The rostrid expansion

With the increase and diversification of the austral forest, rostrids have drastically expanded their range from the few refuges of the Middle Lentocene. Despite having a low species diversity, anatomical and ecological differentiations are very significant in each species. 

Rostrids tend to prefer humid forests with lots of mosses and lichens, which comprise most of their diet. The foliage has become rarer and rarer in their alimentation, due to increasing competition with ducktails, forcing them to move in an even more specialized niche. While lignin digestion was the key to their success and their actual persistence, long and inefficient assimilation make this diet insufficient for their diet, especially during winter when energetic food is scarce. During summer and autumn, their diet comprises a large number of hard fruits, making them good seed dispersers: lots of plant species have since started symbiotic interactions with these large birds, especially austral oaks. In the near-temperate areas of Belgrano, horned rotbills (Silvavis bryophagus) are the main rostrids. They possess a large keratinous horn on their head, mostly used as a display. They are very territorial; males possess a home range of up to 5 km2 (1,9 sq mi), which covers the smaller territories of females. They spend most of their life alone, except during the breeding period, when males of neighboring territories clash each other for female possession. They rarely use their beaks in a fight, since they could easily break the neck of the opponent. After reproduction, females create a large nest, where they'll lay 8-12 eggs.

At the opposite part of Polarica, between Aurora forest and Amery Bay, Aurora debarkers (Wilkesornis humidus) have evolved a more docile behavior. Still being mostly solitary, juveniles create small social groups that separate when reaching adulthood. This rostrid has adapted to eat pine cones, since conifers are the dominant tree association of their distribution.  Feet are short and strong, used for digging up roots and tubers.

Two rostrid lineages are more divergent from the ancestral rostrid body plan. The tree mossbrusher (Minorampho scandus), a descendant of the dwarf mossbrusher, is one of them.
It's the most widespread species of rostrid and also the most abundant, numbering several millions. It's a very small species with a partially arboreal lifestyle. Strong legs are both useful for running in the forest undergrowth and to climb large trees, usually at no more than 5 meters of height. The beak is also used as a grasping hand, giving to this rostrid a sort of brachiation locomotion. The second digit of its feet is also partially prehensile, to better cling on branches. Arboreal movements are slow and precise, to avoid the risk of falling. Its mixed lifestyle enhances this bird to feed abundantly on leaves with virtually no competition from ducktails. They live in small family groups, with no more than 10 individuals. 

While mossbrusher is typically a forest-dweller, the way larger and taller headbutt rotbill (Gigarampho savanicus) lives in the large savannah-like habitats of the Scrubring. They are the most cursorial rostrids, capable of reaching 60 km/h (37.28 mph). Legs are their main offensive weapon, capable of striking deadly kicks. Low shrubs and mosses are the main food of this large rostrid, which can eat as much as 4 kg of plant matter daily. Like all rostrids, headbutt rotbills can't digest grass efficiently; still, these C4 plants are eaten ordinarily for vitamin requirements.
Their names are derived from the enlarged frontal bone of their skull. However, this isn't used for headbutting rivals; instead, it assists them in navigating through shrubs and thickets as they search for the best buds and leaves.