Internal links: Angiosperms > Monocots > Asparagales > Iridaceae
External links: Angiosperms > Monocots > Asparagales > Iridaceae
Wikipedia links: Angiosperms > Monocots > Asparagales > Iridaceae
Other links:
Common name: Iris
Etymology:
Flowers:
x
Fruit:
x
Leaves:
x
Habit:
x
Habitat:
x
Distribution:
Species:
World: S, G
Australia: S, G
Additional notes:
General
Iridaceae is a family of plants in order Asparagales, taking its name from the irises, meaning rainbow, referring to its many colours. There are 66 accepted genera with a total of c. 2244 species worldwide (Christenhusz & Byng 2016[2]). It includes a number of other well known cultivated plants, such as freesias, gladioli and crocuses.
Members of this family are perennial plants, with a bulb, corm or rhizome. The plants grow erect, and have leaves that are generally grass-like, with a sharp central fold. Some examples of members of this family are the blue flag and yellow flag.
Name and history
The family name is based on the genus Iris, the largest and best known genus in Europe. This genus dates from 1753, when it was coined by Swedish botanist, Carl Linnaeus. Its name derives from the Greek goddess, Iris, who carried messages from Olympus to earth along a rainbow, whose colours were seen by Linnaeus in the multi-hued petals of many of the species.
The family is currently divided into four subfamilies but the results from DNA analysis suggest that several more should be recognised:
Subfamily Crocoideae is one of the major subfamilies in the family Iridaceae. It contains many genera, including Afrocrocus, Babiana, Chasmanthe, Crocosmia, Crocus, Cyanixia, Devia, Dierama, Duthiastrum, Freesia, Geissorhiza, Gladiolus, Hesperantha, Ixia, Lapeirousia, Melasphaerula, Micranthus, Pillansia, Romulea, Sparaxis, Savannosiphon, Syringodea, Thereianthus, Tritonia, Tritoniopsis, Xenoscapa and Watsonia. They are mainly from Africa, but includes members from Europe and Asia. The rootstock is usually a corm, they have blooms which sometimes have scent are collected in inflorescence and contain six tepals. The nectar is produced mostly in the base of the bloom from the glands of the ovary, which is where the flower forms a tube-like end. In some species there is no such end and the plant only provides pollen to pollinating insects. Members of this subfamily have the sword-shaped leaves typical of Iridaceae.
Subfamily Isophysidoideae contains the single genus Isophysis, from Tasmania. It is the only member of the family with a superior ovary and has a star-like yellow to brownish flower.
Subfamily Nivenioideae contains six genera from South Africa, Australia and Madagascar, including the only true shrubs in the family (Klattia, Nivenia and Witsenia) as well as the only myco-heterotroph (Geosiris). Aristea is also a member of this subfamily. It is distinguished by having flowers in small, paired clusters among large bracts, slender styles that are divided into three slender branches and nectar (when present) produced from glands in the ovary walls. The flowers are always radially symmetrical, with separate tepals (petals) and the rootstock is a rhizome.
Subfamily Iridoideae is distributed throughout the range of the family and contains the large genera Iris and Moraea. It is the only subfamily that is represented in North and South America. The species have flowers in solitary clusters among large bracts, styles that are often petal-like or crested and nectar (when present) is produced from glands on the tepals. Most species have separate petals and the rootstock is usually a rhizome or rarely a bulb. The flowers are almost always radially symmetrical. Bobartia, Dietes and Ferraria belong to this subfamily.
Ecology
Members of Iridaceae occur in a great variety of habitats. About the only place they do not grow is in the sea itself, although Gladiolus gueinzii occurs on the seashore just above the high tide mark within reach of the spray. Most species are adapted to seasonal climates that have a pronounced dry or cold period unfavourable for plant growth and during which the plants dormant. As a result, most species are deciduous. Evergreen species are restricted to subtropical forests or savannah, temperate grasslands and perennially moist fynbos. A few species grow in marshes or along streams and some even grow only in the spray of seasonal waterfalls.
The above ground parts (leaves and stems) of deciduous species die down when the bulb or corm enters dormancy. The plants thus survive periods that are unfavourable for growth by retreating underground. This is particularly useful in grasslands and fynbos, which are adapted to regular burning in the dry season. At this time the plants are dormant and their bulbs or corms are able to survive the heat of the fires underground. Veld fires clear the soil surface of competing vegetation, as well as fertilise it with ash. With the arrival of the first rains, the dormant corms are ready to burst into growth, sending up flowers and stems before they can be shaded out by other vegetation. Many grassland and fynbos irids flower best after fires and some fynbos species will only flower in the season after a fire.
The family has a very diverse pollination ecology. Most species are pollinated by various species of solitary bees but many are adapted to pollination by sunbirds. These species typically have red to orange, trumpet-like flowers that secrete large amounts of nectar. Other species are adapted to pollination by butterflies and moths, carrion flies and long-proboscid flies, and even monkey-beetles.
List of general
Up to 80 genera have been recognised in the family, with a total of around 1500 species, worldwide. The Afrotropical realm, and in particular South Africa, have the greatest diversity of genera. The spice saffron comes from the stigma of the saffron crocus, Crocus sativus.
They are represented in Australia by 5 genera and 26 species in the primitive tribes Isophysideae and Sisyrinchieae (ref: Flora of Australea, Vol 46). Includes Galaxia, Gynandriris, Homera, Hexaglottis, Homoglossum, synnotia and highlighted genera below.
Afrocrocus
Ainea
Alophia
Anapalina
Aristea
Babiana, Baboon flower
Bobartia
Calydorea, Violet-lily
Chasmanthe, African cornflag
Cipura
Cobana
Crocosmia, Montbretia
Crocus
Cyanixia
Cypella
Devia
Dierama, Fairy-wand
Dietes, Fortnight lily, African iris
Diplarrena
Duthiastrum
Eleutherine
Ennealophus
Ferraria
Freesia (syn. Anomatheca)
Geissorhiza
Gelasine
Geosiris
Gladiolus
Herbertia
Hesperantha
Hesperoxiphion
Iris
Isophysis
Ixia, African cornlily
Klattia
Lapeirousia
Larentia
Lethia
Libertia
Mastigostyla
Melasphaerula
Micranthus
Moraea
Nemastylis
Neomarica
Nivenia
Olsynium, Grasswidow
Orthrosanthus
Patersonia
Pillansia
Pseudotrimezia
Radinosiphon
Romulea
Savannosiphon
Sisyrinchium, Blue-eyed grass, Yellow-eyed grass
Solenomelus
Sparaxis, Wandflower, Harlequin flower
Syringodea
Thereianthus
Tigridia Tiger flower, Mexican shell flower
Trimezia
Tritonia
Tritoniopsis
Watsonia, Bugle-lily
Witsenia
Xenoscapa
Zygotritonia
Source:
Systematic Botany (2008), 33(3): pp. 495–508
© Copyright 2008 by the American Society of Plant Taxonomists
Iridaceae ‘Out of Australasia’? Phylogeny, Biogeography, and Divergence Time Based on Plastid DNA Sequences
Peter Goldblatt1,7, Aaron Rodriguez2, M. P. Powell3, T. Jonathan Davies4, John C. Manning5, M. van der Bank6, and Vincent Savolainen
Source:
Source:
The Southern heaths are a Gondwana group distributed in Australia, South Asia and Argentina.
The early botanists noted that this Gondwana group was similar to the hardy plants such as heather (family: Ericaceae) that dominate the heathlands and moorlands of Europe. The Australian plants were collectively called the Southern Heaths and assigned to the new family Epacridaceae. On the basis of findings in 2002 (in particular, from results of DNA analysis) the Australian heath plants were included in the same family as the northern heaths, the Ericaceae, but placed in a separate subfamily Styphelioideae this subfamily is further sub-divided into various tribes not shown in the diagram. In 2012 a change in the botanical rules of nomenclature made this classification invalid and the subfamily name was changed to Epacridoideae. For more details see ref at end. [Typification of some names in Epacridoideae (Ericaceae) by Darren M. Crayn, Kathleen A. Kron and Benjamin C. M. Potter (Telopea, Volume 17: 319-321, November 2014).)
Shown in this diagram are those genera that occur in NSW (Those found in Ku-ring-gai Wildflower Garden are indicated in bold letters.)
Distribution Members of the Australian Ericaceae occur:-
• All states except the Northern Territory,
• Mostly absent from the arid interior.
• Well represented in south west Western Australia
• Well represented in Tasmania,
• Present along the southern and eastern coasts of the continent extending to northern Queensland at elevation.
Habitat
They can be found in a range of habitats
• Coastal sandplains
• Sandstone ridges
• Common component of low, exposed vegetation known as heathland
• Nutrient deficient soils, often acidic.
They are well adapted to surviving in the prevailing harsh conditions of the high exposed sandstone ridges around the Sydney region.
General Features
The Australian heath plants are mostly low shrubs – just a few are small trees.
Leaves are arranged alternately and the venation in the leaves is narrowly palmate. (The venation is
sometimes described as “parallel” this being the appearance due to the thin leaves.)
Flowers are usually tubular or cup-shaped and occur in spikes or racemes (occasionally solitary).
The fruits are either drupes or capsules that split open when ripe.
Some Australian members of the family produce succulent fruits which were eaten by Aborigines and
the early settlers.
The abundant nectar on the flowers was also valued by Aborigines and birds
Cultivation
Heath plants are some of the showiest of Australian plants but they are relatively difficult to propagate and not widely available as nursery plants. The most commonly cultivated genus is Epacris. They tend
to have straggling growth and are best grown beside large rocks. They like moist, well-drained, acid soil and some shade.
EXAMPLES: