Family NITIDULIDAE Latreille, 1802

+ = 化石属・種 / Fossil

Series CUCUJIFORMIA

Superfamily CUCUJOIDEA Latreille, 1802

Family NITIDULIDAE Latreille, 1802

I. Subfamily CALONECRINAE Kirejtshuk, 1982

•   Calonecrus Thomson, 1857

II. Subfamily MAYNIPEPLINAE Kirejtshuk, 1998

• Maynipeplus Kirejtshuk, 1998

Carpophilinae-lineage

III. Subfamily EPURAEINAE Kirejtshuk, 1986

Tribe Epuraeini Kirejtshuk, 1986

• +Baltoraea Kurochkin et Kirejtshuk, 2010

• +Crepuraea Kirejtshuk in Kirejtshuk et Ponomarenko,1990

• Amedanyraea Kirejtshuk et Pakaluk, 1996

• Amystrops Grouvelle, 1906

Subgenus Amystrops Grouvelle, 1906

Subgenus Mandipetes Kirejtshuk, 1997                                        

• Baloghmena Kirejtshuk, 1987

• Ceratomedia Kirejtshuk, 1990

• Ecnomaeus Erichson, 1843

• Epanuraea Scudder, 1892

• Epuraea Erichson, 1843

Subgenus Africaraea Kirejtshuk, 1989

Subgenus Aphenolia Reitter, 1884

Subgenus Apria Grouvelle, 1919                                            

Subgenus Blackburnaea Kirejtshuk et Kvamme, 2001

Subgenus Ceroncura Kirejtshuk, 1994

Subgenus Dadopora Thomson, 1859

Subgenus Epuraea Erichson, 1843

  Subgenus Epuraeanella Crotch, 1874

Subgenus Haptoncus Murray, 1864

Subgenus Haptoncurina Jelínek, 1977

Subgenus Horniraea Kirejtshuk et Pakaluk, 1996

Subgenus Marinexa Kirejtshuk, 1989

Subgenus Micruria Reitter, 1874

Subgenus Ommoraea Kirejtshuk, 1998

Subgenus Orthopeplus Horn, 1879

Subgenus Polinexa Kirejtshuk, 1989

Subgenus Strophoraea Kirejtshuk et Kvamme, 2001

• Grouvellia Kirejtshuk, 1984

• Mystronoma Kirejtshuk, 1990

• Parepuraea Jelínek, 1977

• Trimenus Murray, 1864

Subgenus Trimenus Murray, 1864

Subgenus Schawalleria Kirejtshuk, 2017

• Tritesus Heller, 1916

• Platychorina Grouvelle, 1905

• Stauromenus Kirejtshuk et Kvamme, 2001

Tribe Taenioncini Kirejtshuk, 1998

• +Nitoraeopsis Kirejtshuk et Nel, 2018

• Carpocryraea Kirejtshuk, 1998

• Eutaenioncus Kirejtshuk et Kvamme, 2001

• Raspinotus Kirejtshuk, 1990

• Taenioncus Kirejtshuk, 1984

• Taeniolinus Kirejtshuk, 1998

IV. Subfamily CARPOPHILINAE Erichson, 1842

• Caplothorax Kirejtshuk, 1997

  Subgenus Caplothorax Kirejtshuk, 1997

  Subgenus Plapennipolus Kirejtshuk, 1997

• Carpophilus Stephens, 1829

Subgenus Askocarpolus Kirejtshuk, 2008

Subgenus Carpophilus Stephens, 1829

Subgenus Ecnomorphus Motschulsky, 1858

Subgenus Gaplocarpolus Kirejtshuk, 2008

Subgenus Megacarpolus Reitter, 1919

Subgenus Myothorax Murray, 1864

Subgenus Semocarpolus Kirejtshuk, 2008

• Ctilodes Murray, 1864

• Nitops Murray, 1864

Subgenus Nitops Murray, 1864

Subgenus Urocarpolus Kirejtshuk, 1997

• Loriarulus Kirejtshuk, 1987

• Vulpixenus Kirejtshuk, 1990

• Urophorus Murray, 1864

Subgenus Urophorus Murray, 1864

Subgenus Anophorus Kirejtshuk, 1990

V. Subfamily AMPHICROSSINAE Kirejtshuk, 1986

• Amphicrossus Erichson, 1843

Nitidulinae-lineage

VI. Subfamily PROMETOPIINAE Böving and Craighead, 1931 

• Platychora Erichson, 1843

(Prometopia-complex)

• +Palaeometopia Kirejtshuk et Poinar, 2007

• Parametopia Reitter, 1884

• Prometopia Erichson, 1843

(Megauchenia-complex)

• Axyra Erichson, 1843

• Megauchenia Macleay, 1825

• Megaucheniodes Audisio et Jelínek, 1993

• Pseudoplatychora Grouvelle, 1890

• Taraphia Audisio et Jelínek, 1993

VII. Subfamily MELIGETHINAE Thomson, 1859

•  Acanthogethes Reitter, 1871

• Afrogethes Audisio & Cline, 2009

• Anthystrix Kirejtshuk, 1981

• Aristogethes Audisio & Cline, 2009

• Asterogethes Audisio & Cline, 2009

• Astylogethes Kirejtshuk, 1992

• Bolbocerogethes Audisio & Cline, 2009

• Boragogethes Audisio & Cline, 2009

• Brassicogethes Audisio & Cline, 2009

• Chromogethes Kirejtshuk, 1989 

• Clypeogethes Scholz, 1932

• Cornutopria S. Endrödy-Younga, 1978

• Cryptarchopria Jelínek, 1975

• Cyclogethes Kirejtshuk, 1979

• Fabogethes Audisio & Cline, 2009

• Genistogethes Audisio & Cline, 2009

• Horakia Jelínek, 2000

• Idiogethes Kirejtshuk, 1977

• Indogethes Audisio & Cline, 2009

• Jelinekigethes Audisio & Cline, 2009

• Kabakovia Kirejtshuk, 1979

• Lamiogethes Audisio & Cline, 2009

• Lariopsis Kirejtshuk, 1989

• Lechanteuria S. Endrödy-Younga, 1978

• Lucanopria Audisio & Cline, 2009

• Meligethes Stephens, 1830 

• Meligethinus Grouvelle, 1906 

• Microporodes S. Endrödy-Younga, 1978

• Microporum C. Waterhouse, 1876

• Micropria Grouvelle, 1899 

• Neolariopsis Audisio & Cline, 2009

• Odontholariopsis Audisio & Cline, 2009

• Paleogethes Audisio & Cline, 2009

• Palmopria S. Endrödy-Younga, 1978

• Pria Stephens, 1830

• Rubiogethes Audisio & Cline, 2009

• Sagittogethes Audisio & Cline, 2009

• Sebastiangethes Audisio, Kirk-Spriggs & Cline, 2008 

• Stachygethes Audisio & Cline, 2009

• Tarchonanthogethes Audisio & Cline, 2009

• Teucriogethes Audisio, Sabatelli, Liu & Cline, 2020

• Thymogethes Audisio & Cline, 2009

• Xerogethes Audisio & Cline, 2009

• Xenostrongylogethes Audisio & Cline, 2009

VIII. Subfamily NITIDULINAE Latreille, 1802

Tribe Nitidulini Latreille, 1802

• Tetrisus Murray, 1864

Nitidula-complex

• Nitidula Fabricius 1775

• Omosita Erichson, 1843

Soronia-complex

• +Microsoronia Kirejtshuk et Kurochkin, 2008

• +Sorodites Kirejtshuk and Chetverikov, 2018

• Annachramus Kirejtshuk, 1995

• Amphotis Erichson, 1843

• Hisparonia Kirejtshuk, 2003

• Lobiopa Erichson, 1843

• Macleayania Kirejtshuk, 2003                                        

• Omosiphila Kirejtshuk, 1990

• Ornosia Grouvelle, 1899

• Pleoronia Kirejtshuk, 2003

• Sebastianella Kirejtshuk, 1995

• Soronia Erichson, 1843

• Stenoronia Kirejtshuk, 2003

• Temnoracta Kirejtshuk, 1988

• Ipidia-complex

• Ipidia Erichson, 1843

Subgenus Ipidia Erichson, 1843

Subgenus Hemipidia Kirejtshuk, 1992

• Perilopa Erichson, 1843

• Psilotus Fischer, 1829

• Taracta Murray, 1867

• Phenolia-complex

• Aethinodes Blackburn, 1891

• Cacconia Sharp, 1890

• Gaulodes Erichson, 1843

• Lasiodites Jelínek, 1999

• Ostomarcha Kirejtshuk, 2006

• Phenolia Erichson, 1843

Subgenus +Palaeoronia Kirejtshuk et Jenkins Shaw, 2023

• Plesiothina Kirejtshuk, 1990

• Stelidota Erichson, 1843

• Temnelytron Lawrence, 2019

• Ussuriphia Kirejtshuk, 1992

Perilopsis-complex

• Cratonura Reitter, 1875                                       

• Epuraeopsis Reitter, 1875

• Perilopsis Reitter, 1875

• Testudoraea Kirejtshuk, 1986

Aethina-complex

• Anister Grouvelle, 1901                                      

• Aethina Erichson, 1843

Subgenus Aethina Erichson, 1843

Subgenus Circopes Reitter, 1873

Subgenus Cleidorura Kirejtshuk et Lawrence, 1999

Subgenus Idaethina Gemminger et Harold, 1868

Subgenus Ithyra Reitter, 1873

• Lasiodactylus Perty, 1830-1834

Subgenus Lasiodactylus Perty, 1830-1834

Subgenus Lordyra Gemminger & Harold, 1868

• Neopocadius Grouvelle, 1906

Subgenus Neopocadius Grouvelle, 1906

Subgenus Brounthina Kirejtshuk, 1997

Subgenus Australaethina Kirejtshuk et Lawrence, 1999                                        

• Psilonitidula Heller, 1916

Pocadius-complex

• +Menatoraea Kirejtshuk et Nel, 2018

• +Omositoidea Schaufuss, 1891

• +Palaeolycra Kirejtshuk et Nel, 2018

• Aenigmoronia Lawrence, 2019

• Atarphia Reitter, 1884

• Australycra Kirejtshuk et Lawrence, 1992 

• Austronitidula Lawrence, 2019

• Bolivitoxus Kirejtshuk, 2009                                        

• Hebasculinus Kirejtshuk, 1992

• Hebascus Erichson, 1843

Subgenus Hebascus Erichson, 1843

Subgenus Teichostethus Sharp, 1891

• Hyleopocadius Jelinek, 1977

• Kryzhanovskiella Kirejtshuk, 2006

• Neothalycra Grouvelle, 1899

• Niliodes Murray, 1868

• Parapocadius Kirejtshuk, 2008

• Physoronia Reitter, 1884

• Pleuroneces Olliff, 1891

• Pocadiolycra Kirejtshuk et Leschen, 1998

• Pocadionta Lucas, 1920

• Pocadites Reitter, 1884

• Quadrifrons Blatchley, 1916

• Rixerodes Kirejtshuk et Lawrence, 1992 

• Tagmolycra Kirejtshuk et Leschen, 1998

• Thalycra Erichson, 1843

Subgenus Thalycra Erichson, 1843

Subgenus Pseudothalycra Howden, 1962

• Thalycrinella Kirejtshuk in Kirejtshuk et Leschen, 1998

• Thalycrodes Blackburn, 1891

Tribe Cychramini Gistel, 1848

• +Cychramites Wickham, 1913

• Axychramus Kirejtshuk, 1996

• Ceratochramus Kirejtshuk, 1986 

• Cychramus  Kugelann,  1794

• Kirejtshukostrongylus Audisio & Jelínek in Audisio, Mariotti, Jelínek & DeBiase 2001

• Oxystrongylus Reitter, 1911

• Xenostrongylus Wollston, 1854

Subgenus Strongylolasius Reitter, 1911

Subgenus Xenostrongylus Wollston, 1854

Tribe Mystropini Murray, 1864

• Anthepurops Kirejtshuk, 1996

• Anthocorcina Kirejtshuk, 1996

• Cychrocephalus Reitter, 1873

• Mystrops  Erichson,  1943

• Nitidulora Reitter, 1873

• Palmostrops Kirejtshuk et Jelínek, 2000

• Platychorodes Reitter, 1884

Tribe Cyllodini Everts, 1898

• +Cyllolithus Kirejtshuk in Kirejtshuk et Ponomarenko, 1990

• Cyclocaccus Sharp, 1891                                      

• Somatoxus Sharp, 1891

Cyllodes-complex

• Apsectochilus Reitter, 1874                                      

• Camptodes  Erichson,  1840

• Camptomorphus Grouvelle, 1908

• Carinocyllodes Leschen, 1999

• Ceramphosia Kirejtshuk et Kirk-Spriggs, 1996

• Coxollodes Kirejtshuk, 1987

• Cyllodes Erichson, 1843

• Cyllopallodes Lawrence & Kirejtshuk, 2019

• Cyllodesus Reitter, 1877

• Eusphaerius Sharp, 1891

• Lordyrops Reitter, 1875                                      

• Meoncerus Sharp, 1891

• Neopallodes Reitter, 1884

• Pallodes Erichson, 1843

• Viettherchnus Kirejtshuk, 1985 

Oxycnemus-complex

• Eugoniopus Reitter, 1884

• Gymnocychramus Lea, 1922

• Interfaxia Kirejtshuk, 2008

• Monafricus Kirejtshuk, 1995

• Oxycnemus Erichson, 1843

• Psilopyga LeConte, 1853

• Pycnocnemus Sharp, 1891

• Triacanus Erichson, 1843

Amborotubus-complex

• Amborotubus Leschen et Carlton, 2004                                     

Tribe Cychramptodini Kirejtshuk et Lawrence, 1992

• Cychramptodes Reitter, 1874

• Cylindroramus Kirejtshuk et Lawrence, 1992

• Miskoramus Kirejtshuk et Lawrence, 1992

Tribe Lawrencerosini Kirejtshuk, 1991

• Koryaga Kirejtshuk, 1990 

• Koryaginus Kirejtshuk, 1990

• Krakingus Kirejtshuk, 1990

• Lawrencerosus Kirejtshuk, 1990

IX. Subfamily CILLAEINAE Kirejtshuk et Audisio, 1986

Tribe Cillaeini Kirejtshuk et Audisio, 1986

• Adocimus Murray, 1864

Subgenus Adocimus Murray, 1864

Subgenus Belonotus Kirejtshuk et Kovalev, 2022

• Allenipeplus Kirejtshuk et Kovalev, 2016                                       

• Apetinus Scott in Sharp & Scott, 1908

• Brachypeplus Erichson, 1842

• Brittonoma Kirejtshuk et Kovalev, 2017

• Caledomus Kirejtshuk et Kovalev, 2017

• Campsopyga Murray, 1864

• Carpophilops Grouvelle, 1898

• Cillaeopeplus Sharp, 1908

• Cillaeus Laporte de Castelneau, 1835

• Colopterus Erichson, 1842

Subgenus Colopterus Erichson, 1842

Subgenus Cyllopodes Murray, 1864                  

• Cyrtostolus Sharp in Sharp et Scott, 1908

• Gonioryctus Sharp, 1878

• Grammorus Murray, 1868

• Halepopeplus Murray, 1864 

• Idosoronia Schaufuss, 1891

• Ithyphenes Murray, 1864

• Laferollaeus Kirejtshuk et Kovalev, 2022

• Leiopeplus Murray, 1864

• Liparopeplus Murray, 1864

Subgenus Liparopeplus Murray, 1864

Subgenus Grouvellepeplus Kirejtshuk, 2001

• Macrostola Murray, 1864

• Matthewsianus Kirejtshuk et Kovalev, 2022

• Notopeplus Sharp in Sharp et Scott, 1908

• Onicotis Murray, 1864

• Oniphenes Kirejtshuk et Kovalev, 2022

• Orthostolus Sharp in Sharp et Scott, 1908

• Paracillaeopsis Kirejtshuk, 2001

• Prosopeus Murray, 1864                                      

• Tokocillaeus Kirejtshuk, 2001

• Xanthopeplus Fairmaire, 1880                          

Tribe Conotelini Kirejtshuk et Kovalev, 2022

• Conotelus Erichson, 1843

incertae sedis

• Cillaeopsis Grouvelle, 1899

• Macrostolops Grouvelle, 1916 

• Teloconus Grouvelle, 1916

X. Subfamily CRYPTARCHINAE Thomson, 1859

Tribe Cryptarchini Th omson, 1859

• Cnips Philippi, 1864

• Cnipsarcha Jelínek, 1982 

• Cryptarcha  Shuckard,  1839

• Glischrochilus  Reitter,  1873

Subgenus Glischrochilus Reitter, 1873 

Subgenus Librodor  Reitter,  1884  

Subgenus Gymnoparomius Kirejtshuk, 1987 

• Homepura Broun, 1893 

• Kaszabena Kirejtshuk, 1987

• Paromia Westwood, 1850 

• Paromidia Reitter, 1873 

• Pityophagus Shuckard, 1839 

Tribe Platyarchini Kirejtshuk, 1998

• Amlearcha Kirejtshuk, 1987            

• Platyarcha Kirejtshuk, 1987

Tribe Arhinopini Kirejtshuk et Bouchard, 2018

• Arhinops Kirejtshuk et Bouchard, 2018            

• Arhinella Kirejtshuk, 1981

• Ceratarhina Kirejtshuk, 1981           

Tribe Eucalosphaerini Kirejtshuk, 1987

• Eucalosphaera Jelínek, 1978

  INCERTAE SEDIS

• +Protonitidula Zhao, Huang et Cai, 2022

• +Miophenolia Wickham, 1916

• +Oligamphotis Theobald, 1937

• Nodola Brethes 1925

• Prioschema Reitter, 1976

*基本的にKirejtshuk (2008), Kirejtshuk et Kavalev (2022) に従っているが，Meligethinae はAudisio, et al. 2009に，Cybocephalidaeの扱いはCline et al., に，Prometopinae はCline, 2014に従った．

**本HPではPrometopinae亜科を採用したため，Kirejtshuk (2008) に定義されたPrometopia-complexとMegauchenia-complexを削除，Megauchenia-complex に含まれていたTetrisus Murray は所属の再検討が必要．

References

• • Audisio, 2009.

  • Bouchard, P., Bousquet, Y., Davis, A.E., AlonsoZarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A.& Smith, A.B.T. (2011) Family-group names in Coleoptera (Insecta).

  • Cline, 2014

  • Jelinek, J., Carlton, C.C., Cline, A.R. & Leschen, R.A.B. (2009). 10.26. Nitidulidae Latreille, 1802. In: Handbook of Zoology 

  • Kirejtshuk, A.G. (2008) A current classification of sap beetles (Coleoptera, Nitidulidae). Zoosystematica Rossica 17(1): 107-122.

  • Kirejtshuk, A.G. & Kavalev, A.V. (2022 ) Monograph on the Cillaeinae (Coleoptera: Nitidulidae) from the Australian Region with comments on the taxonomy of the subfamily. Zootaxa, 5103 (1): 001–133.