Handshape Features:
Palm: [-compact, +flat]
Thumb: [+twisted, -extant, -arc, -contact]
Finger: [+index, -concave, +ulnar, +spread, +ring, -medio]
Orientation Features:
Palm: [-visible, +opaque]
Carpal: [-bilateral, -ante bilateral, +addressee, -signer, -sky]
Location Features:
[+ipsi, -contra, -high, -low, -close, -full]
Movement Features:
Direction: [-vertical, -horizontal, -to-fro, -vector, -symmetric]
Contour: [-straight, +circle, -semi-circle]
Dynamics: [-small, +large]