With the extinction of carnieri, the remaining geotters lost any resemblance to their ancient ancestors: they were no longer terrestrial otters and did not even remotely resemble them.
Today, only two very distinct groups have survived the extinction events that struck Antarctica: the sheardogs, whose bodies are adapted for running and derived from the ottofox, and the ragos, whose bodies are adapted for climbing and feeding mostly on plant material, descended from the terminal rago. From just two surviving species, the geotters have now diversified into around 30 species, with the radiation still ongoing. With the extinction of any small carnivorous species however, geotters have lost a great piece of biodiversity, filled now by several species of rodents, like the topungers.
The antarctic ottofox gave rise to a new group of sheardogs known as the wonderlonts, which have achieved one the highest degree of running adaptations ever seen among sheardogs: despite the undisputed running efficiency reached by beasters, wonderlonts also shows new unique adaptations, like the lost of one digit, which reduce friction while running and the partial fusion of metatarsal bones, which makes forelimbs sturdier and more capable of absorbing impacts.
Among the current largest species there's the open wonderlont (Vulpilutra honestus), adapted to the semi-open landscapes of the Great Depresseaon, feeding mainly on small vertebrates like burrowing rodents and bellydruggers, and only rarely on small stottmice. They are coyote-sized but with a narrower and more delicate skull, similar to a fox, helping them snatch rodents from their dens. Open wonderlonts are currently the only social and territorial wonderlont species, which live in breeding pairs with their first-year young, although partners usually feed alone and not in pack. Like all species in the genus Vulpilutra, they have a mostly red camouflage and a long, bushy tail that provides balance while pursuing prey. In southern populations, the tail is also useful for insulation against the cold ground during winter. An opportunistic feeder, the open wonderlont can also eat fruits, seeds, and carrion provided by larger predators, like borax.
Some open wonderlont species (Vulpilutra spp.) inhabit woodlands, but in these environments, their role as mesocarnivores is usually replaced by another wonderlont genus, Arboricanis, the largest member of which is the dense wonderlont (Arboricanis maculatum) mostly found in the shady montane forests of Transantarctic Mountains. While still retaining running adaptations from their ancestors, dense wonderlonts are trending toward sturdier bodies, using brute force to subdue prey. A rapid adaptation for their scansorial lifestyle was the evolution of retractable claws, which prevent their consumption while walking.
Thanks to strong limbs and long claws, dense wonderlonts are excellent climbers, preying on arboreal animals like nesting birds, ramos, opossumus, and reptiles. They are also large enough to take down relatively big prey, such as many stottmice species and even groundlifters' calves, which are usually attacked from the top of a branch. Dense wonderlonts are also more carnivorous than open wonderlonts, with over 80% of their diet consisting of meat.
They are solitary animals with a very strong social life: communication between neighbours is extremely important in the life of a dense wonderlonts, in order to reduce the risk of intraspecific competition. Encounters are usually peaceful, and any issue is usually solved with some growls without any clash.
Ragos, the second and last surviving group of geotters, are common prey for dense wonderlonts. These arboreal and mostly herbivorous mammals have also diversified into a wide range of forms, from scansorial to fully arboreal species. Despite competition with opossumus, Antarctica's varied environments prevented excessive niche overlap. Ragos species are generally larger than possumus, ranging from 500 grams to as much as 10 kilograms. They all have squirrel- or monkey-like bodies that help them move along branches. Their fingers are well developed and lack claws, ending instead in nails like those of primates, allowing a stronger grip on branches.
Tails come in various shapes but are always very long. The most remarkable tail's function is found in the gliding rago (Simionyx lutrinus), a small-sized rago with browny and white coloration that inhabits the semi-open environments of the Great Depresseaon. As an arboreal species, living in such open areas is dangerous, so the gliding rago has evolved a peculiar way of moving through wooded savannas: it glides using its tail. The caudal vertebrae are laterally compressed and covered in long fur, forming a parachute-like structure for gliding. The gliding system is still primitive, allowing little control in the air and permitting slow glides of no more than 20 meters between trees. However, with time, this innovative locomotion will surely become more refined. The development of a tail instead of a patagium as a way of gliding has several benefits, like maintaining the functionality of limbs. In case a gliding rago fall on the ground, it can gallop at high speed to the next tree.
Among ragos, the gliding rago is one of the most omnivorous, with about half of its diet consisting of plant material and the rest from animal sources, mostly invertebrates. Its strong canines and incisors act as scrapers to uncover bugs and larvae hidden beneath bark, which are then extracted from their tunnels with long fingers. On rare occasions, gliding ragos have been observed ambushing prey such as small possumus species or resting insects, launching airborne attacks from the trees.
With increasing heat and dryness in Antarctica due to tropicalization, the gliding rago’s adaptations for life in fragmented woodlands will likely prove very advantageous in the long run. Yet, as history has shown, the future is not always easy to predict.