Helotidae

オオキスイムシ科

Type genus. Helota MacLeay, 1825

Helotidae Chapuis, 1876:

【形態 / Diagnosis】

【生態 / Bionomics】

• Adults and larvae of Helota gemmata Gorham feed on sap exuding from trees (Oliff, 1883; Fukuda, 1943; Lee & Sato, 2006). Sweeping flowers was found to be a good method for collecting members of the genus Neohelota (Lee & Sato, 2006).

【分布 / Distribution】

• Tropical and  subtropical  areas  of  the  Old  World,  but  excluding  New  Guinea  and  Australia (Wegrzynowicz, 2000). Palearctic and Oriental Regions (Helota Maclay, Neohelota Ohta, Metahelota Kirejtshuk); Afrotropical Region (Afrohelotina Kirejtshuk，Scrophohelota Kirejtshuk). 

【備考/ Notes】 

• 世界から5属約100種が知られる（Wegrzynowicz, 2000） ．

成虫写真 / Photos, adults

国外分布図 / Distribution Map, World

Genera:

• • +Burmahelota Liu, Ślipiński, Ren & Pang, 2019

  • +Palaeohelota Liu, Ślipiński, Ren & Pang, 2019

  • +Trihelota Tihelka, Huang et Cai, 2020

• Helota Macleay, 1825

• Neohelota Ohta, 1929

• Afrohelotina Kirejtshuk, 2000

• Metahelotella Kirejtshuk, 2000

• Scrophohelota Kirejtshuk, 2000

【参考文献 / References】

• • Achard, J. (1920) Description d’une nouvelle espèce d’Helota du Congo.— Rev. Zool. Afr. 8: 270-272.

  • Chapuis, F., 1876. Famille des Érotyliens, des Endomychides et des Coccinellides. In: T. Lacordaire and F. Chapuis  (eds.) Histoire Naturelle des Insectes. Genera des Coléoptères ou exposé méthodique et critique de tous les  genres proposés jusqu’ici dans cet ordre d’insectes. Tome Douzième: 1-424.— Paris.

  • Fukuda A.  1943.  Description of the larva of Helota gemmata Gorham.  Trans. Nat. Hist. Soc. Taiwan 33: 15-18. (In Japanese.)

  • Gorham,  H.S.,  1874.  Observations  on  the  genus  Helota,  M’Leay,  with  description  of  a  new  species from Japan.— Trans. ent. Soc. London 1874 (4): 447-449.

  • Heller,  K.M.,  1923.  Die  Coleopterenausbeute  der  Stötznerschen  Sze-Tschwan-Expedition  (1913-1915).— Ent. Bl. Biol. Syst. Käfer 19 (2): 61-79.

• • Hope, F.W., 1840. The Coleopterist’s Manual, Part the Third, Containing Various Families, Genera, and Species, of Beetles, Recorded by Linnaeus and Fabricius, also, Descriptions of Newly Discovered and Unpublished Insects: 1-191.— London.

  • Kirejtshuk, A. G. 2000. On origin and early evolution of the superfamily Cucujoidea (Coleoptera, Polyphaga). Comments on the family Helotidae. Kharkov Entomol. Soc. Gaz. 8: 8–38.

  • Kolbe, H.J., 1886. Beiträge zur Kenntniss der Coleopteren-Fauna Koreas, bearbeitet auf Grund der von Herrn  Dr.  C. Gottsche  während  der  Jahre  1883  und  1884  in  Korea  veranstalteten  Sammlung; nebst Bemerkungen über die zoogeographischen Verhältnisse dieses Faunengebietes und Untersuchungen über einen Sinnesapparat im Gaumen von Misolampidius morio.— Arch. Naturgesch. 52 (1): 139-240.

  • Kôno, H., 1939. Helotidae of Japan, Korea, and Formosa (Coleoptera).— Philipp. J. Sc. 69 (2): 157-160.

  • Lafer, G.Sh., 1992. 67. Sem. Helotidae: 274-276. In: Ler, P. A. (ed.) Opredelitel nasekomyh Dalnego Vostoka SSSR v shesti tomah. Tom III. Zhestkokrylye, ili zhuki. Chast 2: 1-704.— St. Petersburg.

  • Lawrence, J.F., 1991. Helotidae: p. 463. In: Stehr, F. W. (ed.), Immature Insects. Volume 2: i-xvi + 1-

  • 975.— Dubuque.

  • Lawrence, J.F. & Newton, A.F.Jr., 1995. Families and subfamilies of Coleoptera (with selected genera, notes,  references  and  data  on  family-group  names):  779-1006.  In:  J.  Pakaluk  and  S.A.  Slipinski (Eds.), Biology, Phylogeny and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Part 2: 1-1006.— Warszawa.

  • Lee, C.-F., 2007. Revision of Family Helotidae (Coleoptera: Cucujoidea): I. Gemmata Group of Genus Helota Annals of the Entomological Society of America, 100(5): 623-639.

  • Lee, C.-F., 2008. Revision of the Family Helotidae (Coleoptera: Cucujoidea) II: The Vigorsii Group of the Genus Helota. Annals of the Entomological Society of America, 101(4): 722-742.

  • Lee, C.-F., 2009. Revision of the Family Helotidae (Coleoptera: Cucujoidea) III: The Thibetana Group and a Checklist of Helota MacLeay Species. Annals of the Entomological Society of America, 102(1): 48-59.

  • Lee, C.-F., 2009. Revision of the Family Helotidae (Coleoptera: Cucujoidea): IV. The Genus Metahelotella. Annals of the Entomological Society of America, 102(5): 785-796.

  • Lee, C.-F., 2010. Revision of the Family Helotidae (Coleoptera: Cucujoidea): V. Species Group Classification of the Genus Neohelota Ohta and Revisions of N. laevigata and N. helleri Species Groups. Annals of the Entomological Society of America, 103(4): 500-510.

  • Lee, C.-F., 2011. Revision of Family Helotidae (Coleoptera: Cucujoidea): VI. Candezei Group of the Genus Neohelota. Annals of the Entomological Society of America, 104(4): 658-665.

  • Lee, C.-F., 2013. Revision of the Family Helotidae (Coleoptera: Cucujoidea): VII. The Attenuata Species Group of the Genus Neohelota. Annals of the Entomological Society of America, 106(2): 152-163.

  • Lee, C.-F. & Votruba, P., 2014. Revision of the Family Helotidae (Coleoptera: Cucujoidea): IX. The Culta Species Group and a Checklist of Neohelota Species. Annals of the Entomological Society of America, 107(2): 315-338.

  • Lee, C.-F. & Votruba, P., 2013. A revision of the family Helotidae (Coleoptera: Cucujoidea): VIII. The guerinii species-group of the genus Neohelota. Entomologica Basiliensia et Collectionis Frey, 34: 269-308.

  • Lee, C.-F. & Sato, M., 2005. The Helotidae of Taiwan (Coleoptera: Cucujoidea).  Zoological Studies 45(4): 529-552.

  • Lee，C.-F., Hisamatsu, S.-T. & Yang, P.-S., 2007. Morphology and Ontogeny of Immature Stages of Helotidae Based on Descriptions of Helota thoracica Ritsema and H. gemmata Gorham (Insecta: Coleoptera: Cucujoidea). Zoological Studies 46(6): 760-769.

  • Lewis, G., 1881. A new species of Helotidae from Japan.— Entomologist’s mon. Mag. 17: 255-256.

  • Mader,  L.,  1926.  Clavicornia:  687-752.  In:  A.  Winkler  (ed.)  Catalogus  Coleopterorum  regionis palaearcticae. Pars 6: 625-752.— Wien.

  • Mader, L., 1955. Neue Coleopteren aus Fukien (China). Helotidae, Languriidae, Erotylidae, Endomychidae, Coccinellidae. Koleopt. Rdsch. 33 (1-6): 62-78.

  • Nguyen-Phung,  T.,  1984.  Contribution  a  la  connaissance  du  genre  Helota Mac  Leay  (Coleoptera, Helotidae).— Nouv. Revue Ent. (Nouv. Sér.) 1 (2): 165-170.

  • Nguyen-Phung, T., 1985. Deux nouveaux Helota MacLeay du Laos (Coleoptera, Helotidae).— Rev. fr. Ent. (Nouv. Sér.) 7 (1): 15-18.

  • Oberthür, R., 1883. Trois espèces nouvelles du genre Helota.— Coleopt. Nov. 1: 59-61.

  • Ohta, Y., 1929a. Beitrag zur Kenntnis der Helotiden-Fauna von Japan, Korea und Formosa.— Insecta Matsum. 3 (2-3): 108-110.

  • Ohta, Y., 1929b. Einige Neue Helotiden- und Coccinelliden-Arten aus Formosa.— Insecta Matsum. 4(1-2): 66-70.

  • Ohta, Y., 1931. Ueber einige Coelophoren- und Helotiden-Arten aus Formosa, mit Beschreibung von 2 Neue Arten.— Insecta Matsum. 5 (3): 134-136.

  • Olliff, A.S., 1883. Descriptions of two larvae and new genera and species of Clavicorn Coleoptera, and a synopsis of the genus Helota, MacLeay.— Cistula ent. 3 (27): 49-61, pl.Reitter, 1876: 5.

  • Olliff, A.S., 1884a. Description of an African Species of the Coleopterous Genus Helota, MacLeay.—Ann. Mag. nat. Hist, Ser. 5 13 (78): 479-480.

  • Olliff,  A.S.,  1884b.  Additional  Notes  on  the  genus  Helota,  MacLeay,  and  a  Synonymic  List  of  the described species.— Cistula ent. 3: 99-101.

  • Ritsema, C., 1891a. Synopsis and alphabetical list of the described species of the Coleopterous genus Helota McL.— Notes Leyden Mus. 13 (3): 223-232.

  • Ritsema, C., 1911a. Fam. Helotidae: 104-106. In: W Junk and S. Schenkling (Eds.) Coleopt. Cat. 34: 1-106.— Berlin.

  • Wegrzynowicz, P. (2000) Catalogue of the Helotidae (Coleoptera: Cucujoidea). Zool. Med. Leiden 73 (27): 391-411, figs 1-2.

  • Weise, J., 1903. Helota Pauli n. sp.— Dt. ent. Z. 1903: 171.

  • Westwood, J.O., 1842. Insectorum novorum Centuria.— Ann. Mag. nat. Hist. 8: 120-130.

  • Westwood, J.O., 1848. The Cabinet of Oriental entomology: 1-88, 42 pls.— London.