; Loch Duntelchaig, J.A.Macdonald, 27 viii 1936). Recorded in Stevenson as P. bistortae from Tay (Mar Lodge, Greville 1821), Tay (Glenshee Buchanan White) and Moray. As Puccinia conopodii-bistortae, Edinb. (experimental inoculations), 9 v 1951.

TFTaDMSSoCAR P. brachypodii var. brachypodii Including collections labelled P. baryi on Brachypodium sylvaticum from Forth (Stirling University, D.M.Henderson,6 x 1968; II Critchton, D.M.Henderson, 4 x 1959; II Pittenweem, M.Wilson?, viii 1919), Tay ( Dunkeld, D.M.Henderson, 23 viii 1953; II Perthshire, Banks of Tay, D.M.Henderson, 18 ix 1959 & 3 ix 1961; Edzell, U.Duncan,ix 1944; St.Andrews Boarhills, J.A.Macdonald, 22 ix 1939; near Blair Atholl, D.M. Henderson, 14 vii 1968), Solway (near Newton Stewart, M.Wilson?, ix 1929; II south of Newton Stewart, D.M.Henderson 21 ix 1933; Langholm, D.M.Henderson, 30 ix 1951). Records from Rasaay, Skye, Mull, Arran as P. baryi (Boyd) and Islay; also Garvellochs and Orkney. In Stevenson Add. 5 recorded from Dee (Aberdeen, Prof. Trail).

TFTaDMSSoCAR var. arrhenatheri On Arrhenatherum elatius Forth (Edinb., M.Wilson?, 7 viii 1921 & xii 1924 & Barnton, Edinb.,vii 1922; East Lothian Gladsmuir, D.M.Henderson, 24 vi 1963; II, Newbattle, M.Noble 10 viii 1955; II Dalmahoy Estate wall, D.M.Henderson, 29 vii 1954; Granton, D.M.Henderson, 26 vii 1971; Penicuik, R.W.G. Dennis, 14 ix 1941; Kirkliston, J.A.Mcdonald, 20 vi 1935), Arran (Lochranza, I.Jorstad, 12 ix 1959), Clyde (Chatelherault), Dee (Brechin, G.Wallace, 21 vi 1946), Moray (Beauly, M.Wilson?, 1925), Tweed (Peebles, G.B.Wallace, 6 ix 1925), Tay (Tayport, M.Wilson?, vi 1932; II Kittocks Den, J.A.Mcdonald, 8 x 1936 and Burn’s Nursery, J.A.Mcdonald, 18 ix 1938 both St Andrews); also Skye, D.M.Henderson,12 vii 1968, Mull, D.M.Henderson, 10 ix 1968 and Shetland 29 viii 1983 incl. Tramere Reserve, R.Watling, 30 viii 1983. On Deschampsia caespitosa II Moray (Inshriach, Aviemore, 23 ix 1959), Forth (Penicuik as Uredo airiae 14 ix 1941, R.W.G.Dennis; Kippen, 12 ix 1959, D.M.Henderson; Balerno, 31 v 1959, D.M.Henderson & Thruipmuir, Balerno, 18 xii 1954; Pathhead 21 ix 1925, G.B.Wallace), Tweed (Glentress, Peebles, 17 viii 1954, D.M.Henderson; Stow, ix 1928; Dawyck, 7 v 2006, R.Watling), Argyll (I Ben Lui, 457m., 16 ix 1959), Tay (Edzell, J.A.Macdonald, 13 ix 1958; Creag an Lochan, Ben Lawers, 609m., 17 ix 1959; Glen Lochay, Perthshire, 229m., 1.ix 1961, D.M.Henderson) and as P. deschampsiae Lewis, Skye, Rhum, Mull, Islay, Arran and Gigha.

TFTaDMSSoCAR var. poae-nemoralis Collections labelled P. poae-nemoralis belong here.

On Anthoxanthum odoratum, Arran, Tay (Glen Doll, 913m., 1 vii 1946; Cliffs St Andrews, 23 x 1947; Meall nan Tarmachan,  M.Wilson?, 9 x 1919; Ballinluig, M.Wilson?, viii 1915; II Spittal of Glenshee, R.Watling, 6 vii 1976), Sutherland (Ben Loyal, M.Wilson?, ix 1931 as Uredo anthoxanthina), Forth (as Uredo anthoxanthina Edinburgh, M.Wilson?; RBGdn., Edinb. D.M.Henderson, 25 v 1955), Tweed (Peebles, Glentress, M.Wilson?, vii 1929), Ross (Fort Augustus, M.Wilson?, v 1925), Moray (II Glen Affric,D.M.Henderson,23 viii 1955), Argyll (Ben Lui, M.Wilson?, x 1914 & ix 1921, as U. anthoxanthina) and Clyde (Arran, Brodick, M.Wilson?,  vi 1931, as U. anthoxanthina). As P. poae-nemoralis Tay (Tentsmuir 23 v 1960; Glen Doll, 1 vii 1946, 914m. D.M. Henderson; Cliffs St Andrews, 23 x 1947). On Glyceria maritima as Uredo glycereae, Britain ?, Miss Knagg, v 1925. prob. same as Forth (West Lothian Bo'ness,Miss Knagg, v 1925). On Poa annua, Clyde (II, Hamilton, D.M.Henderson, 14 ix 1959), Tay (Dunkeld, banks of Tay, D.M.Henderson, 18 ix 1959). On Poa annua Kintyre. On P. nemoralis, Forth (West Lothian canalside land West of Ratho, AR & FM.Bennell, 22 vi 1978), Dee (Glentanar, 5 ix 1975), Tweed (Peebles, Glentress, vi 1929)and Tay (Dunkeld banks of Tay, D.M.Henderson, 18 ix 1954). As P. poae-nemoralis Dee (Glentanar 5 ix 1975, A. Angus)

On P. trivialis Tweed (II/III Peebles, Glentress, xii 1929). No host but II present Forth (Midlothian Pathhead, J.Phelps, 20 vii 1921 & vii 1930), Tay (Ballinluig, vii 1915). On Poa sp., Gigha, Tay (I, Blairgowrie, D.M.Henderson, 29 v 1955), Forth (East Lothian Dunbar, D.M.Henderson, iv 1961; Orchard Road, Edinb.,R.Watling, 30 x 1975) and II on Poa sp? Orkney 30 vi 1963. On Puccinellia maritima, Orkney, G.Hadley, 30 vi 1963; also Orkney, F.R.Bullard, vi 1963 and Shetland. On P. procumbens, Dee  (Kincardinshire Gourdon, A.Somerville, 27 viii 1903; 10 24/8/1949).

Also Scotland, no further details, M.Wilson?, vii 1914 as U. anthoxanthina. Also records of this variety on A.elatius and as P. arrhenthericola from Iona, D.M.Henderson, 13 ix 1968, Forth (Aberlady, 8 ix 1959), Tay (Tentsmuir, 23 v 1960), Islay and Orkney. Recorded in Foister (1958) on foddder grasses – Anthoxanthum odoratum and Arrenatherum elatius.

TFTaDMSSoCAR P. bulbocastani On Bunium bulbocastaneum as Aecidium bunii in Stevenson and recorded from Forth and Tay. Not in Check List for Scotland but see Wilson & Henderson.

TFTaDMSSoCAR P. buxi, III, on Buxus sempervirens from Tweed (Dawyck, J.A.Mcdonald, 1 x 1946), Tay  (Fife St.Andrews Boarhills, A.Angus, 12 iv 1969 & 8 v 1981; Den of Airlie, R.Watling, 18 iv 1981; Dunkeld House, D.M.Henderson, 6 ix 1953), Solway (Lockerbie, Miss Maxwell, 1926; on Buxus sp., Castle Douglas, D.S.Anderson, iii 1946), Forth (Arniston, nr. Gorebridge, D.M.Henderson, 7 vi 1954; Arbroath Kinblethmont, U.Duncan, 9 v 1944) and Argyll (Arisaig, G.Anderson, 18 iv 1981). Also Mull Torosay Castle, M.C.Clark, 20 vi 1997 and Islay.

Recorded in Stevenson from Forth & Tay.

TFTaDMSSoCAR P. calcitrapae On Carthamus tinctorius as Puccinia carthami from Forth (Midlothian, Bush Estate, Dr.Gilmour, D.M.Henderson, 13 x 1959). On Carduus crispus as P. carduorum from Tay (II/III from Tay (Stravithie, J.A.Mcdonald). On Centaurea scabrosa, Forth (Largo, 22 vi 1954; II/III Arbroath, Carlingheugh Bay, U.Duncan, viii 1944; Fife, D.M.Henderson, 29 viii 1960), Tay (Angus Seaton, U.Duncan, viii 1944); also Scotland no further details D.M.Henderson 6 vii 1958. On Centaurea nigra some records as P. centaureae Clyde (Ayrshire near Kilmarnock, M.Wilson, 27 vi 1940), Tay (II Redhead near Angus, U.Duncan, v 1944; Ballinluig, M.Wilson, vii 1915; II/III Kindrogan Estate, F.M.Blair, 30 I 1976; Blairgowrie, D.M.Henderson, 20 viii 1973; shores of Loch Tay, D.M.Henderson, 17 ix 1959 & near Loch Tay, 17 ix 1959; II/III St.Andrews, J.A.Macdonald, 11 x 1935; III East Cliffs, St Andrews, J.A.Macdonald, 8 x 1936), Forth (Tantallon Castle, R.W.G.Dennis, ix 1939; III Leith, ix, D.M.Henderson; II Gifford, Saltoun Wood, D.M.Henderson, 8 x 1955; II/III Cove, M.Wilson, viii 1930; Carriber Glen Linlithgow, D.M.Henderson, 27 ix 1957), Ross (Gairloch, vii 1936), Sutherland (III, West Sutherland Oldshore More, B.J.Coppins, 23 viii 1983). Also II, Salen, D.M.Henderson, 8 ix 1968 & Dervaig,4 ix 1968 both Mull and Iona, D.M.Henderson, 13 ix 1968; also Lettermore Farm, M.C.Clark, 24 vi 1971. Records from Harris, Vatersay, Tiree, Coll, Raasay, Skye, Rhum, Mull, Islay, Arran and Kintyre; also Orkney. On Centaurea calciptrapa from Tay (II/III, Perth, Dunning, D.M.Henderson, 20 x 1975). On Cirsium palustre as Puccinia cirsii Moray (II Inverness Loch Duntelchaig, J.A.Macdonald, 1936), Tay (Fife Boarhills, J.A.Macdonald, 16 ix 1939). Also Shetland, Fetlar, R.Watling, 23 viii 1988 & Yell, R.Watling, 8 ix 1986. With other records from Orkney, Lewis, Tiree, Raasay, Skye, Rhum, Mull, Colonsay, Oronsay, Lismore, Islay, Jura, Gigha, Arran and Kintyre. Also recorded as P. cirsii with short description in Stevenson from Tay (Dupplin Castle, M.C.Cooke) and as Trichobasis from samer locality); Moray added in Add.1, Tay (Rannoch, Buchana White) in Add. 2 and Dee in Add. 8. ‘On leaves of Compositae summer-autumn’ as P. compositarum in Stevenson from all Watersheds except Argylle, Ross, Sutherland Solway and Moray and P. syngenesiarum for all but Sutherland, and the western Watersheds; Moray added for P. compositarum in Add.1.

TFTaDMSSoCAR P. calthae, III On Caltha palustris from Moray (Loch Duntelchaig, J.A.Macdonald, 19 viii 1936), Tay (Angus Tarfside, G.B.Wallace, ix 1933; Angus Rosaie Muir, U.Duncan, ix 1944), Sutherland (West coast, Store, R.W.G.Dennis, 22 viii 1944), Clyde (II/III. Loch Lomond, Rossdhu, D.M.Henderson, 11 ix 1959), Argyll (Mallaig, M.Wilson, viii 1929) and Ross (in cove near Gairloch, J.Webster, ix 1975). Also Orkney and Shetland, Fetlar Tresta Links, R.Watling 25 viii 1983, Skye Broadford, vii 1932 and Tiree, M.Wilson, viii 1937. Also records from Lewis, Harris, Coll, Raasay, Islay, Arran and Gigha. In Stevenson recorded from all eastern Watersheds except Sutherland and not from west and as Aecidium calthae all the above incl. Clyde and Argyle. May include rceords of P. calthicola.

TFTaDMSSoCAR P. calthicola, III, On Caltha palustris from Tay (St.Andrews Kinkell Braes,  M. Stewart, 17 vii 1966; Stanley, D.M.Henderson, 7 iv 1965; Glen Fincastle Resevoir Loch, D.M.Henderson, 6 vii 1968 & 14 x 1970), Sutherland (Isbister, R.Watling, date), Forth (RBGdn., Edinb., Demonstration Garden, A.P.Bennell, 20 vii 1981; OI Kirkliston, nr. Edinb. D.M. Henderson, 8 v 1949) and Clyde (0I Arran, Brodick, vi 1931). Also Shetland Mainland Voe, P.James, 10 vi 1981, Iona, D.M.Henderson, 13 ix 1968, II & III, Mull, Killichronan, M.C.Clark, 20 vi 1971, North Uist, D.M.Henderson, 10 viii 1968, II/III Eigg, Edinburgh Univ.Biol. Exped., vii/viii 1949 and Orkney, Stronsay, J.Sinclair, 29 viii 1939. Records from Barra, Vatersay, Mull, Colonsay, Arran and Lismore; also Orkney. As Puccinia zopfii, Argyll (Taynuilt, M.Wilson, vii 1921), Tay (Boarhills, 22 ix 1939; Cameron St Andrews, 20 ix 1958; East Cliffs, St Andrews, 2 v 1936), Moray (Loch Duntelchaig, 19 viii 1936), Sutherland (Tongue, M.Wilson, viii 1931), Forth (Balerno, M.Wilson, 1945), Tay (Ballindaloch, M.Wilson, ix 1937), Dee (Aberdeenshire Lumphanan Bog, M.Wilson, 21 vii 1922) and Tweed (Westruther, M.Wilson, ix 1933); also II/III Skye Skerinish, M.Wilson, vii 1932.

TFTaDMSSoCAR P. campanulae III On Campanula rotundifolia, Tay (Glen Lyon, Meall Gruaidh, D.M.Henderson, 27 vi 1970) and ‘Britain’ ? D.M.Henderson, 27 viii 1958. In Stevenson recorded from Tay, Argylle & Dee.

TFTaDMSSoCAR P. caricina var. caricina Many records simply as P. caricis. On Carex aquatilis from Tay ( Blairgowrie, A.C.Jermy, viii 1990). Recorded by Dennis from Caithness and Inverness-shire. On C. binervis II, Scotland, no further data, D.M.Henderson, 26 viii 1953 and 17 ix 1959), Moray (II Loch Dutelchaig, J.A.Macdonald, 6 vi 1936, 12 vi 1938, 3 & 30 vii 1939, 5 coll., viii 1946), Ross (II Plockton, D.M.Henderson, 17 vi 19550 and Tay (Perthshire, D.M.Henderson, 1 ix 1961). Also known from Lewis, Harris, South Uist, Raasay, Skye, Mull, Ulva and Islay; also Shetland. On C. hirta from Tay (III St.Andrews Kittocks Den, J.A.Macdonald, 25 I 1937) and II Islay. On Carex hostiana for Dee, Sutherland & Wester Ross, Dennis (1980). On C. nigra, ‘Scotland’, D.M.Henderson, 6 x 1969 and Tay (III Kindrogan, D.M.Henderson, 25 ix 1964), Moray (Affric, D.M.Henderson, 10 ix 1963). On C. pallescens II ‘Scotland’, D.M.Henderson, 1 ix 1961, no further data and Tay (O/II Loch Tay, D.M.Henderson, 1 ix 1961). Also may include collections from Harris, Mingulay, Rhum and Mull. See also Puccinia paludosa & P. uliginosa. On C. panicea from Solway (Castle Douglas, R.Watling, 12 ix 1959). On C. rostrata from Dee (Aberdeenshire Glentanar, D.M.Henderson, 5 ix 1975; Dunecht, D.M.Henderson, 9 ix 1975). Possibly also includes collections from Harris and Coll. On unident. Carex spp., Argyll (Argyll, D.M.Henderson, 3 viii 1984), Sutherland (Ben Hope, R.Watling, 17 vii 1988), Tay (Loch Tay, D.M.Henderson, 1 & 2 ix 1961) and Solway (Castle Douglas, R.Watling, 12 ix 1959). Also records for Tiree, Colonsay and Kintyre. As P. striola in Stevenson from Tweed, Forth, Tay and Moray ‘ on carices’.

TFTaDMSSoCAR P. caricina var. magnusiana On C. riparia Tweed (Hoselaw Res., Roxburghshire, J.Muscott, 5 vii 2009).

TFTaDMSSoCAR P. caricina var pringsheimiana On C. goodenovii = C. nigra Solway (III Perthshire, Thornhill, M.Wilson, viii 1932, see above and again as on C.aquatilis, M.Wilson, viii 1932). Needs checking. On Carex spp, Tay (Perthshire Thornhill, Ferguson, 1931?) and Moray (Ballindalloch, M.Wilson, 5 ix 1937). On Ribes rubrum from Moray (I Ballindalloch, D.F.Mackenzie, vii 1936). As P. pringsheimiana Moray (Dallas manse, 1984; Coulmany Moray, 1946) and Tay (Fincastle nr. Pitlochry 1953; Fettercairn, Kincardineshire, 1954) fide Foister (1958). On R. sanquineum from Mull, M.Wilson, 1933 and Skye, Skirinish, M.Wilson, vii 1932. On R. spicatum (as R.petraeum); recorded by Wilson. On Ribes uva crispa Lettermore Farm, Mull M.C.Clark, 24 vi 1971 and Orkney Stromness, G.Hadley, 27 vi 1963. Also Ross (Gairloch, vii 1936), Tay (Perthshire Glenfarg, 7 v 1950), Solway (Dumfriesshire Sanquhar, N.Young, v 1939; Lockerbie, I.Maxwell, vi 1925), Forth (Edinburgh Blinkbonny Avenue, R.Watling, 10 iv 1992; Stirligshire Bridge of Allan, M.Wilson), Clyde (Dunbarton, Miss Broadfoot, iv 1945; Benmore, M.Wilson, v 1932) and Argyll (Gigha, G.Hall, 15 vi 1984). Records from Harris, Skye, Mull, Jura, Islay and Gigha and Shetland. On R. grossulariae as P.pringsheimiana from Arran and Tay (Glen Lyon, 24 vi 1931) and as P. ribesi-carici Forth (Edinb., 2 vi 2004). Records as P. ribis on Gooseberry (R. uva-rispa) in Foister (1958) indicate serious infections in commercial crops in 1927, 1938, 1939, 1940, 1948 & 1957. In Stevenson as Aecidium grossulariae from all Watersheds except Sutherland, Solway, Argylle and Ross. As Puccinia striola recorded for Dee in Add. 8.

TFTaDMSSoCAR var. ribesii-pendulae On Carex pendula Forth (Gifford 2 xi 2008 as P. ribesii-caricina) and Clyde (Chaterherault) and Islay.

TFTaDMSSoCAR var. ribis nigri lasiocarpae Single collection on Carex lasiocarpa from Moray (Aviemore, Loch Pityoulish, D.M.Henderson, 23 ix 1959).

TFTaDMSSoCAR var. ribis nigri paniculatae On Carex paniculata II from Tay (Angus Resteneth Bog, U.Duncan, x 1944) and Forth (Midlothian Crichton Castle, D.M.Henderson, 5 x 1959).

TFTaDMSSoCAR P. chaerophylli, On Anthriscus sylvestris many records as Chaerophyllum. Known from Tay (Stanley, M.Wilson, vii 1930; III Dunkeld, D.M.Henderson, 2 ix 1953; III Forfar Pitscandly Hill, U.Duncan, x 1944; Kirriemuir Longbank, R.W.G.Dennis, 10 v 1944, II/III St.Andrews, J.A.Macdonald, 14 ix 1936 & 22 v 1937; Forfar, D.M.Henderson, 22 vi 1964; Kindrogan, D.M.Henderson, 7262, 25 ix 1964), Dee (Aberdeen,? E.G.Gray v 1945; Aberdeen, D.M.Henderson, 9 ix 1971), Sutherland (Scoune, R.W.Dennis, 26 viii 1944) and Forth (Angus, Lunan Bay, U.Duncan, vi 1944; Kinghorn, M.Wilson, v & vi 1928), Argyll  (II/III Taynuilt, M.Wilson, viii 1934) and Moray (II, Loch Duntelchaig, J.A.Macdonald, 8 vii 1936). Known from S.Uist, Barra, Tiree, Coll, Skye, Mull, Arran, Colonsay and Kintyre; also found on Orkney. On Myrrhis odorata from Clyde (Arran Lagg, Waterson), Tay (Boarhills Fife, A.Angus, 28 v 1982; Perthshire, Kippen, D.M.Henderson, 12 ix 1959; Fife Dura Den, A.Angus, 1 x 1968 & J.A.Macdonald, 1937 & 13 ix 1940; Perthshire, no further data, D.M.Henderson, 20 ix 1965; Strathkiness, A.Angus, 13 v 1966; Arbroath, U.Duncan, v 1944; II/III Dunkeld, Hermitage, R.G.Woods, 24 ix 1985; Aberfeldy roadside, D.M.Henderson, 26 vi 1971; Kirkmichael, R.W. D.G. Dennis, 10 viii 1941; III Angus Den of Airlie, D.M.Henderson, 22 viii 1968; Brechin, G.Walllace, 13 vi 1946 and ? 26 ix 1924; St. Andrews, J.A.Mcdonald, x 1936), Forth (Granton Gasworks, Edinburgh, R.Watling, 13 v 1962; Dunbar, D.M.Henderson, iv 1961; I Fife, Torryburn, B.J, Coppins, 25 v 1985; Dalkeith, M.Wilson, 14 iv 1921; III Crombie Point, B.J.Coppins, 13 ix 1975; II/III Midlothian Crichton, G.B.Wallace, x 1924; II/III Granton foreshore, Edinb., D.M.Henderson, 5 vii 1966; III Balerno, D.M.Henderson, 26 ix 1953; III Cramond, Edinb., B.L.Burt, 4 ix 1951; III Linlithgow, D.M.Henderson, 27 ix 1959; Kirkliston, River Almond, J.A.Macdonald, 27 v 1935; OI Canonmills,A.P.Bennell), Dee (III Aberdeenshire Glentanar,D.M.Henderson, 5 ix 1975 & Aberdeen, M. Wilson) and Moray (Loch Ness Drumnadrochit, U.Duncan, iv 1943). Also ‘Scotland’, D.M.Henderson, 13 vii 1969. Known on this host from Mull, Islay and Kintyre; also Orkney. Host not given Tay (I/II/III Stormont Loch, Blairgowrie, D.M.Henderson, 26 vi 1963). Greville collection in E as P. umbelliferarum and collection labelled ‘Puccinia umbelliferum on Myrrhis odorata, ix 1821’.

TFTaDMSSoCAR P. chrysosplenii III, On Chrysoplenium oppositifolium Tay (St.Andrews Kenely Den, J.A.Macdonald, 9 iv 1938; Glen Farg, D.M.Henderson, 28 ix 1952; Angus, D.M.Henderson, 30 x 1966), Solway (Dumfriesshire, Thornhill, N.L.Alcock, ix & x 1924 & M. Wilson ix 1925) and Forth (Crombie, G.B.Wallace, vii 1933; Bridge of Allan, Dr.Cadman, 27 vi 1943) and Ross (Totaig, M. Wilson, viii 1934). Noted by Foister (1958) on nursery stock of Rockfoils. Also known from Mull, Carsaig, R.Watling, 6 ix 1976 and Ulva, D.M.Henderson, 11 ix 1968, Orkney N. Hoy, Lewis, Eigg, Arran and Kintyre. In Stevenson recorded from Tay, Forth & Moray; Clyde added in App.1.and Dee in Add 3.

TFTaDMSSoCAR P. circaeae II, On Circaea intermedia from Tay (Loch Tay, D.M.Henderson, 1 ix 1961) and Ulva, D.M.Henderson 11 ix 1968. On Circaea lutetiana Tay (Dunkeld, D.M.Henderson, 30 viii 1953; Perthshire, Kippen, D.M.Henderson,12 ix 1959; Blairgowie, U.Duncan, ix 1952; Blaigowrie Craighall, D.M.Henderson, 30 ix 1952), Clyde (Dumbartonshire, ix 1836; Ayr, N.L.Alcock, ix 1926; Culzean, N.L.Alcock, ix 1926). Also ‘Britain’ ? M.Wilson, viii 1923. On Circaea sp., from Solway (Thornhill, Miss Maxwell, ix 1924). Records from Raasay, Skye, Islay, Arran and Mull. In Stevenson recorded from Forth, Tay and Moray; Clyde added in App.1.and Moray in Add 4.

TFTaDMSSoCAR P. clintonii, III On Pedicularis sylvatica Argyll (Kilninver, D.M.Henderson, 28 viii 1983; Ben Lui, D.M.Henderson, 16 ix 1959; Arisaig, D.M.Henderson, 16 v 1964) and Ross (Lochcarron, D.M.Henderson, 11 x 1959; Kishorn, D.M.Henderson, 6 ix 1952 & 27 vi 1957). Also Mull Salen, D.M.Henderson, 16 ix 1968 and ‘Scotland’, no further data, D.M.Henderson, 14 ix 1963. In Stevenson as Aecidium pedicularis on P. palustris for Tay and Dee.

TFTaDMSSoCAR P. cnici On Cirsium palustris from Tay (Glen Loy plantation, R.Watling, 27 vi 1984).

On Cirsium vulgare some records as on Cirsium or Cnicus lanceolatus, Tay (Airlie Castle, A.P.Bennell, 19 v 1981; Perthshire, Kippen, D.M.Henderson, 12 ix 1959;), Clyde (Arran Lochranza, I.Jorstad, 12 ix 1959; Hamilton, D.M.Henderson, 14 ix 1959), Forth (II/III East Lothian, Garleton Hills, A. P. & F.M.Bennell, 1 x 1978; Penicuik, R.W.G.Dennis, 14 ix 1941). Also same host Shetland, Noss, R.Watling, 8 viii 1989 & Yell West Sandwick, R.Watling, 26 viii 1983 and II Orkney South Ronaldsay, G.Hadley, 28 vi 1963. Single collection labelled ‘Scotland’, no further details, D.M.Henderson, 17 ix 1959. As Puccinia cirsii-lanceolati Forth (Eskbank Midlothain, D.M.Henderson, 23 ix 1953 & 13 vi 1954; Gullane, M.Wilson vii 1929), Moray (Aviemore, M.Wilson ?, ix 1927; Inverness near Foyers, U.Duncan, viii 1942; II/III Loch Duntelchaig, J.A.Macdonald, 29 viii 1936) and Tay (II, Spittal of Glenshee, R.Watling, 6 vii 1976; Glen Tilt, R.W.G.Dennis, 26 vii 1942; Angus, U.Duncan, vi 1944; II Fife Dura Den, J.A.Macdonald, 7 ix 1959) and Dee (Aberdeenshire Bridge of Don, R.W.G.Dennis, 11 viii 1944). Also Mull, Glen Aros, M.C.Clark, 22 vi 1971. Also known from Tiree, M.Wilson ?, viii 1937. Records from St Kilda, Lewis, Harris, N. Uist, Vallay, Barra, Mingulay, Tiree, Coll, Raasay, Skye, Rhum, Eigg, Mull, Eigg, Ulva, Geometra, Iona, Lismore, Colonsay, Islay, Jura, Arran and Kintyre; also Orkney and Shetland. Greville collection in E ‘about Edinb. as Uredo suaveolens var. B on C. lanceolatus.

TFTaDMSSoCAR P. cnici-oleracei On Achillea millefolium Clyde (Carnwath, D.M.Henderson, 28 viii 1960), Tay (Black Wd., Rannoch, D.M.Henderson, 20 ix 1960; Perthshire, D.M.Henderson, 16 vii 1962), Dee (Aberdenshire Lumphanan, M.Wilson ?, ix 1922). As Puccinia millefolii, Britain, prob. Clyde D.H.Boyd, ix 1926, no further data, Solway (New Galloway, M.Wilson?, ix 1931). Known from Harris and Kintyre. A collection on Artemisia maritima from Forth (Tyninghame, autumn 1950) had spores agreeing with this taxon. On Aster tripolium as Puccinia tripolii Forth (III Aberlady, M. Wilson ix 1921  & vi 1932 and Miss Miller, 7 vi 1924; O/III, East Lothian Aberlady, D.M.Henderson, 24 v 1953; Tynninghame, D.M.Henderson, 3 ix 1960), Tay (St.Andrews Eden Sanctuary, A.Angus, 20 vii 1965). Known from Mull on this host. On Cirsium heterophyllum Tay (Angus Glen Isla, D.M.Henderson, 19 ix 1963; Kindrogan, B.Coppins, 25 viii 1977 & D.M.Henderson 25 ix 1964; Glen Clova, A.Angus, 28 viii 1976; Glen Tilt, R.Watling, 24 vii 1994), Ross (Kishorn, D.M.Henderson, 6 ix 1952), Moray (Rothiemurchus Inshriach, D.M.Henderson, 23 ix 1959; Aviemore, M.Wilson?, ix 1927). Also known from Mull, P.W.James, 6 ix 1968. An old collection as Puccinia andersoni, Scotland, M.Anderson – is this from Glen Ogle noted by Grove? Also same rust Argyll (Taynuilt, Pass of Bronder, M.Wilson?, viii 1926) and Tay (Glen Lyon, D.M.Henderson, 23 vi 1951). Known from Raasay, Skye and Mull. On Cirsium palustris Forth (III Fife, R.Watling, 5 ix 1978; III, Midlothian Dalmahoy, D.M.Henderson, 1 viii 1954), Solway (Thornhill, poss. N.L. Alcock, ix 1924), Tay (Glen Fincastle, D.M.Henderson, 14 vii 1968). Known from Gigha and Arran on same host. II/III As Puccinia lemonnieriana, Forth (Midlothian Balerno, D.M.Henderson, 2 vi 1952), Clyde (Arran Lag, M.Wilson 1934). As P. tropolii on Aster Forth (Aberlady, 14 vi 1934) and Tay (West Sands, St Andrews, 17 ix 1936). As P. andersoni with short decsiption in Stevenson originally described from Tay (Den of Airlie, Gardiner) and later noted from Moray (Falls of Noran, Glen Ogle) and Tay (Glen Tilt). Also as P. tripoli in Stevenson from Dee (New Pitsligo, Rev. Ferguson).

TFTaDMSSoCAR P. commutata On Valeriana officinalis Argyll (Loch Seil, D.M.Henderson, 3 viii 1954 & I/III 20 vii 1984; Argyllshire, D.M.Henderson, 9 viii 1984) and Forth (Lake of Monteith, D.M.Henderson, 12 ix 1959). Known from Mull.

TFTaDMSSoCAR P. conii On Conium maculatum Tay (Montrose, U.Duncan, vii 1944; II Kenly Den, J.A.Macdonald, 20 iv 1940), Dee (Loch Skene, Prof.Braid, 4 ix 1961; Stonehaven Dunnottar Castle, 4 ix 1961), Forth (Aberlady, D.M.Henderson, 11 viii 1951; Tyninghame, J.A.Macdonald, 9 vii 1938 & D.M.Henderson, 3 ix 1960; Cockburnspath, M.Wilson?, vii 1922; Pittenweem, M.Wilson, vii & viii 1919), Sutherland (Durness, R.W.G.Dennis, 29 viii 1944).

Included here is Puccinia silai on Silaus pratensis from Forth (Dalkeith Caulsland, J.Knapp, viii 1836). This species in Stevenson is recorded from Tweed, Tay and Dee.

TFTaDMSSoCAR P. coronata Recorded in Foister (1958) Economic Plant diseases in Scotland on Agrostis tenuis, Anthoxanthum, Avena, Lolium, Dactylis and Deschampsia. Recorded as such from Tay (Menmuir, Rev. Anderson) in Stevenson. f. sp. agrostis On Agrostis canina from Creagan an Lochan, I.Jorstad, 17 ix 1959 and Clyde (II Benmore Gdn.,D.M.Henderson, 15 ix 1959; Rossdhu, Loch Lomond,I.Jordstad, 11 ix 1959). On A. stolonifera from Forth (East Lothian Tynninghame, D.M.Henderson, 8 ix 1959), Clyde (II Loch Lomond, D.M.Henderson, 11 ix 1959; II Clyde Valley, M.Wilson, ix 1933), Tay (II St Andrews x 1946) and Orkney, Miss Shearer, viii 1930, additionally on var palustris from Tay (Rescobie Loch near Forfar, U.Duncan, viii 1944). On A. tenuis Tay (Tentsmuir, M.Stewart, 19 viii 1966; Loch Tay, D.M.Henderson, 4 ix 1961), Moray (Culbin, D.M.Henderson, 7 ix 1975) Clyde (II/III Balloch Park, Loch Lomond, D.M.Henderson, 11 ix 1959; II Benmore Gdn., D.M.Henderson, 15 ix 1959) and Argyll (Tayniult, M.Wilson, vii 1921). On Argostis spp. Forth (Linlithgow, D.M.Henderson, 27 ix 1959; II/III Pentlandfield, D.M.Henderson, 30 ix 1959), Moray (Culbin Forest, A.Angus, 8 ix 1975). Records from Lewis, Barra, Vatersay, Mingulay, Tiree, Coll, Raasay, Skye, Rhum, and Islay; also Shetland and Orkney incl. Papa Westray. f. sp. alopecuri on Alopecurus pratensis. Record from Skye. f. sp. arrenatheri As Puccinia lolii on Arrenatherum elatius from Forth (II/III North Berwick, M.Wilson, ix 1929; Pathhead, G.B.Walllace, 1924), also Stove near Model Farm, R.Watling, 18 viii 1993 & Arley, M.Wilson, viii 1928), Clyde (Arran Kilmory, M.Wilson, viii 1927) and Tay (Lady Brae nursery, St Andrews 8 vii 1939). Records from Tiree, Raasay, Skye, Colonsay, Garvelloch, Lismore, Gigha, Islay and Kintyre; also Orkney, incl. Papa Westray. f. sp. avenae On Avena sativa (Forth (II Pathhead, G. Wallace, 6 ix 1925 & II A.P.Bennell, 1 viii 1979; Barnton, M.Wilson,ix 1922), Solway (Lockerbie district, R.W.G.Dennis, 6 viii 1943; II/III Gatehouse of Fleet, M.Wilson,  ix 1931) Clyde (Ayr, N.L.Alcock, ix 1926) and Tay (Arbroath Redhead, U.Duncan, x 1944). Record from Rhum and Shetland. f. sp. festucae On F. altissima Clyde (Chatelherault). On Festuca elatior Forth (II Burntisland, M.Wilson, ix 1929), Dee (II/III Brechin, M.Wilson, xi 1933) and Tay (Arbroath, Castle Sea, U.Duncan, viii 1944). On F. gigantea Forth (II Roslin, D.M.Henderson, 20 x 1968,xi 1933; II  Balerno Water of Leith, A.P. & F.M.Bennell, 16 viii 1978) and Clyde (Hamilton, Wilson & Henderson). On F.pratensis Tweed (Coldingham, M.Wilson, ix 1930; II/III; Cove, M.Wilson, vii 1930). On F.rubra Ross (II Lochcarron, D.M.Henderson, 7 viii 1975). On Festuca sp, Forth (Stirling University, D.M.Henderson, 6 x 1968). Record from Orkney.

f. sp. holci On Holcus lanatus Tay (Tentsmuir, M.Stewart, 19 viii 1966), Forth (RBGdn., Edinb. in pot, D.M. Henderson, 12 v 1966), Clyde (Arran Kilmory, M.Wilson). Also known from Mingulay, Raasay and Mull.

On H. mollis Forth (Tyninghame, D.M.Henderson, 8 ix 1959), Dee (II/III, Aberdeen, D.M.Henderson, 9 ix 1975). Records from Tiree, Mull, Arran and Kintyre; also Orkney and Shetland. f. sp. lolii Included here are records of P. lolii on Lolium spp.. On Lolium perenne from Clyde (Ayr, Alloway, N.L.Alcock ix 1926), Tay (II St.Andrews, M.Stewart, 17 vii 1966). On L. italicum Tay (Lady Breas nursery, St Andrews, 18 ix 1930). On Lolium sp, Ayrshire Auchincruive, D.G.O'Brien, x 1931). Known from Skye, Arran and Islay. Reported on Perrennial Ryegrass in North East in 1957 (Foister, 1958). No hosts given: Puccinia coronata. Solway (Kirkcudbrightshire Haugh of Urr, R.W.G.Dennis, 11 viii 1943), Tay (Kinnessburn, St.Andrews, J.A.Macdonald, x 1946; II St.Andrews Boarhills, J.A.Macdonald, 22 ix 1939 and II/III Cameron, J.A.Macdonald, 22 x 1958). Also recorded from Orkney incl. Papa Westray and Shetland. P. coronata & P. lolii on Dactylis glomerata from Clyde (Ayr Alloway, N.L.Alcock,ix 1926), Tay (Boarhills, 22 ix 1939; East cliffs, St.Andrews, J.A.Macdonald, 8 x 1936). On Descampsia caespitosa Forth (Kirknewton, D.M.Henderson, 4 x 1959).

TFTaDMSSoCAR P. crepidicola On Crepis capillaris Tay (Boarhills, 4 ix 1984) and from Colonsay and Islay.

On C. virens Tweed (Kelso? No further data), Tay (Cairn, Fife, 30 vii 1940, R.W.G.Dennis; Ballanluig, viii 1915) and from Kilmory, Arran II, 1927 & as P. crepidis Wilson 1934.

TFTaDMSSoCAR P. cyani O, I, III Forth (St Michael’s, Fife vii 1936), Dee (Dinnett, Aberdeenshire, 19 vi 1961) and Tay (Leuchars, A. Pratt, vii 1936). Noted on cultivated Centaurea cyaneum (Cornflower) by Foister (1958). Puccinia suaveolens var. centaurae is a synonym.

TFTaDMSSoCAR P. difformis On Gallium aparine III Glenforsa, Mull and Sutherland (Thurso, I & II, no further data). Records in Stevenson as Aecidium galii for Tay may belong here and Moray in Add. 8.  See P. punctata below.

TFTaDMSSoCAR P. dioicae var. dioicae O/I on Cirsium dissectum & C. palustre; II/III on Carex dioica. Records from Mull and Dee (Cork Brdige, Aberdeen, 26 vi 1963 along with Carex dioica), Forth (Lumphanan, Dunfermline, 21 vii 1922, M.Wilson), Tay (Ben Lawers 487m., 24 vi 1952; Tentsmuir 18 v 1989, R.Watling), Glenfincastle, 6 vii 1968; Killin, 732m., 28 vi 1964), Moray (Loch Duntelchaig, 1936, JA Mcdonald), Hoy Orkney all on C. palustre.

III On Carex demissa from Rhum. On Carex dioica Tay (Killin, 732m., 28 vi 1964 D.M.Henderson; Creag an Lochan, Ben Lawers, 17 ix 1959, D.M.Henderson), Argyll (Ben Lui, 366m., no date given )and Orkney Halcro Johnson, vi 1928) and Mull (Loch Spelve, ix 1968) and Skye. On C. flava Ross (Gairloch, viii 1934) and Mull Loch na Keal, 7 ix 1968. On C. incurva Moray (Elgin, II & III, G.Gordon as P. schoeleriana. No further details). On Carex lasiocarpa Tay (Creag an Lochan, Ben Lawers, 17 ix 1959, D.M.Henderson). On Carex lepidocarpa Tay (Creag an Lochan, Ben Lawers, 17 ix 1959, D.M.Henderson along with Parnassia), Argylle (Glencoe, 24 ix 1960), Ross (Strome Ferry, 17 x 1959) and Westray Orkney, viii 1913). On C. ovalis Barra (Edin. Univ. Exp. Vii-viii 1936), Forth (as Carex leporina, Gifford, Hope. Res., viii 1946), Argyll (Glen Lochay, 229m., 1 ix 1961, D.M.Henderson), Rhum and Piandrum J.Kirk no further data.

TFTaDMSSoCAR Var. schoeleriana on Carex arenaria Forth (Gullane, 20 ix 1915; Aberlady, 12 ix 1953, D.M.Henderson), Tay (Dunes, St Andrews, 18 vii 1946; Tentsmuir, 7 vi 1938; Guardbridge, St Andrews, 12 vii 1938).

On C. incurva Elgin, G.Gordon II & III. On C. maritima Moray. On Senecio jacobea Forth (Gullane, v 1931, M. Hering & 25 v 1933; Largo, Fife, 17 vi 1954; Aberdlady, 14 vi 1934 & vi 1936, J.A.Mcdonald), Tay (Dunes, St Andrews, 21 v 1938; Tentsmuir, 7 vi 1966, M. Stewart; St Andrews, v 1937, nr. Carnoustie, v 1845; North of Montrose, 15 iv 1946), Orkney, G.Hadley 3 vi 1963. Also two old collections in E ‘Burntisland Fifeshire, 1821 june-sept.’ as Aecidium jacobaea both prob. Greville’s collections. As Aecidium compositarum var. jacobaeae on Senecio jacobaea from Forth and Tay in Stevenson.

TFTaDMSSoCAR Var. sylvatica II/III on Carex capillaris. Glen Doll, 457m., 15 vii 1944, R.W.G.Dennis as Puccinia sylvatica Sutherland (Clashnatoll, B. Flannigan, vii 1955). On Taraxacum offinale agg. Tweed (Galshiels, v 1928), Tay (Ben Lawers, 657m., 28 vi 1950 redet. for P. taraxaci). Tay (Tentsmuir, 7 vi 1938; Glen Lyon, 20 vi 1936) and Forth (Between Gullane & Aberlady, 2 vi 1934). On T. palustre Tay (Ptarmachan, 762m., 10 vii 1965, D.M.Henderson).

On Taraxacum spectabile Tay (Ptarmachan 762m., 10 vii 1965, D.M.Henderson with C. dioica). On Taraxacaum sp. Tay (Ben Lawers, 2 vii 1964 with C. dioica; Killin, 732m., 28 vi 1964, D.M.Henderson) and records from Raasay and Orkney.

TFTaDMSSoCAR P.distincta On Bellis perennis cultivars. Only known from Forth (Blinkbonny Edinb. 12 viii & 13 x 2004, ix 2006 & viii 2007) but spreading from England; should be looked out for in gardens. Also with some doubt recorded from Western Ises. No further data.

TFTaDMSSoCAR P. elymii On Elymus arenarius Tay (St Andrews xi 1924, N.L.Alcock & 25 x 1965, J.A.Mcdonald) and Moray (Nairn, 23 ix 1955, D.M.Henderson).

TFTaDMSSoCAR P. epilobii On Epilobium anagallidifloium Tay (Rait, 3 v 2006), Ross (Beinn Eighe 91.4m., 19 vii 1955; Ben Wyvis, 8 ix 1963). On E. montanum Forth (Edinburgh, vii 1937; Balerno, 31 v 1959) and Tay (Ben Lawers 24 vi 1950; Glen Lyon, 13 ix 1960), Mull and Skye. On E. obscurum Moray (Kinguisse, vi 1922). On E. plaustre Tay (Blairgowrioe, 2 viii 1949; Glenbrantner vii 1929; Glen Shee, 7 viii 1955), Ross (Gairloch, vii 1936), Orkney, Mull and Skye. On Epilobium sp. Forth (S. Medium, West Lothian, 31 v 1959). In Stevenson recorded from Tweed, Forth, Tay & Dee. Collections in E on Epilobium palustre Duddingston Loch, May/June 1821 and E. obscurum ? Edinb. (III) probably both Greville material.

TFTaDMSSoCAR P. eriophori 0/I on Solidago vigaurea Argyll (Ben Nevis, 305m., 15 ix 1956 growing with Scirpus; Ben Vair, Appin, 914m., 4 ix 1941). II/III on Trichophorum caespitosum subsp. germanicum. Known from Harris and Ross (Kishorn as Puccinia confinis 457m., 10 viii 1959, D.M.Henderson).

TFTaDMSSoCAR P, eutremae On Cochlearia alpina/danica. Known only from Scotland but no material in E.

TFTaDMSSoCAR P. fergussonii On Viola palustris Moray (Ballendalloch, Bannf., vii 1936, D.F.Mckenzie), Tay (Creag an Lochan, Ben Lawers, 762m., 8 viii 1958; Stormont Loch, Blairgowrie, 26 viii 1963; Glen Brantner, vii 1929; Glenfincastle, Pitlochry, 14 vii 1948 & 14 vi 1970), Forth (Dunblane, vii 1930; Lumphanan, Dunfermline, vii 1922), Solway (Sandhead, Wigton vi 1930) and Mull. Also from Ben Cruachan 610m., 27 vii 1921. Also record nr Douglas 24 vii 1940. Introduced with short description in Stevenson from Tay (New Pitsligo, Rev. Ferguson type locality), Dee and Moray. Also old collection in E looks like Greville material.

TFTaDMSSoCAR P. festucae Foister (1958) considers this species to be common on commercial crops of Festuca in the S.West & N. East. On Festuca arenaria Argyll (nr. Oban, vi 1930, Rupert Smith). On F. ovina Forth (Swanston Fm., Pentlands, 28 vi 1925) and Argyll (Ben Lui, 457m., 28 vi 1925 & 16 ix 1959). As Uredo festucae on Festuca ovina var. tenuifolia Argyll (Loch Snere, 25 vf 1963, D.M.Henderson). II on F. rubra Harria, Barra, Coll, Skye, Rhum, Islay, and Islay. On F .rubra subsp. rubra Forth (Tyninghame, E.Lothian, 8 ix 1959, D.M.Henderson). On F. rubra var. fallax Forth (East Craigs, 5 iii 1954, M.Noble). On Festuca sp. Forth (Dalry, Edinb., ix 1931), Clyde (Carnwath, ix 1929) and Tweed (Peebles, U. Duncan, 3 x 1946),Tay (Ben Lawers, 616m., 17 ix 1959). I on Lonicera periclymenum Argyll (Loch Suneen, 25 v 1963; Central Argyllshire, 10 vii 1942, M.Noble), Tay (Ceres, 5 vii 1944, R.W.G. Dennis; Parkhill, Arbroath, vii 1948, U.Duncan), Forth (Gifford, 14 vi 1954; Pencaitland, vii 1931) and Moray (Forres, 29 vi 1963; Fochaber, Elgin, vi 1943, RWG Dennis). Also on Falls of Orion 29 vi 1934, Barra Cripg on Mull and Gigha; record from Shetland. As Aecidiuum crassum var. periclymeni on Lonicera in Stevenson from all Watersheds except Sutherland, Solway and Ross.

TFTaDMSSoCAR P. galii-verni On Cruciata laevipes Tay (Ballinluig, shingle islands, 8 viii 2009, M. Clarkson). As Gallium cruciatum Forth (Gifford, vii 1945; Borthwick Castle, viii 1930; Aberlady, II & III, 24 viii 1952), Tay (Perth, Buchanan White; Dunkeld, 23 viii 1953; Kenley Den, St Andrews, v 1937; Den of Airlie, xi 1952 U. Duncan). Also Lagieret, III 17 ix 1965. Further collections under Puccinia valantiae Clyde (Hamilton, 14 ix 1959) and Tay (Kenley Den, St Andrews, 8 v 1937). On G. saxatile (= G. hercynium) Moray (Loch Einich, 640m., 29 viii 1955), Ross (Beinn Eighe, 457m., 17 vii 1955), Solway (Whitecombe, Moffat 457m., 25 vii 1957), Tay (Glen Lyon, 17 vi 1936; Ben Lawers, 24 vi 1950; Creag an Lochan, Ben Lawers, 609m., 17 ix 1959), Argyll (Ben Ledi, 14 v 1950 & 24 v 1951) and Forth (Balerno, Edinb., 4 vi 1921). Known from Harris, Raasay, Skye, Colonsay, Jura, Luing, Islay, Arran, Gigha and Kintyre; also Shetland and Orkney. A P. celakovskyiana about Edinb. Sept. no further data in E probably Greville collection. Recorded in Stevenson as P. valantiae on Gallium cruciatum from Tweed, Forth & Tay and record of P. acuminata from Tay (Rannoch, Buchanan White) introduced with short description in Stevenson and under same name in Add. 8 for Dee.

TFTaDMSSoCAR P. gentianae On Gentiana acaulis Tay (Perth no further data). Noted by Foister (1958) on nursery stock of Gentiana acaulis.

TFTaDMSSoCAR P. glechomatis On Nepeta hederacea Solway (Gatehouse of Fleet, x 1950; Mossguard, Newton Stuart, ix 1929), Tay (St Andrews, 8 x 1936; Den of Airlie, x 1952, U.Duncan; East Cliff St Andrews, 12 i 1936; Raith, ii 1934), Moray (Nairn, 25 ix 1955), Forth (Eskbank, 12 iv 1953; Dalmeny, 6 x 1968; Aberlady, 11 x 1952) and from Skye. Noted from Tay in Stevenson.

TFTaDMSSoCAR P. glomerata On Senecio aquaticus Forth (Duddingston, Edinb., v 1921; Thruipmuir, Balerno, 16 vi 1965), Shetland and Orkney. On S. jacobea Tweed (Dawyck), Tay (St Andrews, v 1937, JA Mcdonald; Taynuilt, vii 1921; Kenmore, no further data; Tentsmuir, x 1930, NL Alcock; Murthley, ix 1925 N.L.Alcock; Glen Lyon, 31 viii 1950), Sutherland (Cape Wrath, viii 1931), Clyde (Milngarvie, 14 vcii 1933, K.Braid), Moray (Beauly, 1925, no further data), Dee (Bridge of Don, viii 1962), Forth (Burntisland, 14 v 1921, JA Mathews; Edinburgh, 25 ii 1954, AE Boyd; Aberlady, vii 1936, J.A.Mcdonald; Gullane, vi 1929; Roslin, ix 1926; Loch Leven, 3 vi 1972, R.Watling) Known from Harris, North Uist, Skye, Rhum, Eigg, and Colonsay; also Shetland and Orkney. As P. expansa Tay (St Andrews Dunes, 21 v 1938), Forth (Flanders Moss, vi 1959 as P. scholeriana, D.M.Henderson) and Islay 30 v 1944. Collection from Roslin, Edinburgh by Greville in E poss. type material but no further data in addition to specimens on S. jacobea ? Greville.

On S. jacobaea recorded in Stevenson from only eastern Watersheds, excluding Sutherland and as P. senecionis on S. aquatica from Tay (Fern, Noran Wd, Rev. Anderson).

TFTaDMSSoCAR P. graminis On Agropyron repens Tay (Balbeggie, Perthshire, 6 xi 1969; Alyth, 6 xi 1969; St Andrews, 7 x 1939 & 9 x 1947) and Forth (Pathhead, 12 viii 1955, M.Noble; Haddington, viii 1943, R.W.G.Dennis; Blinkbonny, Edinb., 27 1 1992;  Aberlady, 15 iv 2006), Dee (North Water bridge, Kincardineshire, 23 viii 1961; Moray (Boat of Garten, ix 1945, M.Noble). and Solway (Dumfries, viii 1924). On Agrostis canina Argyll (Benmore  Gdn.,15 ix 1959). On A.tenuis Tay (Alyth, xi 1928). On A. vulgaris Argyll (Tyndrum, ix 1921). On Agrostis sp. Moray (Rothiemurchus, 23 ix 1959; Inshriach, 20 ix 1965). On Alopecurus pratensis Solway (Thornhill, no further data).

On Anthoxanthum odoratum known from Skye. On Briza media Ttay (Glenshee, viii 1940). On Arrhenatherum ealtius Forth (Pathhead, 10 ix 1961). On Avena sativa ‘oats’ Tweed (Dingleton, Melrose, viii 1942 R.W.G.Dennis ), Dee (North Water Bridge, Kincardineshire, 23 vii 1961), Sutherland (Caithness, ix 1976. Also simply collection noted on oats Clyde (Kilmains Aryshire ix 1924) and ‘viii 1933’ no further data). On A. strigosa Moray (Boat of Garten, ix 1945, M.Noble). On Dactylis glomerata Kilmory, Arran, no date; Forth (Crichton, 15 v 1955). On Festuca elatior Clyde (Alloway, Ayr). On Hordeum vulgare viii 1955, M.Noble, no further data, Dee (North Water Bridge, Kincardineshire, 23 viii 1961, G.Hadley). Collection on ‘barley’ Forth (Pathhead, 12 viii 1955, M.Noble)

On Triticum ‘wheat’  Tay (nr. Arbroath, x 1944, U.Duncan) and old collection in E labelled on wheat, A.Jerdon presumably from Jedburgh area - Tweed. Noted in Foister (1958) on Agropyron repens which acts as a source for forage grasses and cereals, Festuca rubra, Holcus lanatus, Lolium perrne, Avena sativa, Poa trivialis, Triticium aestivum & T. turgidum, Deschampsia, caespitosa, Dactylis glomerata, Agrostis tenuis, Bromus braeziformis & B. inermis, Secale cereale and Alopecurus pratensis and Arrenatherum elatius Also in Foister (1958) as P. avenae on Avena sativa recorded fro S.East, SW and S. Central severe in S East in 1949, in N east  & Orkney in 1950, in S East and S Central N. East and Orkney in 1951 and in S.West in 1955. As var. secalis  on Secales cereale recorded from S.West and Central N.East. Also records from Longforgan 12 viii 1936 (no host given) & Moray (Bridge of Moarnoch, Huntley, 7 x 1942, Dr. Clouston). O on Berberis vulgaris Kirkwall, Orkney, Moray (Rothiemurchus, 23 ix 1959; Glen Lyon, viii 1919), Forth (Aberlady, 10 vi 1939; Gifford, 13 vii 1972; Haddington, viii 1943, RWG.Dennis; Greenlaw, 7 v 1944, RWG Dennis; Largo, Fife, 17 vi 1954) ), Tweed (St Boswells, 13 vii 1941, RWG Dennis), Tweed (Melrose, 2 vii 1950), Solway (Dumfries, viii 1924; Thornhill, N.L.Alcock, ix 1924; ) and Tay (Arbroath, vii 1953, U.Duncan). Old collections as Aecidium berberidis in E on Berberis from Forth (Dalmeny, M.C.Cooke viii 1871), a collection by Sir W. Jardine no further data and a probable Greville collection labelled ‘Edin.’ Also old collection labelled Botanic garden on Berberis – probably Edinburgh. On Mahonia bealei Argyll (Benmore Gdn., v 1943). On Mahonia sp. Forth (Crichton, vi 1936). Recorded by Stevenson for Tweed, Forth, Tay, Dee, Clyde and Ross ‘on leaves of grasses and corn’ – probably refers to several species. Also recorded as Trichobasis linearis in Stevenson with description from all Watersheds except Moray and Sutherland, Solway and Argyle. As Aecidium berberidis in Stevenson from all Watersheds except Sutherland, Solway, Argylle and Ross.

TFTaDMSSoCAR Var. graminicola As P phlei-pratensis Clyde (on Scottish, N.American, and Swedish cultivars of Phleum Alloway, Ayr, ix 1926, N.L.Alcock; also Saxony Avena sativa N.L.Alcock). As variety of P. graminis occasionally severe in South and East central Scotland on Phleum according to Foister (1958); also recorded on Festuca rubra. On Anthoxanthum odoratum as P. anthoxanthi Dee (Craibstone, Aberdeen, x 1921 & x 1942 and Solway (Gatehouse of Fleet, ix 1931).

TFTaDMSSoCAR P. heraclei On Heracleum sphondylium Tay (Killin, vi 1922 & 9 vi 1942; Loch Tay Lawers, 13 viii 1921), Argyll (Drumnadrochit, iv 1943, U. Duncan), Forth (Bridge of Allan vi 1936; Largo, Fife, 18 vi 1954; Pittenweem viii 1919), Sutherland (Cape Wrath, viii 1931; Reay Caithness, 11 vi 1966) Clyde (Glen Buck 29 vi 1941 M.Noble). Also known from Skye and Iona; also Skara Brae Orkney and South Ronaldsay. Old collections in E include material from Mrs Boswell Balmutto prob. Greville collection, holotype from nr. Kirkcaldy vi 1821 and one other collection. In Stevenson recorded from Forth, Tay, Dee, Moray and Argylle.

TFTaDMSSoCAR P.hieracii On Centuarea nigra Tay (Boarhills, 10 x 1969, A.Angus; Blairgowrie, 2 ix 1947 - ssp. nemoralis; Banks of Tay, 18 ix 1959), Clyde (Cadzow Pk., Hamilton, 14 ix 1959) and Forth (Queensferry, 26 viii 1973; Between Hermiston & Ratho on canal bank, 20 ix 1959). On Cirsium heterophyllum 3 ix 1968, no further data.

On Hieracium alpinum Tay (Tarmachan, 9 x 1919) and Argyll (Ben Lui 10 x 1914 & 28 viii 1953). On H. boreale Forth (Barnton, Edinb., ix 1992). On H. peleterianum Tay (Creagg an Lochen, Ben Lawers, 17 vii 1954, D.M.henderson). On H. lawsonii Forth (between Pettycurr & Burntisland July 1821, prob. Greville collection. On H. murorum Tay (Tentsmuir, 26 ix 1937) and Moray (Loch Duntelchaig, 1936) and Tay (Ben y Vrackie, 19 vii 1938 as var. nigrescens). On H. prenanthoides Forth (Roslin, vi 1888). On H.sylvaticum Forth (Roslin July 1821 , prob. Greville collection). On H.subalpinum Argyll (Ben Lui, 9 vii 1995, R.Watling). On Hieracium  spp. Ross (Beinn Eighe, 426m., 19 ix 1955), Dee (Maryculter, 27 x 1966), Tay (Blair Atholl, 274m., date)  and Argylle (Ben Lui 9 vii 1995, R.Watling). Known from Raasay, Skye Croig, Mull, Rhum & Eigg – Edinb. Univ. Exped. vii-viii 1949.

On Leontodon autumnalis Moray (Loch Einich, Aviemore, 305m., 29 ix 1955), Forth (Pittenweem, viii 1920), Tweed (Berwick, ix 1922), Solway (Kintyre, viii 1936; North Uist, Vatersay, Coll, Skye, Mull, Iona, Islay, Arran and Kintrye; also from Stromness Orkney. Material in E as ‘Uredo on Leontodon taraxacoides Edinb. June 1821’ prob. Greville collection as is collection on same host labelled ‘Uredo cichoracearum summar & autumn 1821’ and as ‘Uredo cichoracearum summer 1821’ are probably referred to L. autumnalis; also collections on L. taraxacoides Glen Stocking shore, Colvend, 20 ix 1872 & on L. vulgaris Forth (Longniddry, pot plant 20 x 1955). On Taraxacum officinale Argyll (Fort William, 28 ix 9185; Inverliever, vii 1925), Forth (Dirleton, 7 viii 1986, A.P.Bennell), Solway (Carradale, Kintyre, viii 1931), Tay (Lethan Grange, Arbroath, x 1944, U. Duncan), Sutherland (Tongue, viii 1931) and Torloisk, Mull. On Taraxacum sp. Dee (Linn of Dee, 23 viii 1968; Durris, Kincardineshire) and Tay (Largo, Fife, 15 vi 1954), Forth (Stirling University, 6 x 1968; also Harris, Vatersay, Mingulay, Skye, Rhum, Mull, Islay, Arran and Kintyre. As P. taraxaci Tay (Largo Rd., St Andrews, 11 x 1935; subsp. erythropsepermum; East Sands, St Andrews, 15 v 1937; East Sands, St Andrews, 23 v 1936; West Sands, St Anbdrews, 6 ix 1936; subsp. erthythrospermum, Tentsmuir, 23 v 1960); also known from Shetland and Orkney. Two Greville collections in E unspecified.

More than likely as Aecidium compositarum var. prenanthis on Hieracium in Stevenson for Tweed, Forth, Tay and Dee. As Puccinia tinctoriae on Seeatula tinctoria Tay (St Andrews Bot. Gdn., 11 ix & xi 1980, J.A.Mcdonald).

TFTaDMSSoCAR var. hypochaeridis This variety was the subject of intensive study by D. Scott and T. Jenkins New Zealand Dept. Agric. as possible control organism for the host in N.Zealand. On Hypochaeridis radicata Solway (Glen Trool, 25 v 1986), Moray (Loch Duntelchaig, 19 viii 1936, 23 v 1937 & 21-340 vii 1939, J.A.Mcdonald; Dalwhinnie, 8 vi 1986; nr. Braemore, Inverness, 24 vi 1984; Glenstrathfarrer, 25 ix 1985, A.P.Bennell as P. hieracii), Argyll (Loch Eck, 14 vi 1951; Ardnamarchan, 6 vi 1961), Forth (Aberlady, 10 iv 1986; East Criags, 26 vii 1939 & 2 vii 1941; Edinb., viii 1921), Tay (Ballinluig, vii 1915; Dunes, St Andrews, 6 ix 1936 & 15 v 1937, JA Mcdonald), Ross (Plockton, 20 vi 1955) and Dee (Glen Tannar, 5 ix 1975). Records from Lewis, Harris, Mull, Luing, Arran and Gigha; also Univ. Exped. Vii-viii 1949 Eigg and Orkney

TFTaDMSSoCAR Var. piloselloidarum On Pilosella officinarum Argyll (Loch Scamadale, 20 vii 1983), Forth (Largo, 16 vi 1954; Cammo, Edinb., 3 vi 1985; Turnhouse vi 1985, D.McKean; Dollar Glen, 10 vii 1985), Moray (Glen Strathfarrer, 25 & 26 ix 1985, Ray Woods), Tay (Liunan Bay, Angus, vi 1944, U.Duncan; Tentsmuir, 10 x 1965; Blairgowrie 20 vii 1952 and Glen Tilt, 26 vii 1942, R.W.G.Dennis  both as P. hieracii; Blairgowrtie, 28 v 1955; Pitlochry on subsp. trichosoma, 17 vi 1985), Moray (Black isle vi 1942 U.Duncan0) and Argyll (LochEtive, 24 iv 1964). D. Scott inoculation exp. 1986 - Collections from Moray (Loch Laggan, 8 vi; Spean bridge, 8 vi; Dalwhinnie, 8 vi), Solway (Ballantrae 27 v; Mull of Galloway, 26 v; Torres Warren, 25 v; Glen Trool, 24 v ), Argyll (Fort William, 7 vi; Forth (Glen Dollar, early 1986; Arthur’s seat Edinb., 8 iv;  Stirling 7 vi; Turnhouse, Edinb., 15 v Dolphinston, 23 v; Kirkliston, 30 v), Tay (Tentsmuir, 9 x 1965 A.Angus; Strathyre, 7 vi; River garry, early 1986; St Andrews, 10 v; Kirriemuir, 8 vi; Leuchars 10 v; Blairgowrie 10 v; Arbroath, 20 iv), Moray (Inverness, early 1986), Clyde (Glen Buck 27 v 1986) and Tweed (eebles 27 iv; Selkirk, 27 iv; Galashiels 27 iv; Jedburgh, 27 iv).

TFTaDMSSoCAR P. hordei As P. holcina on Holcus lanatus Tay (Ballinluig, viii 1915; nr. Arbroath, ix 1944 & Parkhill, Arbroath, vii 1944 U. Duncan), Tweed (Broughton, ix 1943; Coldingham, vi 1935), Forth (Crichton, 6 ix 1945; Barnton, ix 1922; Balerno, 23 iv 1961; Arniston Engine, 23 vi 1983; RBGdn., Edinb. 13 vii 1954), Clyde (Medwin, Carnwath, 6 viii 1952) and Moray (Loch Duntelchaig, 29 viii 1936 & viii 1946). Also Greville collection in E as ‘Uredo lanatus common, August’. On H. mollis Tay (Loch Tay 1 ix 1961; Kippen Cardross, 12 ix 1959; Pitlochry, 13 vii 1959; Killiecrankie, 24 ix 1959; Brechin, 21 vi 1966), Sutherland (Lochshin, Larig, 21 viii `1944; Sangobeg, Durness, 30 viii 1944, RWGDennis), Clyde (Cadzow, 19 ix 1959; nr. Hamilton, vii 1930), Forth (Linlithgow, 27 ix 1959; Pentlands, 12 viii 1959; Kirknewton, 10 x 1959; Corstorphine, 22 vii 1952; Penicuik, 20 ix 1959). Also records from Salen Mull and Eigg Edinb. Exp. Vii-viii 1949. On leaves of Hordeum murinum (as P. glumarum) Forth (Whitehall,  Midlothian, 16 ix 1924, G.Wallace; Pathhead, 14 x 1923), Tweed (Berwick, x 1922 M.Y.Orr.

On H. sativum ‘barley’ Forth (Pathhead, 6 ix 1925), and Tweed (Coldingham, viii 1930) and Orkney. As P. simplex Tay (Kinkell Brae, St Andrews, x 1946 on cultivated barley. As P. dispersa on Hordeum vulgare Forth (misident.collection vii 1975). Brown rust recorded as of some concern in E.Central and S.East 1947, 1952 & 1957.

TFTaDMSSoCAR P. horiana Collections only know from Forth (East Craigs, 26 & 27 x 1976) on cultivars Full Gold, Rivalry, Camandra and Mary Shoesmith).

TFTaDMSSoCAR P. hydrocotyles On Hydrocotyle vulgaris from Clyde (Kilwinning, Aryshire, D.A.Boyd), Ulva ferry Mull, 11 ix 1968 & Moray (Loch Duntelchaig 23 v 1937, JA Mcdonald).

TFTaDMSSoCAR P. hysterium On Tragopogon. Most records as Puccinia tragopogoni. Known from Forth (between Kirkliston and Ratho, Edinb., v 1933; Barns Ness, E.Lothian, J. Muscott, 4 viii 2007; Corstorphine, 1 vi 1944, 13 vi 1950, 6 vii 1952; Arniston Engine, West Lothian, 22 vii 1977, AP Bennell; Edinb. no further data; Comiston Quarry, 1 vii 1921, Evans; Lasswade, 22 ix 1960, M.Noble & 20 vii 1966) Tay (Broughty Ferry 4 vii 1938; Guardbridge, St Andrews, 18 vii 1938; Inverkillin, Arbroath, vii 1944, U.Duncan; The Links St Andrews, 31 v 1966, A.Angus; Cairns mill, 3 vi 1966, M. Stewart). Old collection in E but no further data and as Aecidium ‘Foxhall, Cpt. Waunch June-July 1821 both possibly Greville. Known from Tay (East Haven, Rev. Ferguson) in Stevenson and in same publication as Aecidium tragopogonis from Tweed & Tay

TFTaDMSSoCAR P. iridis On leaves of Iris germanica in garden, Rousay Orkney. On I. richenbachii Forth (RBGdn., Edinb., Alpine collection, no further data). Recorded first of all in 1924 fide Foister (1958) and found until 1945 on commercial Iris but not seen since.

TFTaDMSSoCAR P. lagenophorae On Senecio squalidus Forth (Blackhall, 1 ii 2005; Ravelston, 13 vi 1983, APBennell; Carricknowe, D.McKean, 18 vi 1985 – all Edinb.). On S. viscosus Forth (Blackhall, 19 vi 2005; Comley Bank, 26 xi 1988 both Edinb.). On S. vulgaris from Forth (RBGdn., Edinb., 8 x 1987; Longniddry, 26 vi 1968; Lufness mill Hse., Edinb., no date; Blinkbonny,1 xii 1983; Ferry Rd.allotments, , 14 viii 1979, APBennell; Orchard Rd.,1 viii 1976; Orchard Brae, 29 xi 1978 – all Edinb.), Tay (West Sands, St Andrews, 18 ix 1973, A.Angus; St Marys, St Andrews, 9 vii 1981, A.Angus), Clyde (Crossford village & L. Nethan). Records from Skye, Mull, Islay, Arran and Kintyre; also Papa Westray, Orkney. Also O on S. cruentum (Cineraria), Clyde x 1980.

TFTaDMSSoCAR P. lapsanae On Lapsana communis Tay (Dunkeld, 29 ix 1985; Bot. Gdn., St Andrews, 30 iii 1949; Blairgowrie, 8 v 1950, 16 v 1951 & 1 vi 1955; Waulkermills, Arbroath, ix 1944, U. Duncan), Dee (Bieldside, Aberdeen, 19 iv 1981), Forth (Bridge of Allan, vi 1936; Corstorphine,Edinb., 31 iii 1943 & 12 xii 1938; Roslin, 14 iv 1961; East Craigs, 12 vi 1953 D.M.Henderson; Cockburn, 7 v 1949 RWG Dennis), Moray (nr. Foyers, viii 1942, U. Duncan), Argyll (Inverleiver) and Clyde (Hamilton High Parks, Lanark, 19 v 1985 & Chatelherault). Also Skye Mull, Arran and Kintyre. Material in E prob Greville, as ‘Uredo cichoracearum on L. communis Carlowrie 1821’.

As Aecidium compositarum var. lapsanae in Stevenson on Lapsana from Tay (Fern, Brechin etc. Rev. Ferguson); Dee added in App.1.

TFTaDMSSoCAR P. liliacearum On Ornithogalum umbellatum. Known from Forth (Pentcaitland, iv 2008 & 23 iii 2009, J. Muscott) and Tay (Blairgowrie, 8 vi 1950; Whitekirk, East Lothian, v 1935) and Tweed (Roxburgh, v 1921). Scarce in Scotland Wilson & Henderson (1966).

TFTaDMSSoCAR P. longissima On Koeleria cristata Tay (Sands of Barry, 4 vii 1898 – taken from Herb. Specimen).

In Stevenson Add 6 as Puccinia paliformis with description recorded from Dee (Links nr., Aberdeen, Prof. Trail).

TFTaDMSSoCAR P. luzulae. On L. campstris Tay (Dunes 15 v 1937 & Morton Locks 18 vii 1938 both St Andrews)

On L. pilosa Mull and Orkney. On L. sylvatica Arran as P. oblongata, Forth (Saltoun, 4 vi 2000). Recorded with short descripiton for Tay (Glamis) and Moray (Aviemore) in Stevenson ‘on leaves of Luzula and also as Trichobasis oblongata from all but Moray, Sutherland, Solway and Argyle; also in App.1 as P. luzulae from Dee and in Add 4 for Argyle.

TFTaDMSSoCAR P. maculosa as Puccinia chondrillae on Mycelis muralis from Clyde (II Younger BGdn, Benmore J.A.Macdonald, 15 ix 1959) and as P. prenanthes Forth (RBGdn, Edinb. Demonsration Gdn., 15 ix 1959) and Moray (Invergarry, Inverness, 28 ix 1985, BJ Coppins).

TFTaDMSSoCAR. P. magnusiana O/I On Ranunculus flammula var. minimus Tay (Boarhills, St Andrews, 10 v 1966, M.Stewart). On R. lingua ‘Duddingston Loch, vi 1821’ Greville? as Aecidium carmicleanaum.  On R. repens Tay (Boarhills, St Andrews, 8 vi 1956; Blairgowrie, 30 vi 1952) and from Barra Campbell, 1936 and poss. Firth, Orkney.

Probably includes a proportion of the records in Stevenson under Aecidium ranunculearum from all Watersheds except Sutherland, Argylle & Ross. II/III On Phalaris communis Tay (Boarhills, St Andrews, 13 x 1965 & 9 x 1955 also inoculation experiments; Kilconquhar Loch, 16 vi 1966; Blairgowrie, viii 1933 & 10 viii 1953; West Newton, nr. Arbroath, viii 1944, U. Duncan; Parkhill nr. Arbroath, U. Duncan; Pitform, viii 1845; Brechin, ix 1926; Montrose, vii 1930), Argyll (Appin, ix 1941), Solway (Wigton 9 x 1955). Also Glen Forsa, Mull, North Uist and Tiree, Harris, Barra, Coll, Skye, Colonsay, Isaly, Gigha, Arran and Kintyre. Also Greville collection in E 1821 as ‘on ‘Arundo’.

Attacked by Eudarluca caricina Mull (10 viii 1953) and Foula

TFTaDMSSoCAR P. major On Crepis paludosa Ross (Glen Kishorn, 20 vi 1957; Lochailhart, W.Ross, 24 viii 1951, D.M.Henderson; Creag an Lochan, 670m., 17 ix 1959, D.M.Henderson), Tay (Falls of Tummel, vi 1944, U. Duncan; Killin, vi 1922, M.Wilson; Glen Doll, Corrie fee, 22 vi 1985; Loch Lee vi 1954 & Glen Artney 27 v 1939), Argyll (Ben Lui, 24 vi 1951; Loch Sinere, 25 v 1963; Tyndrum 22 vi 1958, 274m.; Glen Branter vii 1929, RWG Dennis), Tweed (St Mary’s Loch, ix 1928), Sutherland (Brora Water, 10 vi 1966), Forth (Thruipmuir, Balerno, vi 1953), Clyde (Ben Vaserlich, Loch Lomond, vi 1915), Moray (Daviot, v 1942, U. Duncan; Loch Duntelchaig, 3 ix 1934, viii 1936 & 22 v 1937v) and Solway (Loch Skene Moffat, 610m., 2 vi 1959). Greville collection in E from Miss Boswell collected at Balmutto on Hieracium paludosa. Also Greville collection ‘on Hieracium prenanthes  1821’ is presumably something different.

TFTaDMSSoCAR P. malvacarum On Althea officinalis Forth (RBGdn., Edinb., 20 vii 1978 & 2 viii 1978.

On A. rosea Forth (Gifford, 14 vi 1954; Afton Terrace, Edinb., 20 iii 1979 & 17 viii 1984; Edinb., 1 vii 1931; Linlithgow, 19 v 1985) and Tay (Univ. Gden, St Andrews, 19 vi 1965), and from Arran on Althaea rosea. On hollyhock Forth (Edinburgh, v 1960; East Craigs no further data). On Lavateria arborea Tay  (St Andrews, 11 xi 1935 & Bot.Gdn St Andrews 9 xi 1967). Noted as on commercial L. trimetris by Foister (1958). On Malva rotundifolia Moray (Findhorn, viii 1927) and Eden Golf Course, 29 v 1937. On Malva sylvestris Sutherland (Helmsdale, 22 vii 1962), Tay (St Andrews dunes, 6 x 1948 & links 20 vii 1965; Auchinthe, Angus, vi 1944, U.Duncan; Guardbridge 27 v 1937, JA Mcdonald; Boarhills, 17 vii 1966, M.Stwart), Moray (Aviemore, ix 1927; Kilmur, Black Isle, vi 1943, U.Duncan), Forth (Kinghorn, v 1936; North Berwick, v 1926; Castle Terrace, Edinb., 27 vi 1984 & vii prob. same year;  Tantallion, ix 1929). Also Islay, Port Charlotte. On cv. Primley Blue’ Solway (Logan BGdn., 6 xii 1984). On Malva sp. Arran. On Sidalcea candida Tweed (Rothes, Peebles, 3 xi 1935 JA McDonald). Recorded in Stevenson Add 2 with short description from Tay (Perth, Buchanan White).

TFTaDMSSoCAR P. menthae On Mentha aquatica Forth (Aberlady, 24 viii 1952), Tay (Kittocks Den, St Andrews, 8 x 1936; Roscobie, Forfar, viii 1944; Loch of the Lowes, 1 ix 1976; Ballinluig, iv 1917; Umbog Bog, nr. St Andrews, A.Angus, no date), Ross (Kishorn, 6 ix 1960) and from N.Uist, Barra, Tiree, Coll, Raasay, Skye, Mull, incl. Iona Colonsay, Lismore, Islay, Kintyre, Arran and Orkney. On M. hirsuta x aquatica about Edinb. Sept. 1821? Possibly Greville Herb. On M. arvensis Tay (Dunkeld, 23 viii 1953), Forth (Flanders Moss, 1 ix 1957). On M. x piperata Tay (Kinbelthmont, nr. Arbroath, x 1944, U. Duncan; Brechin, 1932, G.Wallace) and Ross (Slumbay, 13 x 1959). Old records in E on Mentha piperata ‘nr. Edinb. Oct 1821’ prob. Greville collection. On M. sativa ? Moray (Loch Duntelchaig, 27 viii 1938). On M. verticllata Tay (Blairgowrie, 17 ix 1959), Clyde (Rosshdu, 11 ix 1959). On M. viridis Argyll (Mallaig, viii 1929) and Tay (Pitform vii 1845 prob. R Stark). Also known from Tay (Fearnan Loch Tay, vii 1948 U. Duncan; Boarhills, Fife, 23 vi 1981, A.Angus), Tweed (Peebles 10 vi 1934 & 12 ix 1936), Forth (Crossford, Dunfermline, 3 vii 1979; Gosford Nat. Res., 29 ix 1971). Also on Mentha sp. Forth (Duddingston Loch, 11 ix 2009, J. Muscott). Known on cultivated mint from Forth (Blinkbonny Edinb., 12 v 1980 & 29 iv 1981), Tay (St Andrews Bot Gdn., 4 vi 1966). ‘on Mentha in garden Balmutto Miss Boswell May’ as ‘Uredo labiatarum on M. arvensis Arbay nr Kinross’ & same taxon from Balmutto no date. All poss. Greville collections. First recorded on cultivated mint in 1924 and widespread and common causing concern in 1948 & 1953-1954, also 1956 in North East and 1957 in South West according to Foister (1958). On Origanum vulgare Tay (St Cyrus, 1 viii n1944, U.Duncan; Dunkeld 21 vi 1944, U. Duncan). Recorded from all Watersheds in Stevenson except Solway, Argylle, Ross and Sutherland and as Aecidium menthae from Tay, Dee, Moray and Clyde.

TFTaDMSSoCAR P. microsora II/III Known on Carex vescicaria from Moray (Inshriach, Aviemore, 23 ix 1959 & 12 ix 1964), Tay ( Killin, viii 1900 & x 1919) and Mull.

TFTaDMSSoCAR P. moliniae On Molinia caerulea from Argyll (Ben Lui, D.M.Henderson, 16 ix 1959 and as P. brunellarum moliniae, D.M.Henderson, 28 viii 1953; Glencoe village, D.M.Henderson, 24 ix 1968; Glen Marnie near Arisaig, M.Wilson?, ix 1932; near Bridge of Orchy, x 1922), Ross (Wester Ross, D.M.Henderson,16 x 1959). Also Hoy Orkney, Harris, Coll and Mull. Two collections on Molinia with II/III probably from Scotland simply labelled Britain D.M.Henderson 10 ix 1963 & 14 ix 1964. On Prunella vulgaris Sutherland (Borora River,10 vi 1966). As P. brunellarum-moliniae. On Prunella sp. from Tay (Loch Tay, D.M.Henderson, 1 ix 1961) and III on Prunella seedlings, Edinburgh D.H.Rae, iv 1980. Recorded with short description on M. caerulea from Rannoch - Tay in Stevenson and in Add 4 for Moray.

TFTaDMSSoCAR P. nemoralis Material on Poa nemoralis  from Dee (Glen Tannar, 5 ix 1975) but noted from N.W Scotland.

TFTaDMSSoCAR P. nitida On Petroselinium ‘parsley’ and Heracleum etc. in Stevenson recorded from Forth and Moray as Trichobasis petroselini. Not considered Scottish in Check List.

TFTaDMSSoCAR P. obscura On Luzula campestris Solway (Logan, 9 vii 1972), Tay (Parkhill, Arbroath, ix 1944, U. Duncan; Tentsmuir, 5 vi 1967), Forth (RBGdn., Edinb., 5 viii 1953; Pentlands, 5 vii 1924), Solway (Lumpharan, vii 1922). Also records from North Uist, South Uist, Coll, Rhum, Mull, Colonsay, Gigha and Kerrera. On L. multiforme Argyll (Ben Lui, 265m., 16 ix 1959), Dee (Linn of Dee, 23 viii 1968), Tay (). On L. pilosa Moray (Rothiemurchus, 19 viii 1953), Clyde (Balloch, 11 ix 1959), Tay (Tentsmuir, 1`5 vi 1966, U. Duncan; Kindrogan, 25 ix 1964), Dee (Cairnwell, 1 xi 1959; Aberdeen, x 1921). On Luzula sylvatica incl. some record as P. luzulae Argyll (Ben Lui, 327m., 16 ix 1959), Moray (Aviemore, 12 ix 1964; Rothiemurchus, 12 ix 1957Nairn, viii 1942, U. Duncan; Darnaway, 24 ix 1955), Ross (Strome, iii 1959; Loch Carron, 12 x 1959), Forth (Roslin, vi 1917 & 12 x 1972; ; Carriber Glen, Linlithgow, 27 ix 1959; Humbie WD., 11 v 1941), Tay (Dunkeld, 13 ix 1960; Kindrogan, 30 viii 1976; Kenley Den, Fife, 12 vii 1965, A.Angus; Glen Clova, 21 vi 1965; Drumtochy, Angus, 30 ix 1953; Edzell, ix 1944, U. Duncan; Glen Lyon, 5 iii 1966 & 14 ii 1966; Murthly, ix 1925, N.L.Alcock; Bilbo Wd., St Andrews, N.L. Alcock x 1930), Dee (Moneymusk, 9 ix 1995, A. Angus), Clyde (Rossdhu, 17 ix 1979; Incailloch, Loch Lomond, 17 v 1974; Cadzow Castle, Hamilton, 11 ix 1959), Sutherland (Evanston, 1942 U. Duncan; Loch Skene Lairig, 21 viii 1944, RWG Dennis) and Solway (Kilkerron, Ayr ix 1926). Also records from Lewis, Harris, Scalpay, Barra, Mingulay, Raasay, Arran  Skye,. Rhum, Mull, Garvellochs, Islay, Jura and Kintyre; Orkney (Rousay) and Skye. I on Bellis perennis Tweed (Berwick, ix 1922, M.Orr), Tay (St Andrews 12 viii 1957, 6 vi 1975 & 12 vi 1979), Clyde (Cartland Crags, Chatelherult, Fiddler’s Glen & U. Nethan; Balloch, 11 ix 1959), Forth (Edinb., v 2004; Roslin, 14 x 1961; Balerno, 1 v 1960; RBGdn., Edinb., 29 ix 1961), Sutherland (Thurso, Dunnett Links, v 1943, U. Duncan), South Harris, Benbecula, South Uist, Barra, Tiree, Coll, Skye, Rhum, Mull incl. Iona, and Colonsay; also Orkney incl. Rousay and Shetland, incl. Fetlar & Muckle Roe. Old collection in E possibly Greville ‘Roslin Glen as L. maxima, vii 1821’. In Stevenson as Aecidium compositarum var bellidis from Tay (Fern, Rev. Ferguson). Attacked  by Eudarluca filum Foula

TFTaDMSSoCAR P. opizii On Carex muricata Forth (Pitscandly, nr. Forfar, x 1944, U Duncan). On C. panicea Belford, 1 viii 1982 (taken from Herb specimen), Forth (Arhtur’s Seat, Edinb. vii 1835 taken from Herb. secimen). On C.paniculata Tay (Rae Loch, Blairgowrie, 10 iv 1955).

TFTaDMSSoCAR P. oxalidis II/III on cultivated Oxails adenostylis in garden Forth (Blinkbonny Edinb.1 viii 1999, 22 viii 2000 & 16 viii 2004) and II found on Islay Port Ellen on Oxalis corymbosa. Undoubtedly introduced

TFTaDMSSoCAR P. oxyriae on Oxyria dygina Tay (Lochan nan Cat. Ben Lawers 23 viii 1986; Ben Ledi, 431m., 12 viii 1951 abundant in damp gulleys, 12 viii 1957; Creag an Lochan 610m.-914m., 27 vii 1952 & 13 ix 1960), Dee (nr. Braemar, ix 1929), Argyll (Ben Lui, viii 1925 & 2007) and Moray (Ben Chonzie 732 m., 4 viii 1976, B.J. Coppins). Also record from Skye.

In Stevenson Add 5. With short descripiton for Dee Braemar, Prof. Trail) and Skye (Buchanan White) in Add 5.

TFTaDMSSoCAR P. paludosa Many records as Puccinia caricina var. paludosa. On Carex bigeloweii with III from Easterness, vii 1892 and Moray (II Cairngorm, A.Sommerville, vii 1893). On C.goodenovii from Ross (Kintail, viii 1933), Dee (Aberdeenshire Lamphanan, M.Wilson, x 1922), Argyll (Taynuilt near Loch Etive, M.Wilson, vii 1921) and Moray (Loch Dutelchaig, 1936). On C. nigra from Moray (Inverness Aviemore Loch Pityoulish, D.M.Henderson, 23 ix 1959). On C. panicea known from Coll, Islay and Harris, Dennis, 1986 and Orkney. On Pedicularis palustris from Moray (I Ballindalloch, D.F.Mackenzie, vii 1936), Dee (Aberdeenshire Lumphanan, M.Wilson, vii 1922, D.M.Henderson), Tay (Glen Lyon, D.M.Henderson,16 vi 1956) and Moray (Loch Duntelchaig, 31 vii 1939 & viii 1946). Also I Orkney (Ronaldsay), Skye and Rhum. See P. caricina.

TFTaDMSSoCAR P. pazschkei On Saxifraga aizion cv. Ben’, Forth (Edinb., AP.Bennell) date required. On S. callosa var. leontospathula, Forth (RBGdn., Edinb., 26 iv 1960). On S. cotyledon var caterbariensis, Nursery, St Andrews, 10 vii 1979). On S. crustata var. rochinensis RBGdn. Edinb.). On S. hosta var. rhaetica Forth (RBGdn., Edinb. ix 1920). On S. hammeri Forth (RBGdn., Edinb. x 1960). On S. holhamneni Forth (RBGdn., Edinb., 1 ii 1961).

On S. herzon var. cullata, Forth (RBGdn., Edinb., ix 1920). On S. onnetica Forth (RBGdn., Edinb. 1 ii 1961).

On S. parophylla Forth (R.B.Gdn., Edinb., 1 ii 1961 & cv. ‘Sibthorpisana’ 1 ii 1961). On Saxifraga cvs. ‘Bridget’ and ‘Rigoi’ Forth (RBGdn., Edinb., 1 ii 1961). Noted by Foister (1958) as III on nursery stock of Rockfoils.

TFTaDMSSoCAR var. juliana III On S. aizoides Ross (Loch Lean, Kishorn, 13 & 19 x 1959, 244m., 22 ix 1960 & 19 ix 1963), Sutherland (Bettyhill, 18 vii 1957) and Argylle (Ben Lui, 426m., 16 ix 1959). On S. oppositifolia Tay (Creag an Lochan, Lawers, 610m. & 671m., 17 ix 1959). Only known from Scotland.

TFTaDMSSoCAR P. pelargonii-zonalis Known on cultivated Pelargonium spp. Clyde (Helensburgh, vi 1968), Tweed (Peebles 20 xi 1970) and Forth (Haddington, 24 x 1979).

TFTaDMSSoCAR P. phragmitis On Phragmites communis from Tay (Blairgowrie, iv 1930; nr. Forfar, x1 1944, U. Duncan; Myreside, Blairgowrie, 15 v 1948 as Puccinia trailii; Rae Loch, Blairgowrie, 11 iv 1955; Bot Dept St Andrews by inoculation 27-30 v 1960 & 13 vi 1960; Loch Kilconquaher, St Andrews, 16 vi 1966), Clyde ( Benmore, no further data), Solway (Gatehouse, iv 1944), Ross (Morven and Ardnamurchan); also Coldingham, viii 1930.

In addition old collection labelled Sphaeria arundinacei from ‘Edinburgh, M.J.Stewart, 1820s?). Also old collection labelled Puccinia graminis var. arundinis Greville Edinb. Sept.’ On Rheum raponticum Tay (Bot Gdn. St Andrews, vi 1959). On Rumex acetosa Solway (Thornhill/Dumfries, vi 1930). Also on this host as Aecidium purpurascens vi 1823.

On R. crispus Solway (Thornhill/Dumfries, vi 1930), Tay (Tentsmuir, 27 v 1956 and Rumex sp.? 21 v 1965). On R. hydrolapatum Tay (Usan, Montrose, viii 1930). On R. obtusifolius Dee (Lunan Bay, Kincardineshire, 2 vi 1974), Solway (Castle Douglas, 20 v 1963), Tweed (The Hirsel Berwickshire, 3 v 1970). As Aecidium rubellum in Stevenson probably based on Greville’s collection in E from ‘Duddingston Loch, Edinb. May to June ‘on various spp. of Rumex particularly R. aquaticus’, Tay and Dee and probably records of Puccinia arundinacea from Forth & Tay in Stevenson refer here and App. 1 for Dee.

TFTaDMSSoCAR P. pimpinellae Recorded on Pimpinella saxifraga from Moray (Aviemore, Rev. Keith) and Tweed (Eyemouth, viii 1922). As Aecidium bunii in Greville Herb. and collected by Miss Boswell probably from Fife, even Balmutto. Also record of Greville on Pimpinella in E labelled ‘Aecidium umbelliferarum Autumn’.  In Stevenson Add 2 and Dee in Add. 8.

TFTaDMSSoCAR P. poarum II/III on Poa spp. On P.nemoralis Tay (Spittal iof Glenshee, 6 vii 1963; Blairgowrie, 28 v 1955). On P. pratenis Forth (Stirling, 9 vii 1966; Balerno, 10 viii 1954), Clyde (Loch Lomond, Balloch Pk., 11 ix 1959; Cadzow, Hamilton 14 ix 1959), Ross (Gairloch, 20 vii 1955), Tay (Glen Fincastle, 13 vii 1969; Blairgowrie, 10 viii 1953; Kimblethmont, nr. Arbroath, viii 1944, U. Duncan). On P. trivialis Argyll (Glen Lochay, 16 vii 1962), Tay (Glen Eagles, 27 vi 1970; Guardbridge, St Andrews, 12 vii 1938) and Shapinsay. On unident. grass Forth (Linlthgow, 6 ix 1941, RWG Dennis). Known from Orkney incl Hoy, Barra, Mingulay, Coll, Skye, Colonsay, Islay, Arran, Gigha and Kintyre and. P. nemoralis or P. trivialis. Frequent on lawn Poa spp. according to Foister (1958). O/I on Tussilago farfara Tay (Eden Estuary, St Andrews, 22 v 1966, M. Stewart; Kingsbarn Fife, 3.vii 1960, A. Angus; Eden estate, St Andrews, 22 v 1966; Kingsbarn, 3 vii 1976; Glen Fincastle, 13 vii 1969; Spittal of Glenshee, 6 vii; 1963Parkhill, Arbroath, v 1944, U. Duncan; Glen Eagles, 27 vi 1970), Moray (Loch Duntelchaig, 1936 no further details), Forth (Hillend, Edinb., iv 1926; Balerno, v 1936; Ratho, v & viii 1934; Kirkcaldy, no date; Largo, Fife, 16 vi 1954; Stirling, 9 vii 1966), Argyll (Loch Long, 13 vi 1976; Glen Lochay, 16 vii 1962) and Moray (Loch Duchtelchaig, 1936).

Also on Orkney incl. Hoy, Barra, Mingulay, Coll, Skye, Islay, Arran, Gigha, Kintyre and Eigg). Greville collection as Aecidium tussilaginis, no further details. Other old collections in E as Aecidium compositarum Tay (Carnoustie, ix 1844; Baldovan Wd., Dundee, vi 1845). Also as var. tussilaginis from all Watersheds in Stevenson except Sutherland, Solway, Argylle and Ross; Argyle in Add 4..

TFTaDMAAoCAR P. polygoni-amphibii On Polygonum amphibium from Tay (Arbroath on var. terrestre, ix 1944, U. Duncan; Ballinluig, x 1921; St Andrews 5 x 1935 & xi 1935 - host in one collection re-determined), Forth (Tyninghame, on var. terrestre, 2 x 1954), Clyde (Rothesay, Lister’s nursery, 6 x 1950), Solway (Newton Stweart, Mossyard Bay, ix 1929), Arran, on var. terrestre, ix 1936, Also records from Tiree, Mull, Islay, Arran and Kintyre. In E coll. as Uredo polygorium ‘about Edinb. Sept. 1821’ prob. Greville and coll. ‘Edinb. prob. 1820s Greville?; as P. amphibii introduced with short decription when recorded from Tay (Scone etc. Buchanan White) in Stevenson Add 2. And Dee and Moray in Add. 8.

TFTaDMSSoCAR var. convolvuli Known in Scotland on Fallopia convolvulus only from inoculation  experiemts in Forth (RBGdn., Edinb.).

TFTaDMSSoCAR P. primulae On Primula vulgaris Tweed (Cockburnspath, viii 1932; Broughton, 8 v 1954;), Tay (Belbo Wd., St Andrews, x 1930, NL. Alcock; Dores Foyers Rd., St Andrews, 4 v 1934; nr. Arbroath, v 1944, U. Duncan), Moray (Shores of Loch Ness, 18 iv 1943, U. Duncan), Forth (North Berwick, 1 v 1960; Bridge of Allan, vi 1936; Granthouse, 16 iv 1948; RBGdn., Edinb., vii 1940), Ross (Loch Maree 18 ix 1969), Argyll (Crinan, 25 v 1963; Garnelloch Is., 14 v 1984; Taynuilt, 20 v 1921; Inverliever v 1923; Glen Branter, vii 1929), , Solway (Kintyre, viii 1931, NL Alcock; Ayrshire no further data ix 1926)Barra, Skye, Rhum, Mull, incl. Ulva Colonsay, Islay, Arran, Eigg (vii-viii 1949 Univ. Exped.)and Gigha; also Orkney, incl. Hoy. Collections in E from Forth; ‘nr Kirkcaldy as Uredo primulae July 1821’ and 2nd specimen Puccinia primulae nov. sp. on Primula vulgaris July. Sometimes parasitic on Uredo primulae Kirkcaldy 1821’, Greville); also prob. Aecidium ‘on Primrose no further data. Apparently common throughout area but overlooked’. In Stevenson recorded from Forth and Tay and as Aecidium primulae from Tay and Argyle.

TFTaDMSSoCAR P. prostii O & III on Tulipa sylvestria Forth (RBGdn., Edinb., vi 1914 onwards to 3 v 1971 & 10 iv 1981, D.M.Henderson) and on var. odorata 31 v 1951.

TFTaDMSSoCAR P. pulverulenta On Epilobium hirsutum Forth (East Linton, East Lothian, viii 1922), Tay (River Ericht, ix 1952; Blairgowrie, 31 v 1955; St Vigeans, nr. Arbroath, vi 1944, U. Duncan; Dura Den, St Andrews, 12-13 ix 1948; McIntosh Hall, St Andrews, 19 ixc 1952; Kinkill St Andrews, 17 vii 1966; Balmutto 12 vi 1959; Dumbog bog, nr. ? Thrie, N.Fife, 9 ix 1971) and Moray (Inverness, vii 1942). On E. montanum Forth (Edinb., viii 1933; Balerno, 26 ix 1942, RWG Dennis; Bridge of Allan, vi 1936;  Blinkbonny Av. Edinb., 7 vi 1972 & 1 iv 2005; Pathhead 18 vi 1951), Tay (Ballinluig, vii 1915; Callander, vi 1923; Pass of Killiecrankie, vi 1944, U.Duncan; Priormuir Station, St Andrews, 23 vi 1966), Argyll (Ben Challum, Tyndrum, 304m., 22 vi 1958), Solway (Lumphanan, ix 1922; poss. same host, Thornhill, ix 1924, NL Alcock); also Mull. In E as Aecidium epilobii Pitform vi 1845. On E. palustre, Glen Forsa, Mull. On E. parvifolorum Tweed (Glentress, Peebles, 4 vii 1951), Dee (Moneymusk, 9 ix 1975) and Ross (Strome, viii 1959). On Epilobium sp. Mull and Orkney. Greville material in E as P. obtusata nov. sp. ‘on Epilobium montanum about Edinburgh & on E. hirsutum about Edinb.’. Recorded from Tweed, Forth, Tay, Dee on E. hirsutum and as Aecidium epilobii from Tweed, Forth, Tay, Dee and Moray in Stevenson.

TFTaDMSSoCAR P. punctata On Gallium cruciatum Forth (Aberlady, v 1931; Dalkieth, 13 x 1956; Gullane, 5 xi 2006, M. Clarkson), Clyde (Cadzow Pk., Hamilton, 14 ix 1949) and Tay (Blairgowrie,. 1 x 1947). On G. palustre Moray (poss. this host Tomich, 10 ix 1963), Clyde (Dougalston, 9 ix 1959), Argyll (Loch Seil, 3 viii 1984), Loch Druich, Kintail, viii 1933 & Mull. On G. saxatile Moray (Lundei Loch Black Isle, 13 ix 1942, U. Duncan), Solway  (Thornhill, ix 1924, Scott-Elliot) and Iona. On G. uliginosum Moray (Loch Insh, Aviemore, 11 x 1959), Tay (Loch Craiglush, Dunkeld, 6 ix 1953; Tentsmuiur, 10 x 1965). On G.verum Dee (Bridge of Don, viii 1962), Clyde (Cadzow Pk., Hamilton, 14 ix 1959), Forth (Pathhead, 6 ix 1925. G.Wallace), Moray (Loch Duntelchaig, 29 viii 1935 thru’ 3 & 21-30 vii 1939; Culbin, 9 ix 1963), Tay (St Andrews, 15 v 1937; Blairgowrie, 2 viii 1957; West Bay, St Andrews, v 1937  & x 1937; Redhead nr. Arbroath, v & ix 1944, U. Duncan; Stormont Loch, Blairgowrie, 26 vi 1963; Dunkeld, 18 vii 1959), Ross (Durness 26 vi 1962) and Dee (Dunecht, 9 ix 1975). On G. verum Coirra Seilebost, Harris, RWG Dennis; also North Uist, Vallay, South Uist, Barra, Tiree, Skye, Rhum, Mull, Islay, Kintyre, Shetland and Orkney. As P. galiorum from Forth, Tay, Moray & Clyde in Stevenson and in Add 3 Dee added and Argyle in Add 4.

TFTaDMSSoCAR P. punctiformis Included here records as Puccinia otegens & P. suaveolens. On Cirsium arvense Moray (Strathpeffer 23 vi 1984; Kessock, Black Isle, viii 1942, U.Duncan), Ross (24 vi 2004; Durness, 20 vi 1962), Forth (Balerno, Edinb., v 1936; Glen Corse, Penicuik, viii 1934; Orchard Brae, Edinb., 8 vi 1976; Stirling, 9 v 1971), Sutherland (Lybister, 22 v 1987), Tweed (Melrose 13 vii 1941),Tay (Whitefield, Dundee, 5 x 1975; Kenley Den, St Andrews, 14 vi 1965 & 15 vi 1966; Boarhills, Fife, 7 vi 1966; Glen Dochart, 17 vi 2006; Parkhill Arbroath v 1944 & Kinblethmont both Arbroath, v 1944 & viii 1944 U Duncan; Murthly, ix 1925, NL Alcock; Brechin, 13 vi 1946; nr. Carnoustie, v 1845); also Eigg (vii-viii 1949 Univ. Exp.,) Baghead North Uist, Lewis, Harris, Barra, Raasay, Skye, Arran, Rhum, Eigg, Mull, Iona, Colonsay, Islay, Gigha, Kintyre, Orkney and Shetland. On C. arvense var. setosus Moray (Inverness, ? 1935, J.A.Mcdonald) and Forth (Holyrood, Edinb., 9 viii 2004; Edinb., viii 1871; Crichton, Midlothian, 28 vi 1928; Gulland, vi 1931;  Aberlady, vi 1930 & 14 vi 1934) and Clyde (Gerrard’s Cross v 1911).

Old collections in E by R. Stark ‘on old banks of canal as Uredo suaveolens ix 1836’ and under same name ‘about Edinb. spring & autumn’ Greville collection Undocumented collection poss. Menzies. August 1821. In Stevenson as Trichobasis suaveolens and recorded from Tweed, Forth, Tay and Moray and Add. I for Dee.

TFTaDMSSoCAR P. pygmea On Calamagrostis epigeos Tay (Dunkeld, 23 viii 1953 & 18 ix 1959) and Moray (Loch Morlich, 23 ix 1959).

TFTaDMSSoCAR var. ammophilina On Ammophila arenaria Tay (St Cyrus, no dates; Gatty Lab., St Andrews as Uredo, 3 xi 1953), Forth (Burntisland, xi 1924; Scorghall, East Lothian, 24 ix 1939 RWG Dennis; North Berwick, 20 ix 1972; Aberlady, 21 vii 1951), Ross (Durness, viii 1931 & 30 viii 1944), Solway (Wigtonsahire, Sands of Luce, 20 ix 1953), Siullouth, vii 1923 and from Lewis, Harris, North Uist, South Uist, Vatersay, Tiree, Coll, Iona, Colonsay, Islay, Arran and Kintyre; also Shetland and Orkney. On plantings of Ammophila arenaria; see Foister, 1958.

TFTaDMSSoCAR P. recondita II & III on a range of grasses. f sp. agropyrina On Agropyron caninum, as Puccinia agropyrenia, II, Moray(Loch Duntelchaig, J.A.Macdonald, 27 vii 1939), Arran. As P. dispersa on Agropyron caninum Dee (Forvie Sands, Mouth of Ythen, 10 ix 1975}. On A. donianum Tay (Creag an Lochan, 529m., 28 v iii 1951, 17 ix 1959 & 10 vi 1961). A.junceiforme Arran. Also Forth (nr. Craill, 12 ix 1960; Gullane, 10 x 1963). On Agropyron repens Moray (Ballandalloch, ix 1936), Clyde (nr. Kilmarnock, vii 1940; Cadzow Pk., Hamilton, 19 ix 1959), Forth (Kirknewton, 9 x 1959), Tay (Boarhills, 16 vi 1966) Lewis and Tiree, Mainland Orkney and Papa Westray, St Kilda. and as P. agropyrina on A repens Cameron, J.A.Macdonld, 22 x 1958. As P. dispersa var. agropyrina Tay (Montrose, 29 vii 1944, U. Duncan; St Cyrus, vii 1944, U. Duncan), Forth (Aberlady, 5 ix 1959; East Craigs, 22 viii 1940 & 19 i 1943, RWG Dennis), Dee (nr. Stonehaven, 14 vi 1959). Occurs on Agropyron in Scotland including A acutum (A.junceiforme x A. pungens; Foister, 1958).Attacked by Eudarluca filum Forth (Corstorphine, 16 ix 1939). f. sp. agrostidis On Agrostis stolonifera as P. agrostidis Moray (nr. Forres, 1 viii 1968). On A.vulgaris Tay (Dunkeld, viii 1932). On Agrostis sp.  Dee (Linn of Dee, 23 viii 1968). Isolated from seed from New Zealand as P. agrostidis 1924-37 & 1948. 0/I on various hosts. f.sp. borealis On Thalictrum alpinum Argyll (Ben Lui, 9 vii 1995 & vii 1915 & 1919).f. f. sp. bromina On Bromus asper Solway (Newton Stewart ix 1929). On B. ramosus Forth (Roslin, 28 viii 1943, RWG Dennis; Balerno, 31 vii 1954). On Bromus sterilis Forth (Ferry Rd., Edinb., 16 v 1984; Pathhead, vii 1925, G.Wallace; Currie, 30 v 1959). Clyde (Brand, Glasgow, 26 VI 1931). On B.thomsonii (= mollis) Tay (Lunan Bay, Angus, vi 1944). On Deschampsia caespitosa as P. dispersa on Deschampsia (and Aira) caespitosa Solway (Thornhill, 25 ix 1924, S. Maxwell), Tay (Ballinluig, vii 1915) and Forth (Edinb., xi 1924; Barnton no further data & viii 1922). Probably these records refer to P. brachypodii var. arrhenatheri. .f. sp. holcina On Holcus lanatus Tweed (Dawyck, 15 ix 2002; Duns Castle, 28 ix 2002) Orkney. As P. holcina Chatelherult & H. mollis Hamilton Wilson & Henderson and as P. holcina Wilson, 1934. In Foister (1958) noted also as P. dispersa and on commercial Holcus lanatus (Yorkshire Fog). On H. mollis Tay (Tentsmuir, 18 v 1989), Forth (Ferry Rd., Edinb., no date). f. sp. persistens On Elymus arenarius Tay (Tentsmuir, viii 1958). On Festuca gigantea Forth (Roslin, 14 x 1961), Solway (Newton Stewart, ix 1929), Clyde (Auchincruivre, ix 1933). Records probably refer to P. coronata. On Hordeum vulgare Forth (East Craigs, vi 1976, M.J.Richardson; Pathhead, 1 viii 1979); probably record refers to P. hordei. The first Forth collection attacked by Eudarluca. sp. plexans On Alopecurus pratensis as P. perplexans Foister, 1958. On Anthoxanthum odoratum Ross (Loch Carron, 2 iv 1961), Tay (Brechin, 20 vi 1946, G. Wallace). A collection which needs to be reassessed prob. P. brachypodii var. poae-nemoralis. On Arrhenatherum elatius Forth (Balerno, 23 iv 1961; Corstrophine Hill, Edinb., 27 vii 1952), Tay (Kindrogan, 239m., 25 ix 1964) and Iona. Records need to be reassessed ands probably belong to P. brachypodii var. arrhenatherif. sp. recondita on Secale cerealis Forth (Corstorphine, 1941; East Craigs, 16 x 1940, RWG Dennis), Tay (Brechin ? vii 1925, G.Wallace and as P. secalina St Michaels, Leuchars, 3 xi 1948). Attacked by Eudarluca filum Forth (Corstorphine, 14 xi 1944). On Anchusa arvensis Tay (Leuchars, 3 xi 1948); also see above f. sp. trisetii Forth (Crichton Castle, 3 vii 1964). As P. rubigo-vera Forth (Arthur’s Seat, on Avena flavescens, from Herb specimen 1871; as P. triseti Pathhead, 21 ix 1925, G.Wallace; Liberton, x 1924; Gullane, vii 1928). f. sp. tritici On Tritcium vulgare Dee (North Water Bridge, 23 viii 1961), Forth (Boghall, x 1946), Dee (Craibstone, no date E Gray). On Holcus and other Graminae Arran  (Boyd as P. rubigovera) and Tweed (Peebles, 27 v 1933). Also in Foister (1958) as P. triticina and P. rubigo-vera on Triticium aestivum and T. turgidum. Common in 1947-19511 & 1954, severe in S East and South West 1945; known from S. East, S. West and East Central and N. West. P. dispersa recorded as not severe in Foister (1958) on wheat in S east, S, W & E Central and North West. Old collection labelled Uredo linearis prob Forth (prob. nr. Einb., on Tritcium repens R.Stark). As Aecidium thalictri Clyde (Ben Voirlich Loch Lomond, 1821, Mr.Paterson type in E). On T. dunense Forth (Gullane Pt., 1 vi 1963 & 10 x 1963). In Stevenson as Trichobasis rubigo-vera from Tweed, Forth, Tay, Dee, Ross and Clyde, with description and as Aecidium thalictri from Forth, Tay and Dee and as A. asperifolium on Lycopsis arvensis from Tay (St Andrews, Rev. Anderson; Leuchars, 1938).

TFTaDMSSoCAR P. ribis III on Ribes spicatum Known only from Tay & Moray. Records on Ribes rubrum Moray (Coulmoray, 11 ix 1946 & vi 1947, M.Noble), Dee (Pittolnie Castle Hotel, nr. Inver, 27 viii 1990), Tay Fincastle. Pitlochry, ix 1948) do not belong here.

TFTaDMSSoCAR P. saniculae On Sanicula europea Forth (Saltoun Wd., R.Stark, 1836; Balerno, 26 ix 1942, RWG Dennis; Pathhead, v & x 1923; Eskbank, 12 iv 1953; Gifford, 14 vi 1954; Spott E. Lothian, 31 v 1942, RWG Dennis), Tay (?Bagley Wd., 5 ix 1954 JA Mcdonald), Moray (Loch Ness 18 iv 1943, U.Duncan), Argyll (Glasdrum, 28 ix 1985; Glenbranter, vii 1929) and Clyde (Ayrshire no further locality, ix 1926, NL Alcock) and from Mull, Arran and Kintyre. Greville’s type material from ‘Auchendinny Wd August 1821’ in E. In Stevenson recorded from Tweed, Forth, Tay & Argylle and as Aecidium saniculae from Moray (Grantown, Rev. Keith).

TFTaDMSSoCAR P. saxifragae On Saxifraga granulata Forth (Hopetoun Hse., 25 iv 1960 & 19 iv 1978). On Saxifraga stellaris Tay (Meall nan Tarmachan, 914m., 10 vii 1965; Ben Lawers 1,067m., 15 ix 1956; Ben y Vrackie, 30 vii 1930; Meall Garbh, Lawers, 1 vi 1950), Solway (Loch Skene, Moffat, 488m., 20 vi 1959; Whitecombe, 610m., 25 vii 1957), Ross (Beinn Eighe, 762m., 21 vii 1955), Argylle (Ben Lui, 19 ix 1964; Caenlochan Glen, 25 vi 1950) and from South Uist and Mull. As P. saxifragarum in Stevenson from all but Argylle, Ross, Dee & Sutherland and as Uredo saxifragarum from Forth & Tay.

TFTaDMSSoCAR P. septentrionalis II/III on Persicaria viviparum Tay (Creag an Lochan, Killin, 610m., 26 viii 610m.-761m., 26 vii 1952 & 26 viii 1953; Fincastle, 24 vi 1977; Meal nan Tarmarchan, 9 x 1919; Ben Lawers, viii 1921 & 2 ix 1968; Glen Doll, 1 vii 1946), Sutherland (Laggan, Bonar bridge,487m., 1926 no further data), Ross (Beinn Eighe, 548m., 24 ix 1956), Argylle (Ben Lui, x 1914, vii 1915, ix 1921 & 16 ix 1959). I on Thalictrum alpinum Tay (Meall non Tarmachan, 21 v 1960; Ben Lawers, 609m., vi 1936, 24 vi 1950 & 20 vi 1952; Glen Doll, 1 vii 1946, 7 vii 1977 & vii 1947, U. Duncan; Creag an Lochan, Killin, 610m.,  5 vii 2003 & 16 vi 1968; Creag na Callich, 7 vii 1996; Corrie Fee 20 vi 1966), Argyll (Ben Lui, 26 vi 1979; Beinn Chronach, Tyndrum, 671m., , 22 vi 1954) and Dee (Caenlochan Glen, 25 vi 1950; ; Glen Clova, 21 vi 1965). As Aecidum thalctri Glen Callanter vii 1875 - Dee. Also on P. bistorta from infection on T. alpinum viii 1922.

TFTaDMSSoCAR P. sessilis On Phalaris arundinacea Tay (Lade Brae, St Andrews, x 1955; Dura Den, St Andrews, 13 ix 1940; Boarhills, St Andrews, 20 x 1981; Dunkeld, 18 ix 1959 & 13 ix 1960; Dairie Bridge, Fife, 17 v 1965, A.Angus), Forth (West Linton, 5 ix 1962; nr. Roslin, 16 iv 1961), Solway (Thornhill, ix 1924, NL Alcock), Tweed (Churnside, Duns, iv 1934; Duns Castle, 28 ix 2002), Clyde (Hamilton, 14 ix 1959) and from Tiree, Mull, incl. Iona, Skye, Arran and Kintyre. O/I On Allium ursinum (as P. winteriana) Solway (Castle Douglas, 20 v 1963), Dee (Fyrie Castle, 27 v 1985), Tweed (Dunlaw Dean, Berwickshire, 18 vi 1933), Clyde (Cadzow, Hamilton, 14 ix 1959 19 v 1985; Chatelherult), Tay (Dura Den, St Andrews, v 1937; 13 ix 1944 & 15 vi 1951 & v 1937; Mill on Eden, St Andrews, 16 vi 1966), Moray (Kerries, Inverness 19 v 1935) and Forth (Stirling Campus, 28 v 1872; Waters of Leith, Edinb., 18 v 1983), from Eigg, Skye, Orkney and Arran. On Arum maculatus (as P. phalaridis) Clyde (Auchincruivre, iv 1949; Hamilton High Park, 19 v 1985) and from Islay, Mull and Orkney. Noted by Foister (1958). On Orchis purpurella Solway (Lochinvar, 29 vi 1961, G.Hadley). Greville collections in E as Aecidium allii ‘June-July on Allium ursinum Roslin Wd 1821’ and ‘on Orchis maculata Killin vi 1822’. Attacked by Eudarluca filum Tweed (Duns Castle, 20 ix 2006). In Stevenson as Aecidium allii on A. ursinum from Tweed (Jedburgh, A.Jerdon) and Forth (Edinb. Greville) and as A. orchidearum on Orchis (=prob. Dactylorchis) from Tay (Burghill, Monikie, Rev. Ferguson). On A. paradoxum Forth (Springfield, 11 iv 2009, J. Muscott).

TFTaDMSSoCAR P. smyrinii On Smyrnium olusatrum. Single record from Forth (Weymss Bay, Fife, 25 iv 1953 D.M. Henderson).

TFTaDMSSoCAR P. solidinallae On Solidinella alpina ‘Moonwort’ in Stevenson for Clyde(O, Glasgow BGdn.) and Forth (RBGdn., Edinb.). Considered alien in British Check List.

TFTaDMSSoCAR P. striiformis var. striiformis Record on Arrenatherum elatius (as A. avenaceum, Forth (Boghall 1925) which needs to be reassessed. On Elymus arenarius Tay  (St Andrews, x 1930, NL Alcock; West sands, St Andrews, 6 ix 1936), Forth (North Berwick, 20 ix 1922). On Hordeum distichum, Tay (Kinblethmont, nr. Arboraoth, ix 1944 U.Duncan). On H. murinum Forth (Berwick area, ix 1922, M.Orr; Corstorphine, Edinb., vii 1944, M. Noble; Kirkcaldy, 19 vii 1944), Tay (Dura Den, St Andrews, 6 x 1948; Burns Nursery, St Andrews, 18 ix 1938). On Hordeum vulgare Forth (Barley Balerno, 22 ix 1968; Roslin, 9 ix 1961; East Craigs, 1 ix 1968; North Berwick ix 1929; Pathhead, 1 viii 1979). On Triticium aestivum Forth (Linlithgow, 1 viii 1938; II & III Pathhead 6 ix 1924; Gifford, 10 ix 1961; Barnton, vii 1922; Boghall, 20 vi 1934; Pittenweem, viii 1919; on cv. Victor wheat, 19 vii 1943, RWG Dennis; on Algerian wheat, East Craigs, vi 1923 as P. glumarum; Coldingham, 4 vii 1942), Tay (Cupar, Fife, 6 vii 1943, Kinblethmont, nr. Arbroath, ix 1944, U. Duncan) and Solway (Longforgan Ayr, on cv. Little Joss wheat, 13 vi 1938). Also at Allofearn, 31 viii 1936, JA Mcdonald. On Triticale (Triticium x Secale) Forth (East Craigs, 29 vi 1944). Possible Greville collection in E ‘ Uredo inside calyx Oct. 1821. in S East especially Forth (Lothians). Common and occasionally harmful in 1938, 1939, 1942, 1945 in N.East & East Central, 1948 in N.East, East Central & S.East, 1949 in E & S Central, S.East and 1954 & 1957 in North East.) Recorded by Foister (1958) on Triticium aestivum & T. turgidum, Hordeum distichon & H. vulgare, Bromus carinatus and Agropyron repens Known since 1924 as problem in S.East especially Forth (Lothians), 1938, 1939, 1942, 1945 in N. East and E Central, 1948 in Central N.East E. Central and S. East, 1949 in Central, East and S.Central S east and 1954 & 1957 in N. East. In Stevenson recorded from Tay (Perth, Buchanan White) as P. glumarum. Common

TFTaDMSSoCAR Var. dactylidis As Puccinia coronata - P. glumarum on Bromus giganteus from Forth (Barnton, Edinb., viii 1922). On Dactylis glomerata from Mingulay.

TFTaDMSSoCAR P. tanaceti Probably introduced as yet only found on cultivated plants although found on Artemisia in wild. On A. absinthi Forth (Largo, Fife, viii 1843; Elie, Fife, xi 1915). On Artemisia maritima Forth (Craill, x 1937; East Lothian, no further location, x 1937; Tyninghame, ix 1959). On Artemisia vulgaris Forth (Holyrood, 11 ix 2009, J. Muscott). On Chrysanthemum sp., as P. chrysanthemi from Tay (on C. coccineum Boarhills, St Andrews, 30 x 1981 & 9 vii 1981; Dundee, Storrie Thyne & Co, x 1933) and Solway (Castle Douglas, xi 1931). Noted by Foister as widespread and sporadic on cultivated Chrysanthemum (C. morifolium). In Stevenson recorded as P. discoidearum from Tay (Charleton, Rev. Ferguson).

TFTaDMSSoCAR P. thymii On Thymus drucei from Dee (Aberdeen & Ballater Trail; Ballater, Ogilvie), Clyde (Glasgow) and Ross (as P. schneideri, Ben Eighe, 407m., 19 vii 1935).

TFTaDMSSoCAR P. tumida On Conopodium majus from Skye, Rhum, Arran (poss. misident.) and Mull. As P. umbelliferarum in Stevenson and recorded from all Watersheds except Ross, Sutherland, Argylle and Solway. As Puccinia umbelliferarum, Clyde (Dumbartonshire, viii 1836).

TFTaDMSSoCAR P. uliginosa Many collections incorporated in and labelled as P. caricina or as P. caricina var. uliginosa. On Parnassia palustris from Tay (I Balquidder, vi 1936), Ross (Glen Bog 10 vi 1952), Argyll (Ben Lui, D.M.Henderson, 28 vii 1953; Ben Nevis, R.J.Palling viii 1922; Taynuilt, M.Wilson, vii 1921 & ix 1942), Dee (Aberdeenshire Lumphanan, M.Wilson, vii 1922), Scotland, M.Wilson, 19 vi 1915 and Isle of Eigg, Edinb. Univ. Exped., vii-viii 1949. Also Mainland Orkney. As Aecidium parnassiae with short description in Stevenson for Forth, Tay, Dee and Clyde. On Carex nigra from Tay (III Ben Lui, D.M.Henderson, 28 viii 1953). Also known from Ulva.

TFTaDMSSoCAR P. umbilici In Stevenson recorded on Sedum rhodiola from Tay (Glen Callater, Gardiner) and Moray (Aberdour, Rev. Ferguson). Record on Umbilicus rupestris from Brodick, Arran, vi 1913 and Cotyledon umbilicus M.Wilson, 1934. Greville collection in E on Cotyledon umbilicus P. cotyledones no further data. Also fruiting in RBGE dinb. On plant of U. rupestris from Cornwall 8 11 1977.

TFTaDMSSoCAR P. urticata Recorded as P. caricina  and P. caricis. On Urtica dioica I. Known from Forth (Gifford, 14 vi 1954; Aberlady Gullane, J.A.Macdonald, 25 v 1933 & 14 vi 1934; Bonnyrigg, E.B.Basden, 24 vii 1980; Edinburgh O & I vii 1936), Argyll (Kentallen, A.P.Bennell, iv 1979), Ross (Wester Ross, Strathgarve Farm, coll. K. Grant, 1 vii 1980, R B.Bennell), Tay (Tentsmuir, M.Stewart, 21 v 1965), Moray (Nairn, B.J.Coppins, vii 19870 as P. urticae Loch Duntelchaig, 29 viii 1936.). Also known from Kergord, Mainland Shetland, R.Watling, 16 vi 1987 and Yell, R.Watling, 15 vi 1987, Mainland Orkney, Barra, Tanquray, D.M.Henderson, 23 v 1977 and Skye Quirang, J.A.Paton, 13 vi 1985. Known from Harris, Benbecula, North & South Uists, Barra, Eriskay, Raasay, Skye, Rhum, Mull, Colonsay, Oronsay, Luing, Gigha, Arran and Kintyre. As In Stevenson recorded from Tweed, Forth, Tay, Moray and Clyde as Aecidium urticiae.

var. bisporula

var. urticae-acutiformis

TFTaDMSSoCAR var. urticae-flaccae On Carex flacca several sites on Mainland Orkney and Birsay and on Rhum.

var. urticae-hirtae

var. urticae-inflatae

TFTaDMSSoCAR var. urticae-vescicariae As P. carcina var urtica-vesieaniae II on Carex vesicaria from Argylle (Loch Seil, D.M.Henderson, 20 vii 1984) and Moray (Aviemore, D.M.Henderson, 20 ix 1965 – 2 coll.); also Mull.

TFTaDMSSoCAR P. variabilis On T. palustris Tay (Creag an Lochan, Lawers, 5 vii 2003). On Taraxacum spp. Forth (Balerno, 1 v 1960; Aberlady 6 vi 1936; Borthwick castle, viii 1930; between Gullane & Aberlady, vi 1936), Sutherland (Beinn Eighe, 304m., 19 vii 1955; Tongue viii 1931), Tay (Tentsmuir, 25 vi 1957; Kindrogan 24 viii 1968; Ballanluig spring 1913 as P. silvatica but redet. variabilis vii `1915; Auchterhouse, Angus, 18 vi 1945, U. Duncan; also as P. dioicae nr. Lethan Grange, Angus, v 1944, U. Duncan; Blair Atholl, 304m., 14 vii 1968), Ross (Kishorn, ix 1965), Tweed (Glentress, vi 1930; Liflie’s leap Peebleshire, 16 v 1955) and Argyll (Loch Etive, 24 iv 1964; Tyndrum, 609m., 22 vi 1958). Known from South Uist, Barra, Tiree, Mull and Kintyre; also Orkney incl. Hoy; also Muirkirk, vi 1930. Type material of Greville in E. Recorded from Tweed, Tay, Forth & Moray in Stevenson and as Aecidium compositarum var. taraxaci from Tweed, Forth, Tay and Moray; Dee added in App.1 & Add 3.

TFTaDMSSoCAR P. veronicae On Veronica montana III Forth (Kippen, 12 ix 1959; Roslin, 12 xi 1972, also x 1935 & 28 viii 1943; Eskbank, 15 x 1962), Argyll (Loch Crerar, Argyll, 28 ix 1985 Benmore, ix 1928), Tay (R. Ericht ix 1952, U.Duncan; Blairgowrie 30 ix 1952) and Dee (Durris, 28 ix 2002). Known from Islay, Bridgend.

TFTaDMSSoCAR P. vinciae Known on Vinca sp. from Forth (Gifford, summer 1949). Also noted by Foister (1958) on Cultivated Periwinkle. Recorded from Tay (Maulsden, Rev. Ferguson) in Stevenson.

TFTaDMSSoCAR P. violae On Viola canina Tay (Baldovan Wd., Dundee, vi 1844; ; Cairnsmill, Fife, 28 vii 1966; Bridge of Cally, 3 x 1947), Solway (Thornhill, ix 1925; Kintyre, viii 1925, NL Alcock)), Moray (Loch Duntelchaig, 1936, 22 & 23 v 1937, 21-30 vii 1939) and Sutherland (Enard Bay, 23 viii 1944). On V. curtsii North Uist. On V. hirta Tay (Lundee Carigs, nr. Dundee, ix 1947, U. Duncan). On V. hirta x odorata Tay (St Cyrus, 13 viii 1978). On V. lutea as P. aegra on Forth (Boghall, 30 v 1957), Tweed (Peebles, viii 1941) and Tay (Brechin viii 1933, G.Wallace). On V.riviniana Forth (Balerno, v 1936; Roslin, 9 v 1952; Edinb., vi 1922), Dee (Dinnett, 6 ix 1975), Tay (Corrie fee, Glen Clova, 12 vi 1967 & 20 vi 1966; Glen Lyon, 26 ix 1983; Blairgowrie, 20 v 1984 & Rae Loch Blairgowrie, 31 v 1955), Tweed (Melrose 13 vii 1941; Loch Skene, Moffat, 487m., 20 vi 1959), Moray (Loch Duntelchaig, 3 vi 1966; Dalwhinnie, 7 viii 1987; Glen Strathfarrer, 25 ix 1985; Aviemore, 23 ix 1959; Creag Dunan, Inverness, v 1942, U. Duncan), Argyll (Ben Lui, 9 vii 1995; Benmore, 15 ix 1959; Taynuilt, vii 1921), Ross (Beinn Eighe, 304m., 18 vii 1955), Clyde (Glasgow no further data, Herb. Cooke). Also records from Lewis, Harris, Barra, Vatersay, Raasay, Tiree, Skye, Rhum, Mull, Colonsay, Lismore, Jura, Gigha, Arran, Kintyre, Islay, Shetland and Orkney. O cultivated V. tricolor cv. Forth (Markinch, Edinb., xii 1929). On  Viola spp. Tay (Blairgowrie, vii 1941, RWG Dennis) and Viola spp. in doubt Tay (Prior Muir, St Andrews, 17 & 19 x 1935; in garden Blairgpwrie, 29 v 1950; in garden St Andrews, 2 ix 1968 & on cv. Pickering Blue, St Andrews, 16 ix 1969; Arbroath, v & ix 1944, U. Duncan; Glen Doll, 15 vii 1944), Solway (Sands of Tornie 24 v 1961), Sutherland (Loch Shin, Lairg, 21 viii 1944) and Ross (Durness, 25 viii 1983). On garden plant origianlly collected in Scottish Mts., in Forth (Clermiston Edinb., 7 v 1980). Includes Pucccina depauperatus in Foister (1958) recorded on nursery stock of Viola, violets and pansies. Also under this name for Tweed (Peebles, vi 1935, J.A.Mcdonald). Old Greville collections in E on Viola canina labelled ‘in Roslin & Granton Woods, (Edinb.) May & June 1821’; also ‘nr. Edinb., August’. Also as Uredo violarum  Greville collectionm Miss Boswell, vi 1821 prob. Balmutto. As P. violarum and recorded from all Watersheds in Stevenson except Sutherland and as Aecidium violae from all Watersheds except Sutherland, Solway, Argylle and Ross.

TFTaDMSSoCAR  P. vigae-aureae On Solidago virgaurea from Skye, Orkney and Shetland. Known from Forth (Flockton, Edinb., 30 vi 1955), Moray (Ben Wyvis, 365m., 15 ix 1956) and Argyll (Ben Nevis, 15 ix 1956).

As Puccinia clandestina on Scabiosa succisa (=Succisa pratensis) from Tay & Argylle in Stevenson must be faulty record on such a host or it is possible the error as is outlined for material from Cornwall by Wilson & Henderson being a misidentified host and young rust-structures, especially as this taxon is recorded from Tay at 823m. in Stevenson.

Uromyces

TFTaDMSSoCAR U. acetosae On Rumex acetosa Moray (II/III Loch Duntelchang, viii 1947, J.A.Mcdonald; Abernethy, ix 1925, N.L. Alcock), Clyde (Aryshire no further details, ix 1926, N.L. Alcock), Sutherland (West coast, Echachilles Bay, 26 viii 1944, R.W.G.Dennis), Forth (Lumphuran, ix 1922; I Isle of May, 29 iv 1912), Tay (O,I/II/III Loch Lubnaig, Callander, 1939; Munlochy vii 1942 U.Duncan; Fofar, viii 1902), Argyll (II/III Glen Branter, v 1929), Ross (Inchnadamph, 1 viii 1954, J.A.Mcdonald; II/III Glen Lochy 16 vii 1962, D.M.Henderson,), Solway (Stoer, Dumfries, viii 1951, J.A.,Mcdonald) and Tweed (Innerleithen, v 1923, M.Wilson). Also Shetland, Mid Yell and Orkney, incl. Birsay, South Ronaldsay & Hoy.  On R. acetosella Tay (Glen Dochart, 10 viii 2007, R.Watling).

On R. obtusifolius Tay (Blairgowrie, 5 vi 1952, D.M.Henderson). Records on this host for Lewis and Mull; also Shetland and Orkney.

TFTaDMSSoCAR U. airae-flexuosae On Deschampsia flexuosa; some records as Aira. Known from Tay (Abercairny Wd., Crieff, 17 viii 1889, J.Trail; Kinnoul Hill, 12 ix 1944, R.W.G.Dennis), Forth (Corstorphine Hill, viii & ix 1922; Edinburgh ix 1929), Moray (Beauly, no further data), Solway (Moniavie, Thornhill, ix 1931) and Dee (Brechin G.B.Wallace, 21 vi 1946), Clyde (Dougalstoun, 9 ix 1959,D.M.Henderson; Loch Lomond, Balloch Pk., 11 ix 1959, D.M.Henderson). Known from Skye, South Uist and Mull.

TFTaDMSSoCAR U. aloes Undoubtedly an introduction. On Aloe glauca and 2 un-named Aloe spp. from Natal and one from Swaziland in Forth (RBGdn., Edinb., ix 1938).

TFTaDMSSoCAR U.ambiguus On Allium scorodoprasum Dee (II/III Innerbervie, 23 v 1956, D.M.Henderson), Solway (II/III Gatieston, Kirkcubridgthshire, 1970; III Galloway Hse., Wigtownshire, 12 vi 1970) and Moray (Dyke, Forres, 20 vii 1957). On A. schoenoprasum from Dee (Aberdeen, 30 vi 1948). 

TFTaDMSSoCAR U. anthylidis On Anthylis vulneriana Dee (Aberdeen Links, 1822 Greville collection ?; Gardenstown, Bannf., 9 viii 1969), Tay (Kinkell Braes, St Andrews, x 1946 & St Andrews, 12 x 1935, J.A.Mcdonald), Forth (Boghall, Edinb., vi 1934, J.A.Mcdonald; Redhead, Arbroath, v 1944, U. Duncan). Also recorded from Tresnish Isles and from South Ronaldsay, Orkney and records from South Uist, Vatersay, and Mull. In Foister (1958) recorded as U. jaapianus on Trifolium dubium. Recorded on Anthyllis in Stevenson as Puccinia fallens for Moray (Findhorn & Aviemore, Rev. Keith).

TFTaDMSSoCAR U. ari-triphylli On Peltandra virginea ‘Arrow Arum’ as Aecidium importatum for Forth (RBGdn., Edinb., 1924).

TFTaDMSSoCAR U. appendiculatus Undoubtedly introduced. On Phaseolus from Forth (purchased from Sainsbury’s, Edinb., 13 i 1997, Marissa Main). Also old collection in E as Puccinia phaseoli, poss. Greville material.

Recorded from Tay in Stevenson.

TFTaDMSSoCAR U. armeriae On A. distantifolia from Forth (RBGdn., Edinb., 2 v 1952 & 26 vi 1962).

On Armeria maritima complex. Known from Tay (Ben Hearkgarnish, 467m., 28 vii 1952; Boarshill, Fife, iv 1971 Arbroath, v 1953, U. Duncan;; I Ben Lawers 1954, grown in RBGdn., Edinb. ii 1955), Forth (Gullane, v 1931;  Aberlady, 5 iv 1953; O?/I Tantallon, 31 iii 1953, D.M.Henderson; I on var. alpina, RBG Edinb., 31 iii 1952 & II 10 & 18 vii 1951 & I/II on var. elongata RBG Edinb., 18 vii 1951; I/II on var. typica  = var. leuchaena xi 1924, 13 vi 1924, 31 iii 1952,10 v 1952 & 26 vi 1912), Ross (I Loch Kishorn, 7 iv 1958 & Loch Carron 2 iv 1960, D.M.Henderson; Beinn Eighe, 457m., 18 vii 1955, D.M.Henderson), Tweed (nr. Berwick, ix 1922, Mr Orr) Argyll (Beinn Bhan, summer 1959) and Clyde (Loch Strivnn 12 vi 1951; Ballantrae, ix 1929). Known from Lewis, Barra, Mingulay, Coll, Raasay, Skye, Mull, Geometra, Colonsay, Islay Garvellochs, Jura, Gigha, Kintyre and Orkney. Also known from Arran and as U. limonii on same host. On A. plantaginea from Forth (III RBGdn., Edinb., 18 vii 1951 & 31 iii 1952). As Uredo statices on A. maritima in Stevenson Add 4 and for Dee (Aberdeen, Buchanan White) and Tay in Add. 8; also Forth (Edinb. Greville).

TFTaDMSSoCAR U. behensis On Silene maritima Solway (I Kintyre, viii 1936; Gatehouse of Fleet, ix 1931), Mull and Orkney. Single record on ‘S. inflata var. puberula’ from Forth (Aberlady, R.McStark 1830.

TFTaDMSSoCAR U. beticola All records as U. betae. On Beta vulgaris and cultivars. Known from Tay (Boarshill, 20 x 1985; Fife, Dr.E Gray), Dee (Aberdeen, x 1945), Forth (II/III Crail, 3 v 1985 Mr Ali; Mungails, Falkirk, xii 1945, E. Gray; Inverleith Park, Edinb., 7 xi 1991, cv. Perpetual, M. Dickson), Tweed (Ayton Law, Berwickshire, 19 x 1925), Moray (Nairn, 9 x 1925, N.L. Alcock). Also on Spinach Beet Jura, 13 iv 1949. I,II,III recorded on Sugar Beet, Mangel, Fodder Beet, Red Beet and Seakale between 1928 & 1958 in Foister (1958). Record in Stevenson from Tay (Glamis Den) on Rumex is surely  misidentification.

TFTaDMSSoCAR U. chenopodii On Suaeda maritima var. flexilis. Known only from Forth (Tyninghame, 8 ix 1959 & 3 ix 1960, D.M.Henderson).

TFTaDMSSoCAR U. croci Single collection on Crocus albifolius from Forth (Goldenacre, v 1956, B.L.Burtt), although noted in Henderson (2000) as being on C. vernus in Scotland. 

TFTaDMSSoCAR U. dactylidis On Dactylis glomerata from Forth (III Balerno, 12 ix 1960; II/III Pentlandfield 30 ix 1959; Kilconquhar Loch, Fife, 28 v 1966), Tay (Ballanluig, vii 1915; nr. Stanley, vii 1930) and Clyde (II/III Dalrymple, ix 1933); also Shetland. ‘Dactylis’ form on Mull known on Ranunculus acris, R. bulbosus and R. repens

Records include Uredo festucae and Uromyces poae Solway (Gatehouse of Fleet, viii 1931), Clyde (Cadzow Pk., Hamilton, 14 ix 1959, D.M.Henderson). On Festuca ovina Argyll (Mallaig, on, viii 1929) and Ross (Beinn Eighe, on F. ovina, 365m., 19 vii 1955, D.M. Henderson). Uredo festucae I Solway (Langholm, on F. ovina, 30 ix 1951, D.M. Henderson). On F. rubra  Tweed (nr. Berwick, ix 1922, Mr Orr). On F. viviparum Tay (Caenlochan,30 vi 1952, D.M. Henderson). Known from Shetland II/III Festuca spp. On P. nemoralis Mull. On P. pratensis Tay (Blairgowrie, 16 vii 1962, D.M.Henderson), North Uist and Shetland. On P. trivialis Tay (Rae Loch, Blairgowrie, 30 vi 1952 & 10 viii 1953; Lade Brae, St Andrews, 10 viii 1966) and Forth (II/III Kirknewton, 4 x 1959; Hermitage of Briads, Edinb., 9 x 1982), Orkney and Rousay. Known from Shetland on Poa spp. Poa form on Ranunculus ficaria known from Lewis, Harris, South Uist, Eriksay, Barra, Vatersay, Skye, Rhum, Mull, Gigha, Kintyre, Shetland and Orkney. 0/I On R. acris Orkney and Forth (Roslin, 2 v 1952). On R. ficaria as Uromyces poae Ross (Kilmuir, Ross, v 1942, U. Duncan), Tay (Balmerino, 9 v 1936, J.A.Mcdonald), Tweed (Peebles, iv 1932, J.A. Mcdonald), Forth (Edinb., v 1931; Raith est. Kirkcaldy, 6 v 1951; Elie, 4 iv 1930; Vogrie, 22 v 2006, R.Watling; Midhope Burn, West Lothian, 28 iv 1940, R.W.G.Dennis; Roslin, N.L.Alcock, iv 1926; Fife no further data v 1933; Dalkeith, Eskbank, 12 iv 1953; Dunbar, vi 1936; Arbroath, v 1944, U. Duncan), Tweed (Dawyck Bot. Gdn., 5 v 1985, R.McBeath) and Argyll (Loch Long 14 v 1950); also records from Mull, Gigha and Islay. On R. repens, Scotland, no further details, N.L. Alcock, Forth (Kirkliston, 7 vi 1934, J.A.Mcdonald; Grange Rd., Edinb., 16 v 1966), Tay (Glen Shee, 14 ix 1957; Blairgowrie, 16 vii 1962, D.M.Henderson; Rae Loch, Blairgowrie, 30 vi 1952 & 10 viii 1953), Tweed (Peebles, iv 1931, J.A. Mcdonald).

On R. scleratus from Tay (Fingask Loch, Blairgowrie, 10 v 1953, D.M.Henderson) and Shetland. On Ranunculus spp. Forth (Cove, Cockburnspath, 13 v 1959, R.W.G.Dennis), Solway (Newton Stewart, ii 1945), Tay (Kittocks Den, St Andrews, 2 v 1936, J.A.Mcdonald). There is a strange record on Tussilago farfara from Stirling University 6 x 1968, D.M.Henderson. Old collection as Aecidium confertum on Ficaria ranunculoides = R. ficaria ex Herb. Edinb. Univ. possibly Greville or Menzies and also under same name and on same host in Greville Herb. in E and noted as ‘v-vi 1821’. Also coll. from Lockerbie from Sir W. Gardiner in Greville coll. or Menzies. Probably a proportion of the records under Aecidium ranuculearum in Stevenson from all Watersheds except Sutherland, Ross & Argylle belong here.

TFTaDMSSoCAR U. dianthini. Early records as Puccinia caryophyllinus. Known on cultivated Carnations (Dianthus caryophyllus x plumarius ) Forth (Haddington, iv 1953, A.E.W. Boyd; Dobbies Gdn. Centre, Edinb., 14 iii 1928), Clyde (Glasgow iii 1935). Also on Dianthus barbatus from Forth (Edinb. Parks dept., 23 xii 1979). Recorded by Foister (1958) on nursery stock.

TFTaDMSSoCAR U. ervi On Vicia hirsuta. Known from Tay (Laze Mills, Angus, 22 vi 1964, D.M.Henderson; Strathkinness, 16 ix 1931, J.A.Mcdonald; Lade Braes Nursery, St Andrews, 16 ix 1939 & 1 vii 1946, J.A.Mcdonald; Abroath, Waulkermills, U. Duncan, ix 1944) and Forth (East Neuk, Fife, 18 & 19 iv 1948, A.Angus).

TFTaDMSSoCAR U. fallens On Trifolium pratense. Known as U. trifolii from Forth (Canal bank, Edinb., 1931; ex East Craigs) and from North Uist (Tingharry, 11 viii 1968), Harris, Barra, Vatersay, Coll, Raasay, Mull, Colonsay, Islay, Skye, Orkney  and Shetland. On T. pratense in North East and West Central Scotland according to Foister (1958) under U. trifolii.

TFTaDMSSoCAR U. ficariae on Ranunculs ficaria Tay (Arbroath, v 1944, U. Duncan; Balmerino, 9 v 1936, J.A Mcdonald; Torryburn, Fife, 23 v 1902; Bot Gdn,, St Andrews, spring 1949), Forth (East Calder, 6 iv 2008; Balerno, vii 1936; Dunbar, vii 1934; Pathhead, Midlothian, v 1923, G. Wallace; Cramond Brig, iv 1921; Roslin 2 & 9 v 1952, on bulbiferous form of R. ficaria, D.H.Henderson & 15 iv 1945, M. Noble; Eskbank, 12 iv 1953; Midhope Burn, West Lothian, 28 iv 1940, R.W.G. Dennis; Kirkliston, 8 v 1949, D.M.Henderson; Elie, Fife, 21 iv 1930, N.L.Alcock; Edinburgh, v 1931), Argyll (Ben Lui, on montane ledges, 9 vii 1995, R Watling; Loch Sunart, 12 v 1984, D.M.Henderson; Loch Long 14 v 1950) and Clyde (Chatelhaerult & Falls of Clyde). Only known in Hebrides from Barra. Also known from single collection probably from Greville in E. In Stevenson recorded from Tweed, Forth, Tay and Dee; Moray added in Add.1.

TFTaDMSSoCAR U. geranii On Geranium cinereum Skye. On G. natalense var. album Solway (Logan Bot. Gdn., 22 vi 1970)On G. pratense, Islay, Forth (Aberlady, v 1952; Longnnidry, East Lothian, v 1936), Tay (Fife Coast 26 ix 1995 III; St Andrews Lade Braes, 11 x 1935 & 23 vii 1965-10 viii 1966; East Cliff St Andrews 2 v 1936; St Andrews 17 vii 1966 II & 25 ix 1966; Blair Atholl 14 vii 1960, D.M.Henderson; Ben Vrackie 10 viii 1975). On G. pyreniacum Forth (Craigmillar Castle, Edinb., 2 ix 1962; Dalmeny, Edinb., 6 ix 1968 and as U. kabatianus same locality vi 1925 & vii 1934), Tay (Murthley, ix 1925 N.L.Alcock; St Andrews II/III; Glen Lyon, O-II, 10 vi 1963, D.M.Henderson; nr. Montrose & Carlingheugh Bay, both viii 1944, U.Duncan) and Tweed (St Boswells, 13 vii 1941). Also Strathbroom 2 viii 1942, R.W.G.Dennis. On G. rotundifloium Forth (St David’s Bay, 11 x 2003, R.Watling). On G. sylvaticum Tweed (Duns Castle, 19 vi 1982, A.P.Bennell; Earlston, viii 184; Currie, 2 vi 2002, R.Watling; Tempell, Union Canal, 27 v 1975; Cramond Ferry, Edinb. 13 ix 1962), Solway (Langholm, 10 ix 1951, D.M.Henderson); also Sithnobroan, 2 viii 1942, R.W.G.Dennis), Tay (Aberfeldy, 27 vi 1981), Moray (Loch Duntelchang, 1936 J.A.Mcdonald) and Lismore.

Three old collections in E as Uredo prob. ‘ about Edinb. Sept. on G.pyreniacum’ prob. about 1822 and ‘Kirkcaldy on G. pratense’ and on G. sylvaticum ‘MacBridge about 1822; all Greville. Also Cooke Exsic. 60 2nd edition as Trichobasis from Roslin and as Dothidea gerani Banks of Clunie, Carthton, vii 1845. In Stevenson recorded from Tweed, Forth & Tay and as Aecidium gerani on G. sylvaticum from Tweed and Tay; Dee added in Add. 8.

TFTaDMSSoCAR U. limonii On Statice Forth (RBGdn., Edinb.) as U. statici Grev.

TFTaDMSSoCAR U. lineolatus On Scirpus maritimus Solway (Glen Luce bay, Wigtonshire, 31 viii 1940), Forth (Tyninghame, vi 1939, & 2 x 1954, D.M.Henderson), Tay (Tentsmuir, 30 viii 1947) and Gigha R.W.Dennis 12 v 1981.

f.sp. glaucis-scirpi On Glaux maritima Fife – Forth. f.sp. scirpi-oenanthe-crocati On Oenanthe crocata Islay and Gigha.

TFTaDMSSoCAR U. minor On Trifoloium incarnatum from Moray & Argyle in Foister (1958) as U. trifolii.

TFTaDMSSoCAR U. muscari On Scilla campanula Tay (Dundee, in garden, 11 vi 1952). On S. nutans Tay (Murthly, vi 1930; Arbroath U. Duncan. V 1944; St Andrews, v 1937), Moray (Black Isles, v 1942) and Forth (RBGdn., Edinb., v 1936). On S. verna Southness, Shetland, 14 vi 1981. On Muscari polyanthum Forth (RBGdn., Edinb., vi 1914). Included here are records of U. scillarum on Hyacinthus non-sciptus (= Endymion & Scilla). Records from Ross (Slumbay on bulb scales, 2 iv 1961, D.M. Henderson), Tweed (Dawyck), Clyde (Inchcailloch, Loch Lomond, 17 v 1974, D.M.Henderson), Forth (Blinkbonny Edinb., and on flower scapes 25 vi 1989; RBGdn, Edinb.,  v 1961 R.Watling; on Scilla hispanica x non-scriptus Forth (Edinb., 6 vi 1966 & 20 vii 1984 on hybrid bluebell; Torryburn Wd., 23 v 1982), Tay (Dores Foyers roadside 4 v 1934 & East cliffs, 2 v 1936 both St Andrews, JA Mcdonald,; Boarshill, A.Angus, 181 v 1966; Lade Brae, St Andrews, M. Stewart, 2 vi 1966; Long Walk, St Andrews, A. Angus, 25 v 1979), Clyde (Falls of Clyde & Chatelhaerult), Dee ( Summerhill M. Wilson, 23 iii 1917) and Argyll (Crinan, 25 v 1963). Known from South Uist, Barra, Islay, Raasay, Skye, Mull, Liung, Gigha, Arran, Kintyre and Shetland. Recorded by Foister (1958) on nursery stock of Muscari racemosum. Also on Brimeura fastigiata Forth (RBGdn., Edinb. on specimen brought from Corsica, 21 iv 1971). On Bellevalia Forth (RBGdn., Edinb., D.M.Henderson). As Uromyces concentrica recorded in Stevenson from Tay and Moray.

TFTaDMSSoCAR U. pisi-sativae On Astragalus danicus Tay (Dunes, St Andrews, 6 & 8 x 1948; Sands of Barry, U. Duncan, x 1952 II only, ix 1947 & 22 x 1952; Dunes, St Andrews, 6 x 1948). On Genista anglica Moray (Loch Loy, Nairn, 25 ix 1955, D.M.Henderson; nr. Inverness, ix 1934, J.A.Mcdonald; Loch Duntelchang 29 viii 1935, J.A.Mcdonald; banks of Caledonian Canal, 27 viii 1935 & viii 1946, J.A.Mcdonald last two records as Uromyces genistae-tinctoriae). On Lathyrus pratensis Tay (Tentsdmuir, A. Angus, 9 x 1965; Burn Nursery, St Andrews, 18 ix 1938), Forth (Crichton, 6 ix 1925, M. Wilson; Pathhead, Ford, 6 ix 1925, M.Wilson; East Saltoun, 10 ix 1961; Cove, II & III, viii 19030), Solway (Mull of Galloway, II & III, ix 1929; Stranraer, 14 viii 1943, R.W.G.Dennis), Tweed (Tynehead xi 1919, M. Wilson) and Colonsay. On Lotus corniculatus Vatersay and Arran; also Skye, Canna and Kintyre. Also as Uromyces loti Stravathir Rd, 14 x 1946, West Dunes, 14 x 1946 & St. Andrews dunes, 18 vii 1946, JA Mcdonald; Tentsmuir 120 x 1965) and Moray (Loch Dutelchaig, vii 1939), Sutherland (Dunnett Head, 24 viii 1983, B.J.Coppins), Tweed (Berwick ix 1922, M. Orr) and Clyde (Larg, Arran, 1941) and Torduff Reservoir, 28 vii 1935.

On Medicago lupulinus as Uromyces striatus Tay (Burns Nursery 18 ix 1938 & Strathie Rd., vii 1946, both St Andrews). Noted by Foister (1958) in Economic Plany Diseases. On Sarathamnus scoparius Moray (Banks of Caledonian Canal, viii 1946; Loch Duntelchang, 29 viii 1935, J.A.Mcdonald; Burghhead 7 viii 1959, D.M.Henderson; Inverness viii 1935) and Tay (Banks of Tay, Dunkeld, 18 ix 1959). On Ulex europaeus Moray (Loch Duntelchang, 29 viii 1935, J.A.Mcdonald). Probably as Uromyces sp. on Lotus corniculatus Moray (Aviemore, 24 vi 1964).

Old collection in E could be Greville on Pisum sativum. Recorded under U. fabae on Pisa sativum by Foister (1958); first recoded in 1939 and noted on commercial crops in 1951 & 1953 in North East and North.  I on Euphorbia cyparissis St Andrews no further data.

TFTaDMSSoCAR U. polygoni-avicularis On Polygonum aviculare Tay (St Andrews, no further data; Waukermill, Arbroath, U. Duncan, ix 1944; nr. Ballantrae, ix 1929), Clyde (S. of West Kilbride, 27 viii 1953), Forth (Corstorphine, 1 x 1965, D.M.Henderson; Edinb., viii 1936, JA Mcdonald; Gifford, 25 ix 1965; Musselburgh, xi 1925, N.L.Alcock; Longniddry 26 vi 1960; East Craigs, 8 vii 1941, M.Noble) and Dunrossness, Shetland, Tigharry, N. Uist, Tiree and Lismore. Old collection in E undoubtedly Greville labelled ‘about Edinb. Sept.’.  As Uromyces polygoni from Tay and Moray in Stevenson and in Add 3 for Dee.It is possible that the records in Stevenson as Puccinia vaginalium on P. aviculare from Tweed, Forth & Tay belong here; strange classification as P. vaginalium refers to Puccinia acetosae!

TFTaDMSSoCAR U. rumicis On Rumex crispus Forth (nr. Edinburgh, viii 1821 prob. Greville coll.; Aberlady, 24 viii 1952) and Ross (Kishorn Loch an Loin, 6 ix 1952. On R. conglomeratus Orkney 17 viii 1989. On R.obtusifolius Clyde (Rosshdu, 11 ix 1959, D.M.Henderson), Forth (Kirknewton 4 x 1959, D.M.Henderson; Banks of Canal, presumably Edinburgh, ix 1836 under Uredo), Tay (Waulkermill, Arbroath, U. Duncan, ix 1944; Comrie, 12 ix 1969, D.M.Henderson; Ballanluig, M.Wilson x 1921; Dunvina & 29 ix 1939 & Dura Dun 13 ix 1940 both St Andrews J.A.Mcdonald), Argyll (Ardgour, II & III 14 vi 1961), Tweed (Dawyck Bot. Gdn., 5 viii 2000), Solway (Gatehouse 14 viii 1952) and from Tiree, St Kilda, Skye, Mull, Colonsay, Islay, Kintyre, Shetland and Orkney. On R. sanguineus Clyde (Rosshdu, 11 ix 1959. Two old collections in E both prob. Greville, Foxhall, Edinb. and as Uredo ‘Banks of   Sept. 1936’. As Uromyces apiculosa on dock, clover etc.in Stevenson for Tweed, Forth, Tay, Moray; in Add 3 Dee added. Also said to be on Oxyria but see Puccinia oxyriae. On Ranunculus ficaria Forth (Balerno, 19 iv 1960, D.M. Henderson).

TFTaDMSSoCAR U. scrophulariae On Scrophularia nodosum Tay (Falskally, 6 viii 1957, D.M. Henderson) and Argyll (Inverawe, Taynuilt, vii 1921 & viii 1934). As Uromyces concomitans in Stevenson from Tay (Nora, Rev. Ferguson; Dunkeld, Buchanan White; Monikie, Rev. Ferguson) and as Aecidium scrophulariae for Tay. t is possible that the records as Puccinia scrophulariae in Stevenson from Tay (Fern, Rev. Anderson & Monikie, Rev. Ferguson) belong here.

TFTaDMSSoCAR U. scutellatus III on Euphorbia cyparissias. Single collection from Sutherland (Strathnaver, E.Boa, 24 vi 1979). Also collection by Joyce Pitt v 1982 not localised.

TFTaDMSSoCAR U. sparsus On Spergularia II & III Forth (nr. Port Seton, Leven, vi 1925).

TFTaDMSSoCAR U. trifolii On Trifolium repens known from Skye, Mull and Colonsay; also Orkney As Uromyces flectens Forth (East Craigs, vi 1929 & 24 viii 1940; Borthwick Castle, viii 1930; Corstorphine, iv 1930; Balerno, 17 vi 1954), Ross (Plockton, 20 vi 1955), Dee (Aberdeen, 18 vi 1959), Tay (Blairgowrie, vii 1930; Arbroath, U. Duncan, viii 1944; Loch Nesa, U.Duncan, ix 1942; Ballanluig, x 1921) and Clyde Loch Lomond, Balloch Pk., 11 ix 1959; Corsehill, Kilwinning, no further data), Argyll (Ardnamaurchan, 14 vi 1961), Skaill Bay and Yeasnaby, Orkney  and Skye. Record from Lochephail, N.Uist 9 viii 1968. Often recorded under U. nerviphilus known from Arran and Kintyre, Moray (Loch Duntelchaig, 29 viii 1935), Tay (Kinnell Brae St Andrews, M. Stewart, 8 viii 1966); also under this name on Trifolium repens from Lewis, Harris, North Uist, Barra, Tiree, Coll, Raasay, Skye, Rhum, Shetland, Eigg, Mull, Lismore, Islay, Kintyre, Orkney and Shetland. In Foister (1958) on T. repens in all areas except S West & S East.

As Aeciduum trifoliorum with description in Stevenson for Tay (Menmuir vi 1875 Rev. Anderson; Glen Clova); see Wilson & Henderson for confirmation of aecidia in this tricky group. Probably also as Uromyces sp. on T. repens Tay (Tentsmuir, 1 vii 1947).

TFTaDMSSoCAR U. trifolii-repentis On Trifolium repens in Foister (1958) and as Uromyces trifolii in Foister (1958) on T. hybridum cv. Aliske Clover. Records from Tay (Tentsmuir 1 vii 1947).

TFTaDMSSoCAR U. valerianae On Valeriana officinalis Forth (Pathhead, vii 1925, N.L.Alcock; Ratho, 22 vi 1978, A.P.Bennell; Aberlady, 24 v 1961 D.M.Henderson; Roslin, v 1912 M. Wilson; Lake Menteith, 12 ix 1959, D.M.Henderson; Colstoun Wd., East Lothian, BMS meeting), Tay (Blairgowrie, 10 viii 1953 & 5 vi 1952, D.M.Henderson; Ben Ledi, 1921 no further data; Forest of Alyth, 20 viii 1973, R.Watling; Tentsmuir, 10 x 1965, D.M.Henderson; Rae Loch, Blairgowrie, 11 vi 1950, D.M.Henderson; Monemish Loy, Tay 183m., 1 ix 1961, D.M. Henderson; Roscobie Loch, Fofar, viii 1944, U.Duncan; Banks of Tay, Dunkeld, 18 ix 1958, D.M.Henderson; Callender, vi 1936; Murthly Est., ix 1925, N.L.Alcock), Argyll (Taynult, Etive, vii 1921; I & II Tentsmuir, 10 vii 1955, J.A.Mcdonald, 9 x 1965, A.Angus & 19 viii 1966, M.Stewart), Dee (Loch Kinoul, Aberdeen, 6 ix 1975), Moray (Loch Duntelchaig, viii 1946, J.A.Mcdonald), Tweed  (Cothill, Berwicksh., J.P.Brennan, 8 x 1943; nr. Berwick, no further data) and from Skye, Eigg (vii-viii 1947), Islay, Mull (Glen Forsa & Torloisk), Arran and Orkney. In Stevenson as Lecythea and recorded from Tay and as Aecidium valerianacearum from Tweed and Tay; Moray added in App.1 and Argyle in Add 4 and Dee and Moray in Add. 8.

TFTaDSSoCAR U. vici-fabae On V. cracca Moray (Ballindalloch, ix `1937), Tay (Kingsway, Dundee x 1933) and Shetland. On Vicia faba Forth (Blinkbonny Gdns., Edinb., 7 viii 1990, 10 ix 2001 & 22 viii 2007; East Craigs 24 x 1941, R.W.G.Dennis; Foxcovert, Edinb., R.W.G.Dennis; Dunfermline, J. Maxwell; RBG Edinb., x 1939; Lasswade, x 1968, M. Noble), Tweed (Maxton, St Boswells, viii 1969, A.Hughes), Tay (Arbroath, ix 1944, U.Duncan; St Andrews 10 x 1971 J.A.Mcdonald & 12 x 1968 A.Angus; Parkhill, Boarshill 25 ix 1980 & 2 x 1971 A. Angus; Longforgan, Dundee, 12 viii 1936; Kingburns, St Andrews, ix 1958), Sutherland (Tongue II & III, viii 1931; Scourie, 20 viii 1944), Solway (Grange of Coll, Wigtown viii 1943). Rare on Broad bean in 1924 but recorded from South West, South East, West, South & Central Scotland; especially a problem in 1951, 1953 in North East and North; see Foister (1958).

On V. lathryoides Tay (Invery Angus, 22 vi 1964, D.M.Henderson). On V. sepium Tay (Dura Dun, 13 ix 1940; Glen Lyon 13 ix 1960), Argyll (Appin? M.Wilson), Clyde (Younger Bot. Gdn., Benmore, 14 ix 1959, D.M. Henderson) & Forth (Gladsmuir, I & II, 24 vi 1967, D.M.Henderson Kippen, 12 ix 1959, D.M.Henderson); also Benevenagh II & III, no further data and Mull. Holotype in E and 2 additional collections from Greville, ‘about Edinburgh’ and ‘Kingspark, Edinb.’ In Stevenson as Trichobasis fabae from Tweed, Forth, Tay and Moray and in Add 3 for Dee.

TFTaDMSSoCAR Var. orobi On Lathyrus montanus Moray (Loch Duntelchaig, viii 1933,1936, vii 3 1939, 21-30 vii 1939 & viii 1947; Aviemore, ix 1925 and as Lathyrus macrorhizus, Abernethy 9 v 1943), Tay (on Lathyrus acrorhizus, St Andrews), Forth (Lumphnan, Dunfermline, 21 vii 1922) and Mull. Old collection for Edinb. in E by Greville as Aecidium orobi. Also probably as Uromyces sp. Moray (Loch Duntelcraig, 23 v 1937, J.A.Mcdonald). Recorded as U. fabae on Orobus tubersous (= Lathyrus montanus) in Stevenson from Tweed, Forth, Tay and Moray; see discussion therein. As Aecidium orobae noted in Stevenson from Tay, Dee and Moray