Reference:

Takasuka K. (202x) Global checklist of spider parasitoids and pseudo-parasitoids of the ichneumonid wasp subfamily Pimplinae. Online at https://sites.google.com/site/keizaf/checklist-polysphinctine (accessed on 00 Month 202x).

2026年5月13日更新

下記は各属への小見出しリンク（Googleサイトはページ内リンクを小見出しでしか貼れない。。。本当は属一覧の属名に直リンクを貼りたい。）

クモヒメバチ属群 the Polysphincta group of genera 属一覧

Currently, 25 genera and 299 extant species plus 5 fossil species (Yu et al. 2016 +alpha)

Schizopyga subgroup sensu Matsumoto (2016)

  1. Genus Brachyzapus Gauld & Dubois, 2006 16 spp.

  2. Genus Dreisbachia Townes, 1962 16 spp.

  3. Genus Iania Matsumoto, 2016 1 species + 1 subspecies

  4. Genus Inbioia Gauld & Ugalde Gόmez, 2002 2 species

  5. Genus Piogaster Perkins, 1958 8 spp.

  6a. Genus Schizopyga (Schizopyga) Gravenhorst, 1829 8 spp. + 3 subspecies

  6b. Genus Schizopyga (Schizopygoides) Kasparyan, 1976 1 species

  7. Genus Zabrachypus Cushman, 1920 5 spp. + 1 fossil species

Acrodactyla subgroup sensu Matsumoto (2016)

  8. Genus Acrodactyla Haliday, 1838 29 spp. +1 subspecies

  9. Genus Eruga Townes, 1960 14 spp.

10. Genus Lamnatibia Palacio & Sääksjärvi, 2007 1 species

11. Genus Longitibia He & Yu, 1999 1 species

12.  Genus Megaetaira Gauld & Dubois, 2006 3 spp.

13. Genus Pterinopus Townes, 1969 1 species

Polysphincta subgroup sensu Matsumoto (2016)

14. Genus Acrotaphus Townes, 1960 27 spp.

15. Genus Aravenator Momoi, 1973 1 species

16. Genus Chablisea Gauld & Dubois, 2006 7 spp.

17. Genus Eriostethus Morley, 1914 18 spp.

18. Genus Flacopimpla Gauld, 1991 7 spp.

19. Genus Hymenoepimecis Viereck, 1912 29 spp.

20. Genus Oxyrrhexis Förster, 1869 4 spp. + 1 subspecies

21. Genus Polysphincta Gravenhorst, 1829 33 spp. + 4 fossil species + 2 subspecies

22. Genus Reclinervellus He & Ye, 1998 4 spp.

23. Genus Sinarachna Townes, 1960 4 spp. + 1 subspecies

24. Genus Ticapimpla Gauld, 1991 5 spp.

25. Genus Zatypota Förster, 1869 54 spp. + 8 subspecies

クモフシヒメバチ Pseudo-parasitoids of spider egg masses（直近祖先群・クモ卵塊寄生バチ）属一覧

1. Genus Clistopyga Gravenhorst, 1829 95 spp. +1 subspecies

2. Genus Tromatobia Förster, 1869 33 spp. + 3 subspecies

3. Genus Zaglyptus Förster, 1869 27 spp. +7 subspecies

2024年1月時点のGlobal checklist (Takasuka & Broad 2024)

Takasuka, K. & Broad, G.R. (2024) A bionomic overview of spider parasitoids and pseudo-parasitoids of the ichneumonid wasp subfamily Pimplinae. Contributions to Zoology 93 1-106.

クモヒメバチ属群 the Polysphincta group of genera 25属の種一覧

Schizopyga subgroup sensu Matsumoto (2016)

1. Genus Brachyzapus Gauld & Dubois, 2006 16 spp.

Gauld I.D. & Dubois J. (2006) Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae): a taxonomic revision of spider ectoparasitoids. Systematic Entomology 31 529-564.

Brachyzapus atripedalis (Sheng, 2001)

Brachyzapus carinatus Pham, Broad, Matsumoto & Wägele, 2012

Brachyzapus convergens Choi & Lee, 2019

Brachyzapus convexus Pham, Broad, Matsumoto & Wägele, 2012

Brachyzapus duboisi Pham, Broad, Matsumoto & Wägele, 2012

Brachyzapus fansipanensis Pham, Broad, Matsumoto & Wägele, 2012

Brachyzapus glaber Varga & Reshchikov, 2018

Brachyzapus greengoblin Varga & Di Giovanni, 2018

Brachyzapus hoanglienensis Pham, Broad, Matsumoto & Wägele, 2012

Brachyzapus nikkoensis (Uchida, 1928)

Brachyzapus nitidus (Hao & Sheng, 2002)

Brachyzapus nonareaeidos (Wang, 1997)

Brachyzapus politus Pham, Broad, Matsumoto & Wägele, 2012

Brachyzapus pyramidalis Choi & Lee, 2019 

Brachyzapus tenuiabdominalis (Uchida, 1941)

Brachyzapus unicarinatus (Uchida & Momoi, 1958)

Choi J.K., Kolarov J. & Lee J.W. (2019) Descriptions of two new species of Brachyzapus Gauld & Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from South Korea with key to a Palaearctic species. Zootaxa 4711 293-306.

Hao D. & Sheng M.L. (2002) A new specie of genus Zabrachypus Cushman (Hymenoptera: Ichneumonidae). Journal of Northeast Forestry University 30 95-96 (in Chinese with English summary).

Pham N.T., Broad G.R., Matsumoto R. & Wägele W.J. (2012) First record of the genus Brachyzapus Gauld and Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam, with descriptions of six new species. Journal of Natural History 46 1639-1661.

Sheng M.L. & Pan H.Y. (2001) Two new species of family Ichneumonidae (Hymenoptera). Entomologia Sinica 8 25-29.

Uchida T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido Imperial University 25 1-115.

Uchida T. (1941) Baiträge zur Systematik der Tribus Polysphinctini Japans. Insecta Matsumurana 15 112-122.

Uchida T. & Momoi S. (1958) On the species of Polysphincta Gravenhorst and Zatypota Förster from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 22-30.

Varga O., Reshchikov A. & Giovanni F.D. (2018) First record of the genus Brachyzapus Gauld & Dubois, 2006 (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region, with descriptions of two new species. Zootaxa 4377 581-586.

Wang S.F., Yao J. & Wang G.G. (1997) Hymenoptera: Ichneumonidae. In: X.K. Yang (eds.). Insects of the Three Gorge Reservoir area of Yangtze River. pp. 1617-1646 (in Chinese). Chongqing Publishing Company, Chongqing, China.

2. Genus Dreisbachia Townes, 1962 16 spp. 

Townes H. (1962) A generic name in the Polysphinctine Ichneumonids (Hymenoptera). Proceedings of the Entomological Society of Washington 64 38.

Dreisbachia amurensis Kasparyan, 1976

Dreisbachia anseli (Fernandez-Triana, 2007)

Dreisbachia aperta Momoi, 1966

Dreisbachia avivae Gauld, 1991

Dreisbachia congica (Benoit, 1953)

Dreisbachia curvicauda (Seyrig, 1935)

Dreisbachia delossantosi Ketelaera & Hodecek 2025

Dreisbachia dotae Gauld, Ugalde Gómez & Hanson, 1998

Dreisbachia eberhardi Gauld, Ugalde Gómez & Hanson, 1998

Dreisbachia flavifrontalis (Uchida & Momoi, 1959)

Dreisbachia lutea Gauld, 1984

Dreisbachia mira (Tosquinet, 1903)

Dreisbachia moreirai (Loffredo & Penteado-Dias 2008)

Dreisbachia navaja (Townes, 1960)

Dreisbachia punctata (Uchida & Momoi, 1959)

Dreisbachia slossonae (Davis, 1898)

Benoit P.L.G. (1953) Sur des Polysphinctini du Congo belge (Ichneum. - Pimpl.). Revue de Zoologie et de Botanique Africaines 47 137-141.

Davis G.C. (1897) Descriptions of new species of Trigonalidæ, Stephanidæ and Ichneumonidæ. Transactions of the American Entomological Society 24 349-372 

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biología Tropical 46 (Suppl. 1) 1-189.

Kasparyan D.R. (1976) [Review of Ichneumonids of the tribe Polysphinctini and Poemeniini (Hymenoptera, Ichneumonidae) of the Far East.] Trudy Zoologicheskogo Instituta 67 68-89 (in Russian).

Ketelaere A.D. & Hodecek J. (2025) A new species of Dreisbachia (Hymenoptera: Ichneumonidae) from the Dominican Republic. Bulletin de la Société royale belge d'Entomologie 161 70-75.

Momoi S. (1966) Some new Ichneumonidae (Hymenoptera) from New Guinea and adjacent areas. Pacific Insects 8 152-164.

Seyrig A. (1935) Mission scientifique de l'Omo. Tome III. Fascicule 18. Hymenoptera, II. Ichneumonidae: Cryptinae, Pimplinae, Tryphoninae et Ophioninae. Mémoires du Muséum National d'Histoire Naturelle 4 1-100.

Tosquinet D. (1903) Ichneumonides nouveaux. Mémoires de la Société entomologique de Belgique 10 1-403.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. & Momoi S. (1959) On the species of the genera Laufeia and Lycorina occurring in Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 82-87.

 3. Genus Iania Matsumoto, 2016 1 species + 1 subspecies (Matsumoto, 2016)

Matsumoto R. (2016) Molecular phylogeny and systematics of the Polysphincta group of genera (Hymenoptera, Ichneumonidae, Pimplinae). Systematic Entomology 41 854-864.

Iania pictifrons (Thomson, 1877)

Iania pictifrons rufigaster (Constantineanu, Ciochia, Constantineanu & Ularu, 1967)

Constantineanu M.I., Ciochia V., Constantineanu R.M., Mustata G. & Ularu G. (1967) Contributions a l'etude des Ichneumonides de la montagne de Gutii, région de Maramures. Analele ştiinţifice ale Universităţii “Al. I. Cuza” din Iaşi. Sect. II a 13 47-61.

Thomson C.G. (1877) XXVII. Bidrag till kännedom om Sveriges Pimpler. Opuscula Entomologica, Lund 8 732-777.

 4. Genus Inbioia Gauld & Ugalde Gόmez, 2002 2 spp.

Inbioia himalayensis De Keteraere & Maqbool, 2025

Inbioia pivai Gauld & Ugalde Gómez, 2002

Bhat D.M., Ketelaere A.D. & Maqbool A. (2025) First record of polysphinctine genus Inbioia Gauld & Ugalde Gomez, 2002 (Hymenoptera: Ichneumonidae) from the Palearctic with description of a new species. Zootaxa 5618 424-432. 

Gauld I.D. & Ugalde Gόmez J.A. (2002) Appendix 1. Subfamily Pimplinae. Further supplement to volume I. Memoirs of the American Entomological Institute 66 744-746.

 5. Genus Piogaster Perkins, 1958 8 spp. 

Perkins J.F. (1958) A new genus and three new species of Polysphinctini from Europe (Hym., Ichneumonidae). Entomologist 91 263-267.

Piogaster albina Perkins, 1958

Piogaster daisetsuzana Kusigemati, 1985

Piogaster lucida Constantineanu & Constantineanu, 1969

Piogaster maculata Townes, 1960

Piogaster pilosator (Aubert, 1958)

Piogaster punctulata Perkins, 1958

Piogaster ussuriensis Kasparyan & Khalaim, 2007

Piogaster variegata Bass, Bennett & Schwarzfeld, 2026

Aubert J.F. (1958) Ichneumonide Pimpline d'un genre nouveau, espèce nouvelle, rèpandue sur le rivage mèditerranéen. Bulletin de la Société Entomologique de Mulhouse 1958 79-80.

Bass A., Bennett A.M.R., Spasojevic T. & Schwarzfeld M. (2026) Revision of the world species of Piogaster Perkins (Hymenoptera, Ichneumonidae, Pimplinae), ectoparasitoids of jumping spiders (Araneae, Salticidae) with a description of one new species. Journal of Hymenoptera Research 99 621-701.

Constantineanu M.I. & Constantineanu R.M. (1969) Ichneumonides nouveaux pour la science et pour la faune de la Roumanie. Comunicari de Zoologie, Bucaresti 8 155-165 (in Romanian with French & Russian summary).

Kasparyan D.R. (1981) [New species of the subfamily Pimplinae (Hymenoptera, Ichneumonidae), parasites of spiders in the Far East of the USSR.]. Trudy Zoologicheskogo Instituta 92 108-112 (in Russian).

Kasparyan D.R. & Khalaim A.I. (2007) [1. Subfamily Pimplinae]. In: A.S. Lelej (eds.). [Key to the insects of the Russian Far East. Vol. IV. Neuropteroidea, Mecoptera, Hymenoptera. Pt. 5]. pp. 279-333 (in Russian). Vladivostok, Russia.

Kusigemati K. (1985) A new species of the genus Piogaster Perkins from Japan (Hymenoptera, Icuneumonidae). Kontyû 53 583-586.

Perkins J.F. (1958) A new genus and three new species of Polysphinctini from Europe (Hym., Ichneumonidae). Entomologist 91 263-267.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

 6a. Genus Schizopyga (Schizopyga) Gravenhorst, 1829 8 spp. + 3 subspecies

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Schizopyga alnae Varga & Reshchikov, 2018

Schizopyga circulator (Panzer, 1800)

Schizopyga circulator meridionator Aubert, 1963

Schizopyga circulator pulchra Walley, 1936

Schizopyga coxator Constantineanu, 1973

Schizopyga flavifrons Holmgren, 1856

Schizopyga frigida Cresson, 1870

Schizopyga podagrica Gravenhorst, 1829

Schizopyga podagrica nigrifacies Constantineanu & Ciochia, 1967

Schizopyga quannanica Sun & Sheng, 2011

Schizopyga varipes Holmgren, 1856

Aubert J.F. (1963) Les Ichneumonides du rivage Méditerranéen Français (5e série, Département du Var). Vie et Milieu 14 847-878.

Constantineanu M.I. (1973) Ichneumonidae (Hymenoptera) din Delta Dunarii (Grindul Caraorman). Buletin Muzeul Delta Dunarii, Tulcea. Peuce 3 131-163.

Constantineanu M.I. & Ciochia V. (1967) Ichneumonidae in der Burzenländer Senke und ihrer Umgebung. Muzeul Regional Brasov. Culegere de Studii si Cercetari 1 33-52.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Fernández-Triana J.L. (2007) Notes on Cuban Pimplinae (Hymenoptera: Ichneumonidae), with the description of two new species. Boletin de la Sea 40 211-214.

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Holmgren A.E. (1856) Om slägtet Schizopyga. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar 13 69-72.

Loffredo A.P.S. & Penteado-Dias A.M. (2008) First record of Schizopyga Gravenhorst (Hymenoptera, Ichneumonidae, Pimplinae) from Brazil and a description of a new species. Brazilian Journal of Biology 68 457-458.

Panzer G.W.F. (1800) Faunae Insectorum Germanicae. Heft 79 8-14.

Sun S.P. & Sheng M.L. (2011) A new species of the genus Schizopyga Gravenhorst (Hymenoptera, Ichneumonidae) with a key to species known from China. Acta Zootaxonomica Sinica 36 970-972.

Varga O. & Reshchikov A. (2018) Schizopyga alinae, a genus of pimpline parasitoid wasps (Hymenoptera: Ichneumonidae: Pimplinae) new to the Afrotropical region. Zootaxa 4422 291-295.

Walley G.S. (1936) A new Schizopyga from Canada (Hymenop.; Ichneumonidae). The Canadian Entomologist 68 20-21.

 6b. Genus Schizopyga (Schizopygoides) Kasparyan, 1976 1 species

Schizopyga (Schizopygoides) nitida Kasparyan, 1976

Kasparyan D.R. (1976) [Review of Ichneumonids of the tribe Polysphinctini and Poemeniini (Hymenoptera, Ichneumonidae) of the Far East.] Trudy Zoologicheskogo Instituta 67 68-89 (in Russian).

 7. Genus Zabrachypus Cushman, 1920 5 spp. + 1 fossil species

Cushman R.A. (1920) The North American ichneumon-flies of the tribes Lycorini, Polysphinctini, and Theroniini. Proceedings of the United States National Museum 58 7-48.

Zabrachypus cazorlensis Rey del Castillo, 1996

Zabrachypus moldavicus Constantineanu & Constantineanu, 1969

Zabrachypus primus Cushman, 1920

Zabrachypus rubroabdominalis Mohammadi-Khoramabadi, Talebi & Broad, 2022

Zabrachypus rugatus Mohammadi-Khoramabadi, Talebi & Broad, 2022

(Zabrachypus tumidus Khalaim, 2008 [fossil])

Constantineanu M.I. & Constantineanu R.M. (1969) Ichneumonides nouveaux pour la science et pour la faune de la Roumanie. Comunicari de Zoologie, Bucaresti 8 155-165 (in Romanian with French & Russian summary).

Cushman R.A. (1920) The North American ichneumon-flies of the tribes Lycorini, Polysphinctini, and Theroniini. Proceedings of the United States National Museum 58 7-48.

Khalaim A.I. (2008) Fossil ichneumon wasps (Hymenoptera: Ichneumonidae) from Biamo (Russia), Oligocene. Alavesia 2 101-112.

Mohammadi-Khoramabadi A., Talebi A., Broad G.R. & Zwakhals K. (2022) A study of the genus Zabrachypus Cushman, 1920 (Hymenoptera Ichneumonidae Pimplinae) from Iran with descriptions of two new species. Redia 105 29-36.

Rey del Castillo C. (1996) Addition to the knowledge of Spanish Polysphinctini with description of Zabrachypus cazorlensis sp. n. (Hymenoptera: Ichneumonidae). Boletín de la Asociación Española de Entomología 20 141-147 (in Spanish).

Acrodactyla subgroup sensu Matsumoto (2016)

 8. Genus Acrodactyla Haliday, 1838 29 spp. +1 subspecies

Haliday A.H. (1838) Descriptions of new British insects, indicated in Mr. Curtis's guide. Annals of Natural History 2 112-121.

Acrodactyla aequaria Momoi, 1966

Acrodactyla atricoxa Kusigemati, 1984

Acrodactyla carinator (Aubert, 1965)

Acrodactyla carinator meridionator (Aubert, 1965)

Acrodactyla cursor Gauld, 1984

Acrodactyla degener (Haliday, 1838)

Acrodactyla elongata Pham, Broad, Matsumoto & Böhme, 2012

Acrodactyla gressitti Momoi, 1966

Acrodactyla iliensis Sheng & Bian, 1996

Acrodactyla indica Momoi, 1966

Acrodactyla jubata Townes, 1960

Acrodactyla lydia Kasparyan, 1979

Acrodactyla maiphuquyi Pham, Broad, Matsumoto & Böhme, 2012

Acrodactyla micans Gauld, 1984

Acrodactyla mitis Kasparyan, 1976

Acrodactyla mixta Kasparyan, 1976

Acrodactyla ocellata Townes, 1960

Acrodactyla orientalis (Cushman, 1933)

Acrodactyla phuthoensis Pham, Broad, Matsumoto & Böhme, 2012

Acrodactyla quadrisculpta (Gravenhorst, 1820)

Acrodactyla rufithorax (Györfi, 1944)

Acrodactyla rufotibiator Aubert, 1964

Acrodactyla shawi Pham, Broad, Matsumoto & Böhme, 2012

Acrodactyla similis Horstmann, 2011

Acrodactyla syndromosa Kusigemati, 1984

Acrodactyla taiwanensis (Cushman, 1933)

Acrodactyla takewakii (Uchida, 1927)

Acrodactyla tami Pham, Broad, Matsumoto & Böhme, 2012

Acrodactyla trochanterata Kasparyan, 1976

Acrodactyla zekhem Gauld, 1984

Aubert J.F. (1964) Ichneumonides de France et du Bassin méditerranéen appartenant à un genre nouveau et neuf espèces nouvelles. Bulletin de la Société Entomologique de Mulhouse 1964 35-40.

Aubert J.F. (1965) Ichneumonides d'Europe appartenant à dix espèces nouvelles et plusieurs genres nouveaux. Bulletin de la Société Entomologique de Mulhouse 1965 15-23.

Cushman R.A. (1933) H. Sauter's Formosa-collection: Subfamily Ichneumoninae (Pimplinae of Ashmaed). Insecta Matsumurana 8 1-50.

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

Gravenhorst J.L.C. (1820) Monographia Ichneumonum Pedemontanae Regionis. Memorie della Reale Academia dell Scienze di Torino 24 275-388.

Györfi J. (1944) Vier neue Ichneumoniden-Arten aus Ungarn. Fragmenta Faunistica Hungarica 7 42-45.

Haliday A.H. (1838) Descriptions of new British insects, indicated in Mr. Curtis's guide. Annals of Natural History 2 112-121.

Horstmann K. (2011) Revisionen von Schlupfwespen-Arten XV (Hymenoptera: Ichneumonidae). Mitteilungen der Münchner Entomologischen Gesellschaft 101 5-13.

Kasparyan D.R. (1976) [Review of Ichneumonids of the tribe Polysphinctini and Poemeniini (Hymenoptera, Ichneumonidae) of the Far East.] Trudy Zoologicheskogo Instituta 67 68-89 (in Russian).

Kasparyan D.R. (1979) [A new species of Acrodactyla Haliday (Hymenoptera, Ichneumonidae).] Trudy Vsesoyuznogo Entomologicheskogo Obshchestva 61 132-134 (in Russian).

Kusigemati K. (1984) Some Ephialtinae of South East Asia, with descriptions of eleven new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 5 126-150.

Momoi S. (1966) Some new Ichneumonidae (Hymenoptera) from New Guinea and adjacent areas. Pacific Insects 8 152-164.

Pham N.T., Broad G.R., Matsumoto R. & Böhme W. (2012) First record of the genus Acrodactyla Haliday (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam, with descriptions of six new species. Zootaxa 3207 40-53.

Sheng M.L. & Bian X. (1996) A new species of genus Acrodactyla (Hymenoptera: Ichneumonidae) from China. Entomotaxonomia 18 216-218 (in Chinese).

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. (1927) Zwei neue Schmarotzerhymenopteren der Spinnen. Insecta Matsumurana 1 171-174.

Uchida T. & Momoi S. (1958) On the species of Polysphincta Gravenhorst and Zatypota Förster from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 22-30.

 9. Genus Eruga Townes, 1960 14 spp.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Eruga albanoi Gauld, 1991

Eruga atrata Townes, 1960

Eruga draperi Gauld, 1991

Eruga erratica (Seyrig, 1932)

Eruga gutfreundi Gauld, 1991

Eruga haseltinei Gauld, 1991

Eruga kauli Gauld, 1991

Eruga lineata Townes, 1960

Eruga paulisiana (Benoit, 1953)

Eruga rufa Townes, 1960

Eruga straussi Gauld, 1991

Eruga telljohanni Gauld, 1991

Eruga unilabiana Pádua & Sobczak, 2018

Eruga viatrix (Seyrig, 1932)

Benoit P.L.G. (1953) Notes Ichneumonologiques Africaines IV. Revue de Zoologie et de Botanique Africaines 47 153-159.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Seyrig A. (1932) Les Ichneumonides de Madagascar. I Ichneumonidae Pimplinae. Mémoires de l'Académie Malgache. Fascicule 11 1-183.

Sobczak J.F., Pádua D.G., Costa L.F.A., Carvalho J.L.V.R., Ferreira J.P.S., Sobczak J.C.M.S.M. & Messas Y.F. (2018) The parasitoid wasp Eruga unilabiana Pádua & Sobczak, sp. nov. (Hymenoptera: Ichneumonidae) induces behavioral modification in its spider host. Entomological Science 21 59-65.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

10. Genus Lamnatibia Palacio & Sääksjärvi, 2007 1 species

Lamnatibia andina Palacio & Sääksjärvi, 2007

Palacio E., Sääksjärvi I.E. & Vahtera V. (2007) Lamnatibia, a new genus of the Polysphincta group of genera from Colombia (Hymenoptera: Ichneumonidae; Pimplinae). Zootaxa 1431 55-63.

11. Genus Longitibia He & Yu, 1999 1 species

Longitibia sinica He & Ye, 1999

He J.H. & Ye S.F. (1999) A new genus of Polysphinctini (Hymenoptera: Ichneumonidae) from China. Entomologia Sinica 6 8-10.

12. Genus Megaetaira Gauld & Dubois, 2006 3 spp.

Gauld I.D. & Dubois J. (2006) Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae): a taxonomic revision of spider ectoparasitoids. Systematic Entomology 31 529-564.

Megaetaira inoperta (Kusigemati, 1984)

Megaetaira madida (Haliday, 1838)

Megaetaira varicarinata (Uchida & Momoi, 1958)

Haliday A.H. (1838) Descriptions of new British insects, indicated in Mr. Curtis's guide. Annals of Natural History 2 112-121.

Kusigemati K. (1984) Some Ephialtinae of South East Asia, with descriptions of eleven new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 5 126-150.

Uchida T. & Momoi S. (1958) On the species of Polysphincta Gravenhorst and Zatypota Förster from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 22-30.

13. Genus Pterinopus Townes, 1969 1 species

Pterinopus scambus Townes, 1969

Townes H. (1969) The genera of Ichneumonidae, part 1. Memoirs of the American Entomological Institute 11 1-300.

Polysphincta subgroup sensu Matsumoto (2016)

14. Genus Acrotaphus Townes, 1960 27 spp.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Acrotaphus amajari Pádua, 2020

Acrotaphus amazonicus Pádua & Sääksjärvi, 2020

Acrotaphus bodoquenaensis Pádua, 2020

Acrotaphus chedelae Gauld, 1991

Acrotaphus cuzconus Pádua & Sääksjärvi, 2020

Acrotaphus dolichopus Pádua, 2020

Acrotaphus fasciatus (Brullé, 1846)

Acrotaphus fascipennis (Cresson, 1865)

Acrotaphus ferruginosus (Cresson, 1865)

Acrotaphus franklini Gauld, 1991

Acrotaphus fuscipennis (Cresson, 1865)

Acrotaphus homeofranklini Pádua, 2020

Acrotaphus jackiechani Pádua & Sääksjärvi, 2020

Acrotaphus japi Higa & Penteado-Dias, 2020

Acrotaphus kourou Pádua & Sääksjärvi, 2020

Acrotaphus latifasciatus (Cameron, 1911)

Acrotaphus mexicanus (Cameron, 1886)

Acrotaphus micrus Pádua, 2020

Acrotaphus monotaenius Pádua, 2020

Acrotaphus nambilloensis Pádua, 2020

Acrotaphus pseudoamazonicus Pádua & Sääksjärvi, 2020

Acrotaphus pseudomexicanus Pádua, 2020

Acrotaphus tibialis (Cameron, 1886)

Acrotaphus venezuelanus Pádua, 2020

Acrotaphus wagnerianae Pádua, 2020

Acrotaphus wiltii (Cresson, 1870)

Acrotaphus zampieronae Pádua, 2020

Brullé M.A. (1846) Tome Quatrième. Des Hyménoptères. Les Ichneumonides. In: A. Lepeletier de Saint Fargeau (eds.). Histoire Naturelle Des Insectes. Hyménoptères. pp. 56-521. Paris, France.

Cameron P. (1911) On the Hymenoptera of the Georgetown Museum, Britich Guiana. Part I. Timehri 1 152-186.

Cameron P. (1886) Hymenoptera. In: F.D. Godman & O. Salvin (eds.). Biologia Centrali-Americana, or, Contributions to the knowledge of the fauna and flora of Mexico and Central America. Zoology.). pp. 241-328. 

Cresson E.T. (1865) On the Hymenoptera of Cuba. Proceedings of the Entomological Society of Philadelphia 4 1-200.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Higa P.T. & Penteado-Dias A.M. (2020) Altitudinal effects on diversity of Pimplinae (Hymenoptera, Ichneumonidae) from Southeast Brazil and description of new species. Brazilian Journal of Biology 80 377-385.

Pádua D.G., Sääksjärvi I.E., Monteiro R.F. & Oliveira M.L. (2020) Review of the new world genus Acrotaphus Townes, 1960 (Hymenoptera: Ichneumonidae: Pimplinae), with descriptions of fifteen new species. Zootaxa 4719 1-62.

Pádua D.G. & Kloss T.G. (2020) A new species of Acrotaphus Townes (Hymenoptera: Ichneumonidae: Pimplinae) from the Brazilian Amazon, with notes on its host spider behavioral modification. Zootaxa 4810 161-168.

15. Genus Aravenator Momoi, 1973 1 species

Aravenator kamijoi Momoi, 1973

Momoi S. (1973) Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei. 331. Einige mongolischen Arten der Unterfamilien Ephialtinae und Xoridinae (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica 26 (Suppl.) 219-239.

16. Genus Chablisea Gauld & Dubois, 2006 7 spp.

Gauld I.D. & Dubois J. (2006) Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae): a taxonomic revision of spider ectoparasitoids. Systematic Entomology 31 529-564.

Chablisea albifacialis (Kusigemati, 1984)

Chablisea concava Liu, He & Chen, 2009

Chablisea condimenta Pham, Broad, Matsumoto & Wägele, 2011

Chablisea imbiba Gauld & Dubois,2006

Chablisea khuatdanglongi Pham, Broad, Matsumoto & Wägele, 2011

Chablisea shaanxiensis (Liu, He & Chen, 2009)

Chablisea varicolor Liu, He & Chen, 2009

Gauld I.D. & Dubois J. (2006) Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae): a taxonomic revision of spider ectoparasitoids. Systematic Entomology 31 529-564.

Kusigemati K. (1984) Some Ephialtinae of South East Asia, with descriptions of eleven new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 5 126-150.

Liu J.X., He J.H. & Chen X.X. (2009) Two new species of genus Chablisea Gauld et Dubois, 2006 (Hymenoptera: Ichneumonidae: Pimplinae) from China. Biologia 64 1165-1169.

Pham N.T., Broad G., Matsumoto R. & Wägele W. (2011) Two new species of the genus Chablisea Gauld et Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam. Biologia 66 1134-1139.

17. Genus Eriostethus Morley, 1914 18 spp.

Morley C. (1914) A Revision of the Ichneumonidae Based on the Collection in the British Museum (Natural History) Part III. Tribes Pimplides and Bassides. British Museum, London, U.K. 148 pp.

Eriostethus babai (Kusigemati, 1987)

Eriostethus bifasciatus (Baltazar, 1964)

Eriostethus carinatus Baltazar, 1964

Eriostethus chinensis (He, 1985)

Eriostethus fasciatus (Momoi, 1966)

Eriostethus infumatus (Momoi, 1966)

Eriostethus krombeini (Baltazar, 1964)

Eriostethus maximus Gauld, 1984

Eriostethus mesorufus Momoi, 1966

Eriostethus minimus Gauld, 1984

Eriostethus nigropictus Baltazar, 1964

Eriostethus nodatus (Baltazar, 1964) 

Eriostethus perkinsi (Baltazar, 1964)

Eriostethus pulcherrimus Morley, 1914

Eriostethus rufus (Uchida, 1932)

Eriostethus samoanus (Fullaway, 1938)

Eriostethus sinuatus (Momoi, 1966)

Eriostethus trifasciatus (Baltazar, 1964)

Baltazar C.R. (1964) Eriostethus Morley and a new polysphinctine genus (Ichneumonidae, Hymenoptera). Pacific Insects 6 388-401.

Fullaway D.T. (1938) New species from the Bishop Museum collection of Samoan parasitic Hymenoptera. Proceedings of the Hawaiian Entomological Society 10 49-52.

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

He J.H. (1985) [A new species of Millironia Baltazar (Hymenoptera: Ichneumonidae).] Wuyi Science Journal 5 63-65 (in Chinese with English summary).

Kusigemati K. (1987) Some Ephialtinae, Banchinae, Porizontinae, Mesochorinae, Metopiinae and Acaenitinae of Formosa (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 8 21-33.

Momoi S. (1966) Some new Ichneumonidae (Hymenoptera) from New Guinea and adjacent areas. Pacific Insects 8 152-164.

Morley C. (1914) A Revision of the Ichneumonidae Based on the Collection in the British Museum (Natural History) Part III. Tribes Pimplides and Bassides. British Museum, London, U.K. 148 pp.

18. Genus Flacopimpla Gauld, 1991 7 spp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Flacopimpla barathrica Fritzén, 2014

Flacopimpla kasparyani Khalaim & Ruíz-Cancino, 2011

Flacopimpla gerardoi (Gauld, Ugalde Gómez & Hanson, 1998)

Flacopimpla nigriceps (Walsh, 1873)

Flacopimpla parva (Cresson, 1870)

Flacopimpla sulina Graf & Kumagai, 1998

Flacopimpla varelae Gauld, 1991

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Fritzén N.R. (2014) Two new species of the Polysphincta genus-group (Hymenoptera: Ichneumonidae: Pimplinae) reared from their spider (Araneae) hosts in Europe. Zootaxa 3894 117-130.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biologia Tropical 46 (Suppl. 1) 1-189.

Graf V. & Kumagai A.F. (1997) The occurrence of Flacopimpla Gauld in Brazil (Hymenoptera, Ichneumonidae, Pimplinae). Revista Brasileira de Zoologia 14 737-777.

Khalaim A.I. & Ruíz-Cancino E. (2011) North and Central American species of the genus Flacopimpla Gauld (Hymenoptera: Ichneumonidae: Pimplinae), with description of a new species from Mexico. Proceedings of the Zoological Institute RAS 315 70-74.

Walsh B.D. (1873) Descriptions of North American Hymenoptera. Transactions of the Academy of Sciences of St. Louis 3 65-166.

19. Genus Hymenoepimecis Viereck, 1912 29 spp.

Viereck H.L. (1912) Descriptions of five new genera and twenty six new species of Ichneumon-flies. Proceedings of the United States National Museum 42 139-153.

Hymenoepimecis amazonensis Pádua & Oliveira, 2015

Hymenoepimecis andina Pádua & Sääksjärvi, 2020

Hymenoepimecis argyraphaga Gauld, 2000

Hymenoepimecis atriceps (Cresson, 1865)

Hymenoepimecis atlantica Pádua, 2022

Hymenoepimecis bicolor (Brullé, 1846)

Hymenoepimecis cameroni Townes, 1966

Hymenoepimecis castilloi Pádua & Sääksjärvi, 2020

Hymenoepimecis dolichocarinata Pádua & Sääksjärvi, 2020

Hymenoepimecis duckensis Pádua & Onody, 2015

Hymenoepimecis ecuatoriana Pádua & Sääksjärvi, 2020

Hymenoepimecis heidyae Gauld, 1991

Hymenoepimecis heteropus (Kriechbaumer, 1890)

Hymenoepimecis japi Sobczak, Loffredo, Penteado-Dias & Gonzaga, 2009

Hymenoepimecis jordanensis Loffredo & Penteado-Dias 2009

Hymenoepimecis kleini Pádua & Sobczak, 2015

Hymenoepimecis longilobus Pádua & Sääksjärvi, 2020

Hymenoepimecis manauara Pádua & Oliveira, 2015

Hymenoepimecis neotropica (Brues & Richardson, 1913)

Hymenoepimecis pinheirensis Penteado-Dias & Pádua 2022

Hymenoepimecis pucallpina Pádua & Sääksjärvi, 2020

Hymenoepimecis rafaelmartinezi Pádua & Sääksjärvi, 2020

Hymenoepimecis ribeiroi Pádua & Sobczak, 2015

Hymenoepimecis robertsae Gauld, 1991

Hymenoepimecis silvanae Loffredo & Penteado-Dias 2009

Hymenoepimecis sooretama Sobczak, Loffredo, Penteado-Dias & Gonzaga, 2009

Hymenoepimecis tedfordi Gauld, 1991

Hymenoepimecis uberensis Pádua & Onody, 2015

Hymenoepimecis veranii Loffredo & Penteado-Dias 2009

Brues C.T. & Richardson C.H. (1913) Descriptions of new parasitic Hymenoptera from British Guiana. Bulletin of the American Museum of Natural History 32 485-503.

Brullé M.A. (1846) Tome Quatrième. Des Hyménoptères. Les Ichneumonides. In: A. Lepeletier de Saint Fargeau (eds.). Histoire Naturelle Des Insectes. Hyménoptères. pp. 56-521. Paris, France.

Cresson E.T. (1865) On the Hymenoptera of Cuba. Proceedings of the Entomological Society of Philadelphia 4 1-200.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D. (2000) The re-definition of pimpline genus Hymenoepimecis (Hymenoptera: Ichneumonidae) with a description of a plesiomorphic new Costa Rican species. Journal of Hymenoptera Research 9 213-219.

Kloss T.G., Pádua D.G., Almeida S.D.S., Penteado-Dias A.M., Mendes-Pereira T., Sobczak J.F., Lacerda F.G. & Gonzaga M.O. (2022) A new Darwin wasp (Hymenoptera: Ichneumonidae) and new records of behavioral manipulation of the host spider Leucauge volupis (Araneae: Tetragnathidae). Neotropical Entomology 51 821-829.

Kriechbaumer J. (1890) Ichneumoniden-Studien. Neue Ichneumoniden des Wiener Museums. II. Annalen des Naturhistorischen Hofmuseums Wien 5 479-491.

Loffredo A.P.S. & Penteado-Dias A.M. (2009) New species of Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Brazilian Atlantic forest. Revista Brasileira de Entomologia 53 11-14.

Pádua D.G. (2022) First record of the Darwin wasp Hymenoepimecis Viereck, 1912 (Hymenoptera: Ichneumonidae: Pimplinae) from Argentina and Bolivia, with description of a new species. Zootaxa 5169 49-60.

Pádua D.G., Oliveira M.L., Onody H.C., Sobczak J.F., Sääksjärvi I.E. & Gómez I.C. (2015) The Brazilian Amazonian species of Hymenoepimecis Viereck, 1912 (Hymenoptera: Ichneumonidae: Pimplinae). Zootaxa 4058 175-194.

Pádua D.G., Sääksjärvi I.E., Monteiro R.F. & Oliveira M.L. (2020) Seven new species of spider-attacking Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Ecuador, French Guiana, and Peru, with an identification key to the world species. ZooKeys 935 57-92.

Sobczak J.F., Loffredo A.P.S., Penteado-Dias A.M. & Gonzaga M.O. (2009) Two new species of Hymenoepimecis (Hymenoptera: Ichneumonidae: Pimplinae) with notes on their spider hosts and behaviour manipulation. Journal of Natural History 43 2691-2699.

Townes H. & Townes M. (1966) A catalogue and reclassification of the Neotropic Ichneumonidae. Memoirs of the American Entomological Institute 8 1-367.

20. Genus Oxyrrhexis Förster, 1869 4 spp. + 1 subspecies

Förster A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 25 135-221.

Oxyrrhexis carbonator (Gravenhorst, 1807)

Oxyrrhexis carbonator texana (Cresson, 1870)

Oxyrrhexis eurus Kasparyan, 1977

Oxyrrhexis rugosa Liu, He & Chen, 2009

Oxyrrhexis zephyrus Fritzén, 2014

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Fritzén N.R. & Fjellberg A. (2014) Natural history of Oxyrrhexis zephyrus sp.n. (Hymenoptera, Ichneumonidae), a parasitoid of Enoplognatha serratosignata (Araneae, Theridiidae), with notes on taxonomy and other host species of Oxyrrhexis. Arthropoda Selecta 23 135-144.

Gravenhorst J.L.C. (1807) Vergleichende Übersicht des Linneischen und einiger neurern zoologischen Systeme, nebst dem eingeschalteten Verzeichnisse der zoologischen Sammlung des Verfasser und den Beschreibungen neuer Thierarten, die in derselben vorhanden sind. Göttingen. 476 pp.

Kasparyan D.R. (1977) [Ichneumonids of the subfamilies Pimplinae and Tryphoninae (Hymenoptera, Ichneumonidae) new for Mogolia and Transbaikalia.] Nasekomye Mongolii 5 456-469 (in Russian).

Liu J.X., He J.H. & Chen X.X. (2009) Revision of genus Oxyrrhexis Foerster, 1869 (Hymenoptera: Ichneumonidae: Pimplinae) from China. Annales Zoologici 59 171-177.

21. Genus Polysphincta Gravenhorst, 1829 33 spp. + 4 fossil species + 2 subspecies

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Polysphincta albipes Cresson, 1880

Polysphincta areolaris Ratzeburg, 1844

(Polysphincta atavina Cockerell, 1921 [fossil])

Polysphincta bonita Pádua & Sääksjärvi, 2021

Polysphincta boops Tschek, 1869

Polysphincta burgessii Cresson, 1870

Polysphincta cosnipata Pádua & Sääksjärvi, 2021

Polysphincta dizardi Gauld, 1991

Polysphincta gutfreundi Gauld, 1991

Polysphincta inca Pádua, Sääksjärvi & Spasojevic, 2021

(Polysphincta? inundata Brues, 1910 [fossil], see Spasojevic et al. (2022))

Polysphincta janzeni Gauld, 1991

Polysphincta jundiai Higa & Penteado-Dias, 2020

Polysphincta koebelei Howard, 1892

Polysphincta latistriata Ratzeburg, 1844

Polysphincta limata Cresson, 1870

Polysphincta longa Kasparyan, 1976

Polysphincta macroepomia Pádua & Sääksjärvi, 2021 

Polysphincta mascoi Gauld, 1991

Polysphincta naranjae Gauld, Ugalde Gómez & Hanson, 1998

Polysphincta organensis Pádua & Sääksjärvi, 2021

Polysphincta pichincha Pádua, Sääksjärvi & Spasojevic, 2021

Polysphincta punctigaster Varga & Reshchikov, 2015

Polysphincta purcelli Gauld, 1991

Polysphincta rufipes Gravenhorst, 1829

(Polysphincta saxea (Scudder, 1878) [fossil])

Polysphincta shabui Gauld, 1991

Polysphincta sinearanea Pádua, 2018

Polysphincta sirena Khalaim, 2024

Polysphincta soaresi Higa & Penteado-Dias, 2020

(Polysphincta statzi (Meunier, 1923) [fossil])

Polysphincta teresa Pádua & Sääksjärvi, 2021 

Polysphincta thoracica (Cresson, 1874)

Polysphincta townesorum De Santis & Millan, 1968

Polysphincta tuberosa Gravenhorst, 1829

Polysphincta tuberosa bruneti Provancher, 1873

Polysphincta tuberosa meridionator Aubert, 1963

Polysphincta vexator Fitton, Shaw & Gauld, 1988

Polysphincta xena Khalaim, 2024

Aubert J.F. (1963) Les Ichneumonides du rivage Méditerranéen Français (5e série, Département du Var). Vie et Milieu 14 847-878.

Brues C.T. (1910) The parasitic Hymenoptera of the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology at Harvard University 54 1-125.

Cockerell T.D.A. (1921) Fossil Arthropods in the British Museum. V. Oligocene Hymenoptera from the Isle of Wight. Annals and Magazine of Natural History 7 1-25.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Cresson E.T. (1874) Descriptions of Mexican Ichneumonidae. Proceedings of the Academy of Natural Sciences of Philadelphia 1873 374-413.

Cresson E.T. (1880) Report of the commissioner of agriculture. Report of the Entomologist of the United States Department of Agriculture 1879 208, 235, 238.

De Santis L. & Millan E. (1968) Adicones a la fauna Argentina de Icneumonidos (Hymenoptera). Notas de la Comision de Investigacion Cientificas de la Provincia de Buenos Aires 5 1-7.

Fitton M.G., Shaw M.R. & Gauld I.D. (1988) Pimpline ichneumon-flies Hymenoptera, Ichneumonidae (Pimplinae). Handbook for the Identification of British Insects, Royal Entomological Sciety of London, London, U.K. 110 pp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Higa P.T. & Penteado-Dias A.M. (2020) Altitudinal effects on diversity of Pimplinae (Hymenoptera, Ichneumonidae) from Southeast Brazil and description of new species. Brazilian Journal of Biology 80 377-385.

Howard L.O. (1892) The hymenopterous parasites of spiders. Proceedings of the Entomological Society of Washington 2 290-302.

Khalaim A.I., RuÍz-Cancino E. & Coronado-Blanco J.M. (2024) Darwin wasps of the subfamily Pimplinae (Hymenoptera: Ichneumonidae) of Mexico: Polysphincta genus-group. Zootaxa 5458 151-196. 

Kloss T.G., Pádua D.G., Lacerda F.G., Oliveira L.S., Cossolin J.F.S., Serrão J.E. & Gonzaga M.O. (2018) Suppression of orb-web building behavior of the spider Metazygia laticeps (O. Pickard-Cambridge, 1889) (Araneae: Araneidae) by a new parasitoid wasp. Zoologischer Anzeiger 276 100-106.

Meunier F. (1923) Sur quelques insectes de l'Aquitanien de Rott (Sept-Monts, Rhénanie). Miscellanea Entomologica, Revue Entomologique Internationale 26 82-88.

Pádua D.G., Sääksjärvi I.E., Spasojevic T., Kaunisto K.M., Monteiro R.F. & Oliveira M.L. (2021) A review of the spider-attacking Polysphincta dizardi species-group (Hymenoptera, Ichneumonidae, Pimplinae), with descriptions of seven new species from South America. ZooKeys 1041 137-165.

Provancher L. (1873) Les Ichneumonides de Québec avec description de plusieurs especès nouvelles. Le Naturaliste Canadien 5 470-477.

Ratzeburg J.T.C. (1844) Die Ichneumonen der Forstinsecten in forstlicher und entomologischer Beziehung. Berlin, Germany. 224 pp.

Scudder S.H. (1878) The insects of the tertiary beds at Quesnel. Report of Progress. Geological Survey Canada 1876-1877 266-280.

Spasojevic T., Broad G.R. & Klopfstein S. (2022) Revision of 18 ichneumonid fossil species (Hymenoptera, Ichneumonidae) highlights the need for open nomenclature in palaeontology. Fossil Record 25 187-212.

Tschek C. (1869) Beiträge zur Kenntniss der österreichischen Pimplarien. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 18 269-280.

Varga O. & Reshchikov A. (2015) New records of the genus Polysphincta Gravenhorst, 1829 (Hymenoptera: Ichneumonidae: Pimplinae) from the Oriental region. Zootaxa 3955 435-443.

22. Genus Reclinervellus He & Ye, 1998 4 spp.

He J.H. & Ye S.F. (1998) Reclinervellus, a new genus of Polysphinctini (Hymenoptera: Ichneumonidae: Pimplinae) from China. Entomotaxonomia 20 153-156 (in Chinese with English summary).

Reclinervellus dorsiconcavus He & Ye, 1998

Reclinervellus masumotoi Matsumoto & Konishi, 2007

Reclinervellus nielseni (Roman, 1923)

Reclinervellus tuberculatus (Uchida, 1932)

He J.H. & Ye S.F. (1998) Reclinervellus, a new genus of Polysphinctini (Hymenoptera: Ichneumonidae: Pimplinae) from China. Entomotaxonomia 20 153-156 (in Chinese with English summary).

Matsumoto R. & Konishi K. (2007) Life histories of two ichneumonid parasitoids of Cyclosa octotuberculata (Araneae): Reclinervellus tuberculatus (Uchida) and its new sympatric congener (Hymenoptera: Ichneumonidae: Pimplinae). Entomological Science 10 267-278.

Roman A. (1923) Nya Polysphincta-former ur de Nielsen'ska kläckningarne. Entomologiske Meddelelser 14 206-210.

Uchida T. (1932) Beiträge zur Kenntnis der japanischen Ichneumoniden. Insecta Matsumurana 6 145-168.

23. Genus Sinarachna Townes, 1960 4 spp. + 1 subspecies

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Sinarachna ceylonica (Ashmead, 1896)

Sinarachna maculata He & Ye, 2006

Sinarachna nigricornis (Holmgren, 1860)

Sinarachna pallipes (Holmgren, 1860)

Sinarachna pallipes strigis (Howard, 1892)

He J.H., Chen X.X., Ma Y. & Ye S.F. (2006) Ichneumonidae. In: Z. Li & D. Jin (eds.). Insects from Fanjingshan landscape (Insect Fauna from National Nature Reserve in Guizhow Province VI). pp. 571-575. Guizhow Science and Technology Publishing House, Guiyang, Guizhou, China.

Holmgren A.E. (1860) Försök till uppställning och beskrifning af Sveriges Ichneumonider. Tredje Serien. Fam. Pimplariae. (Monographia Pimplariarum Sueciae). Vetenskapsakademiens Handlingar, (B) 3 1-76.

Howard L.O. (1892) The hymenopterous parasites of spiders. Proceedings of the Entomological Society of Washington 2 290-302.

Howard L.O. & Ashmead W.H. (1896) On some reared parasitic hymenopterous insects from Ceylon. Proceedings of the United States National Museum 18 633-648.

24. Genus Ticapimpla Gauld, 1991 5 spp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Ticapimpla amazonica Palacio, Broad, Sääksjärvi & Veijalainen, 2010

Ticapimpla carinata Palacio, Broad, Sääksjärvi & Veijalainen, 2010

Ticapimpla matamatae Palacio, Broad, Sääksjärvi & Veijalainen, 2010

Ticapimpla soinii Palacio, Broad, Sääksjärvi & Veijalainen, 2010

Ticapimpla vilmae Gauld, 1991

Palacio E., Broad G.R., Sääksjärvi I.E. & Veijalainen A. (2010) Western Amazonian Ticapimpla (Hymenoptera: Ichneumonidae: Pimplinae): four new species from Colombia, Ecuador, and Peru, with a key to species of the genus. Psyche: A Journal of Entomology 2010 Article ID 161595.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

25. Genus Zatypota Förster, 1869 54 spp. + 8 subspecies

Förster A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 25 135-221.

Zatypota albicoxa (Walker, 1874)

Zatypota albicoxa meridicolor Aubert, 1989

Zatypota alborhombarta (Davis, 1895)

Zatypota anomala (Holmgren, 1860)

Zatypota arizonica Townes, 1960

Zatypota baezae Pádua & Sobczak, 2019

Zatypota baragi Matsumoto, 2010

Zatypota bayamensis Fernández-Triana, 2007

Zatypota bingili Gauld, 1984

Zatypota bohemani (Holmgren, 1860)

Zatypota brachycera Matsumoto, 2010

Zatypota capicola Benoit, 1959

Zatypota celer Gauld, 1984

Zatypota chryssophaga Matsumoto, 2010

Zatypota cingulata Townes, 1960

Zatypota crassipes Townes, 1960

Zatypota dandiensis Gauld, 1984

Zatypota dendrobia Matsumoto, 2010

Zatypota dichroa (Marshall, 1892)

Zatypota discolor (Holmgren, 1860)

Zatypota discolor helveticator Aubert, 1989

Zatypota discolor nigrithorax Constantineanu, Ciochia, Constantineanu, Mustata & Ularu, 1967

Zatypota discolor obscura (Roman, 1923)

Zatypota elegans Matsumoto, 2010

Zatypota exilis Townes, 1960

Zatypota favosa Townes, 1960

Zatypota flamma Fritzén, 2014

Zatypota fonsecai Gauld, 1991

Zatypota gracilipes Uchida & Momoi, 1958

Zatypota inexpectata (Seyrig, 1932)

Zatypota kauros Gauld, 1984

Zatypota kerstinae Fritzén, 2010

Zatypota lagiralda Gauld Menjivar, Monro & Gonzalez, 2002

Zatypota luteipes Townes, 1960

Zatypota maculata Matsumoto & Takasuka, 2010

Zatypota medranoi Gauld, 1991

Zatypota mongolica Sedivy, 1971

Zatypota morsei Gauld, 1991

Zatypota mulunguensis Pádua & Sobczak, 2019

Zatypota orbitalis Varga, 2025

Zatypota pallipes Schmiedeknecht, 1888

Zatypota patellata Townes, 1960

Zatypota percontatoria (Müller, 1776)

Zatypota percontatoria celticator Aubert, 1989

Zatypota percontatoria meridianator Aubert, 1989

Zatypota petronae Gauld, 1991

Zatypota phraxos Gauld, 1984

Zatypota picticollis (Thomson, 1888)

Zatypota picticollis gallicator Aubert, 1989

Zatypota picticollis meridiator Aubert, 1989

Zatypota prima Benoit, 1953

Zatypota rennefer Gauld, 1984

Zatypota riverai Gauld, 1991

Zatypota solanoi Gauld, 1991

Zatypota stellata Gauld, 1984

Zatypota sulcata Matsumoto, 2010

Zatypota takayu Matsumoto & Takasuka, 2010

Zatypota talamancae Gauld, Ugalde Gómez & Hanson, 1998

Zatypota tropica (Morley, 1912)

Zatypota velata Gauld, 1984

Zatypota walleyi Townes, 1960

Zatypota yambar Matsumoto, 2010

Aubert J.F. (1989) Ichneumonides non pétiolées inédites et quatrième suppl. aux Scolobatinae (Ctenopelmatinae): les Homaspis Foerst. Bulletin de la Société Entomologique de Mulhouse 1989 1-11.

Benoit P.L.G. (1953) Notes Ichneumonologiques Africaines IV. Revue de Zoologie et de Botanique Africaines 47 153-159.

Benoit P.L.G. (1959) Hymenoptera: Ichneumonidae (Banchinae, Bassinae, Pimplinae et Tryphoninae). South African Animal Life 6 441-482.

Constantineanu M.I., Ciochia V., Constantineanu R.M., Mustata G. & Ularu G. (1967) Contributions a l'etude des Ichneumonides de la montagne de Gutii, région de Maramures. Analele ştiinţifice ale Universităţii “Al. I. Cuza” din Iaşi. Sect. II a 13 47-61.

Davis G.C. (1895) Two new species of Clistopyga. Entomological News 6 198-199.

Fernández-Triana J.L. (2007) Notes on Cuban Pimplinae (Hymenoptera: Ichneumonidae), with the description of two new species. Boletin de la Sea 40 211-214.

Fritzén N.R. (2010) Natural history and description of Zatypota kerstinae sp.nov (Hymenoptera: Ichneumonidae) reared from Theridion palmgreni Marusik et Tsellarius (Araneae: Theridiidae) in Finland. Zootaxa 2487 52-60.

Fritzén N.R. (2014) Two new species of the Polysphincta genus-group (Hymenoptera: Ichneumonidae: Pimplinae) reared from their spider (Araneae) hosts in Europe. Zootaxa 3894 117-130.

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biologia Tropical 46 (Suppl. 1) 1-189.

Gauld I.D., Menjivar R.U.H., Monro A.K. & González M.O. (2002) Guía para la identificación de los Pimplinae de cafetales bajo sombra de El Salvador (Hymenoptera: Ichneumonidae). Natural History Museum, London, U.K. 76 pp.

Holmgren A.E. (1860) Försök till uppställning och beskrifning af Sveriges Ichneumonider. Tredje Serien. Fam. Pimplariae. (Monographia Pimplariarum Sueciae). Vetenskapsakademiens Handlingar, (B) 3 1-76.

Marshall T.A. (1892) Énumerations de quelques Hyménoptères du Venezuela. Annales de la Société Entomologique de France 61 60-76.

Matsumoto R. & Takasuka K. (2010) A revision of the genus Zatypota Förster of Japan, with descriptions of nine new species and notes on their hosts (Hymenoptera: Ichneumonidae: Pimplinae). Zootaxa 2522 1-43.

Morley C. (1912) No. XII. Hymenoptera, Ichneumonidae. In: J.S. Gardiner (eds.). Transactions of the Linnean Society of London 15. pp. 169-179. Linnean Society of London, London, U.K.

Müller O.F. (1776) Zoologiae Danicae prodromus, seu anumalium Daniae et Norvegiae indigenarum characteres, nomina et synomyma imprimis popularium. Hafniae, 282 pp.

Roman A. (1923) Nya Polysphincta-former ur de Nielsen'ska kläckningarne. Entomologiske Meddelelser 14 206-210.

Schmiedeknecht O. (1888) Die europäischen Gattungen der Schlupfwespen Familie Pimplariae. Zoologische Jahrbücher Abteilung für Systematik 3 419-444.

Sedivy J. (1971) Ergebnisse der mongolisch-tschechoslowakischen entomologisch-botanischen Expeditionen in der Mongolei: 24. Hymenoptera, Ichneumonidae. Acta Faunistica Entomologica Musei Nationalis Pragae 14 73-91.

Seyrig A. (1932) Les Ichneumonides de Madagascar. I Ichneumonidae Pimplinae. Mémoires de l'Académie Malgache. Fascicule 11 1-183.

Sobczak J.F., Pádua D.G., Villanueva- Bonilla G.A., De Sousa Nóbrega F.A. & Messas Y.F. (2019) Two new species of Zatypota (Hymenoptera: Ichneumonidae, Pimplinae) sharing the same host spider in Northeast Brazil. Zootaxa 4609 169-177.

Thomson C.G. (1888) XXXVII. Bidrag till Sveriges insectfauna. Opuscula Entomologica. Lund. XII 1202-1265.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. & Momoi S. (1958) On the species of Polysphincta Gravenhorst and Zatypota Förster from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 22-30.

Varga O. (2025) A new species of the genus Zatypota Förster, 1869 (Hymenoptera: Ichneumonidae: Pimplinae) from South Africa. Journal of Insect Biodiversity 61 58-61. 

Walker F. (1874) Descriptions of some Japanese Hymenoptera. Cistula Entomologica 1 301-310.

クモフシヒメバチ Pseudo-parasitoids of spider egg masses（直近祖先群・クモ卵塊寄生バチ）3属の種一覧

 1. Genus Clistopyga Gravenhorst, 1829 95 spp. + 1 subspecies

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Clistopyga aenigmatica Bordera & González-Moreno 2014

Clistopyga africana Benoit, 1956

Clistopyga albovittata Bordera & Palacio, 2019

Clistopyga alutaria Townes, 1960

Clistopyga amazonica Bordera & Sääksjärvi, 2016

Clistopyga andaki Palacio & Bordera, 2025

Clistopyga andina Bordera & Palacio, 2025

Clistopyga angustipleura Bordera & Palacio, 2025

Clistopyga arctica Kusigemati, 1985

Clistopyga arhuaca Palacio & Bordera, 2025

Clistopyga arizonica Bordera & González-Moreno, 2014

Clistopyga atrata Cushman, 1922

Clistopyga aurantia Bordera & Palacio, 2025

Clistopyga brevicaudata Bordera & Palacio, 2025

Clistopyga californica Khalaim & Hernández, 2008

Clistopyga calixtoi Gauld, 1991

Clistopyga canadensis Provancher, 1880

Clistopyga caramba Castillo & Sääksjärvi, 2015

Clistopyga carinata Bordera & Palacio, 2019

Clistopyga carvajali Gauld, Ugalde Gómez & Hanson, 1998

Clistopyga catarina Bordera & Palacio, 2019

Clistopyga chaconi Gauld, 1991

Clistopyga cinnamoptera Bordera & Sääksjärvi, 2016

Clistopyga crassicaudata Sääksjärvi, Bordera & Palacio, 2018

Clistopyga cuscoensis Bordera, Castillo & Sääksjärvi, 2016

Clistopyga declinata Palacio & Bordera, 2019

Clistopyga diazi Porter, 1979

Clistopyga eldae Gauld, 1991

Clistopyga emphera Kusigemati, 1985

Clistopyga femorator Bordera & Palacio, 2025

Clistopyga fernandezi Gauld, 1991

Clistopyga ghatensis Manjusha, Sudheer & Ghosh, 2019

Clistopyga hayesiana Bordera & Sääksjärvi, 2016

Clistopyga henryi Gauld, 1991

Clistopyga horcoana Bordera & Palacio, 2025

Clistopyga inca Bordera & Palacio, 2025

Clistopyga incitator (Fabricius, 1793)

Clistopyga incitator mongolica Kasparyan, 2007

Clistopyga inti Bordera & Palacio, 2025

Clistopyga isabellae Bordera, 2025

Clistopyga isayae Díaz, Bordera & Palacio, 2018

Clistopyga jakobii Graf, 1985

Clistopyga juliana Bordera & González-Moreno 2014

Clistopyga kalima Palacio, 2018

Clistopyga kogabae Palacio & Bordera, 2025

Clistopyga laevis Kasparyan, 1981

Clistopyga lapacensis Bordera & Palacio, 2019

Clistopyga laplatense Bordera & Palacio, 2025

Clistopyga latifrontalis (Uchida, 1941)

Clistopyga linearis (Wollaston, 1858)

Clistopyga lobulata Palacio & Bordera, 2025

Clistopyga longifemoralis Varga & Reshchikov, 2015

Clistopyga lopezrichinii (Blanchard, 1941)

Clistopyga maculifrons Cushman, 1922

Clistopyga magninsula Higa, Victorino, Dias & Dias, 2024

Clistopyga manni Cushman, 1922

Clistopyga marcapatensis Bordera & Palacio, 2019

Clistopyga maya González-Moreno & Bordera 2014

Clistopyga melanoptera Castillo, Sääksjärvi & Bordera, 2016

Clistopyga meridae Bordera & Palacio, 2025

Clistopyga misionensis Bordera & Sääksjärvi, 2016

Clistopyga mocaguae Palacio & Bordera, 2016

Clistopyga mochica Bordera & Palacio, 2025

Clistopyga moraviae Gauld, Ugalde Gómez & Hanson, 1998

Clistopyga muisca Palacio & Bordera, 2025

Clistopyga nasa Palacio & Bordera, 2025

Clistopyga nigrifrons Cushman, 1922

Clistopyga nigriventris Palacio, 2018

Clistopyga nubiphila Palacio & Bordera, 2025

Clistopyga oaxacana González-Moreno & Bordera 2014

Clistopyga orellanae Bordera & Sääksjärvi, 2016

Clistopyga panchei Palacio, 2018

Clistopyga peruandina Sääksjärvi & Pádua 2025

Clistopyga plana Morley, 1914

Clistopyga plaumanni Bordera & Palacio, 2025

Clistopyga polita Bordera & Palacio, 2019

Clistopyga porteri Bordera & Sääksjärvi, 2016

Clistopyga pseudoclistopyga (Varga, 2018)

Clistopyga quechua Bordera & Palacio, 2025

Clistopyga recurva (Say, 1835)

Clistopyga rondoniae Bordera & Sääksjärvi, 2016

Clistopyga rufator Holmgren, 1856

Clistopyga rugulosa Kusigemati, 1985

Clistopyga serricauda Khalaim & Hernández, 2008

Clistopyga speculata Bordera & Palacio, 2019

Clistopyga splendida Palacio & Bordera, 2018

Clistopyga stanfordi Gauld, 1991

Clistopyga subtenella Bordera & Palacio, 2025

Clistopyga sziladyi Kiss, 1959

Clistopyga taironae Palacio, Bordera & Díaz, 2018

Clistopyga telamoniae Ranjith and Sushama 2026

Clistopyga teresitae Díaz, Palacio & Bordera, 2019

Clistopyga teresopolitana Pádua 2025

Clistopyga valida Bordera & Palacio, 2025

Clistopyga wounan Palacio & Bordera, 2025

Clistopyga yabuquensis Bordera & Sääksjärvi, 2016

Bajáry N.E. (1959) Revision der Ichneumoniden-Typen von Kiss und Szépligeti II (Hymenoptera). Annales Historico-Naturales Musei Nationalis Hungarici 51 415-418.

Benoit P.L.G. (1956) Ichneumonidae nouveaux ou interessants de l'Afrique du Sud. Annals of the South African Museum 43 123-135.

Bordera S., González-Moreno A., Khalaim A.I. & Sääksjärvi I.E. (2014) Revision of North American species of Clistopyga (Hymenoptera: Ichneumonidae: Pimplinae). The Canadian Entomologist 146 355-414.

Bordera S., Sääksjärvi I.E., Castillo C., Palacio E. & González-Moreno A. (2016) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part I: the C. chaconi species group, with the description of eleven new species. European Journal of Taxonomy 206 1-37.

Bordera S. & Palacio E. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part IV: the C. eldae species group, with the description of three new species. Zootaxa 4564 327-346.

Bordera S., Palacio E. & Martínez J.J. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part V: the C. diazi species group, with the description of three new species. Zootaxa 4661 545-565.

Bordera S., Palacio E. & Herrera-Flórez A.F. (2025) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part VI: the C. calixtoi species group, with the description of twenty-four new species. Zootaxa 5662 1-115. 

Cushman R.A. (1922) North American Ichneumon-flies of the genera Clistopyga and Schizopyga. Proceedings of the United States National Museum 60 1-14.

Fabricius J.C. (1793) Entomologia systematica emendata et aucta. Tom. II. Hafniae, 519 pp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biologia Tropical 46 (Suppl. 1) 1-189.

Graf V. (1985) Ichneumofauna do sudeste e sul do Brasil. VI. Nova espécie de Clistopyga (Ephialtinae, Hymenoptera). Revista Brasileira de Entomologia 29 349-350.

Higa P.T., Victorino B.I., Dias M.M. & Dias A.M.P. (2024) New species and new records of Clistopyga (Ichneumonidae: Pimplinae) in Brazil. Zootaxa 5523 354-362.

Holmgren A.E. (1856) Entomologiska anteckningar under en resa i södra Sverige ar 1854. Kongliga Svenska Vetenskapsakademiens Handlingar 75 1-104.

Kasparyan D.R. & Khalaim A.I. (2007) [1. Subfamily Pimplinae]. In: A.S. Lelej (eds.). [Key to the insects of the Russian Far East. Vol. IV. Neuropteroidea, Mecoptera, Hymenoptera. Pt. 5]. pp. 279-333 (in Russian). Vladivostok, Russia.

Khalaim A.I. & Hernández S.A. (2008) Review of the genus Clistopyga Grav., 1829 of Mexico (Hymenoptera: Ichneumonidae: Pimplinae). Russian Entomological Journal 17 311-315.

Kusigemati K. (1985) Three new species of Clistopyga from Japan (Hymenoptera: Ichneumonidae). Memoirs of the Faculty of Agriculture, Kagoshima University 21 189-194.

Manjusha B.M., Sudheer K. & Ghosh S.M. (2019) A new species of Clistopyga Gravenhorst (Hymenoptera: Ichneumonidae: Pimplinae) from the Western Ghats. Records of the Zoological Survey of India 119 60-63.

Morley C. (1914) A Revision of the Ichneumonidae Based on the Collection in the British Museum (Natural History) Part III. Tribes Pimplides and Bassides. British Museum, London, U.K. 148 pp.

Pádua D.G., Sääksjärvi I.E., Kaunisto K.M. & Monteiro R.F. (2025) Two new species of the Clistopyga henryi species-group (Hymenoptera, Ichneumonidae, Pimplinae) from South America, with a key to species of the group. ZooKeys 1260 23-36. 

Palacio E., Bordera S., Sääksjärvi I.E. & Díaz F. (2018) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part II: the C. isayae species group, with the description of seven new species. Zootaxa 4442 101-121.

Palacio E., Bordera S. & Díaz F. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae) Part III: the C. henryi species group, with the description of three new species. Zootaxa 4563 103-118.

Porter C.C. (1979) Ichneumonidae de Tarapacá. I. Subfamilia Ephialtinae (Hymenoptera). Idesia 5 157-187.

Provancher L. (1880) Faune canadienne. Les insectes - Hyménoptères. Naturaliste Canadien 12 33-48.

Sääksjärvi I.E., Castillo C., Bordera S. & Broad G.R. (2015) Clistopyga caramba sp. nov. (Hymenoptera: Ichneumonidae; Pimplinae), an astonishing example of mimicry in spider-attacking parasitoid wasps. Zootaxa 4013 287-292.

Say T. (1835) Descriptions of new North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1 210-305.

Sushama V., Ranjith A.P., Rajmohana K., Sen S. & Dinesh K.P. (2026) A new species of Clistopyga Gravenhorst (Hymenoptera: Ichneumonidae: Pimplinae) parasitizing Telamonia dimidiata (Simon), with notes on host association and biology. Journal of Natural History 60 165-175. 

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. (1941) Beiträge zur Systematik der Tribus Polysphinctini Japans. Insecta Matsumurana 15 112-122.

Varga O. & Reshchikov A. (2015) New records of the genus Clistopyga Gravenhorst, 1829 (Hymenoptera: Ichneumonidae: Pimplinae) from the Oriental region, with description of a new species. Zootaxa 3964 561-568.

Varga O. (2018) Afroanomalia pseudoclistopyga, a new genus and species of pimpline parasitoid wasp (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region. Zootaxa 4461 146-150.

2. Genus Tromatobia Förster, 1869 33 spp. + 3 subspecies

Förster A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 25 135-221.

Tromatobia argiopei Uchida, 1941

Tromatobia argiopes (Marshall, 1892)

Tromatobia blancoi Gauld, 1991

Tromatobia carballoi Gauld, Ugalde Gómez & Hanson, 1998

Tromatobia commutabilis Förster, 1888

Tromatobia dilutula (Benoit, 1953)

Tromatobia flavistellata Uchida & Momoi, 1957

Tromatobia forsiusi (Hellén, 1915)

Tromatobia huebrichi (Brèthes, 1913)

Tromatobia hutacharerni Kusigemati, 1988

Tromatobia koehleri (Blanchard, 1941)

Tromatobia lineata (Wollaston, 1858)

Tromatobia lineatoria (Villers, 1789)

Tromatobia lineatoria decorata Obrtel, 1953

Tromatobia lineatoria rubella (Costa, 1886)

Tromatobia lineiger Morley, 1914

Tromatobia maculata Momoi, 1970

Tromatobia nipponica Uchida, 1928

Tromatobia notator (Fabricius, 1804)

Tromatobia onerosa Förster, 1888

Tromatobia ornata (Gravenhorst, 1829)

Tromatobia ornata nigrescens Constantineanu, Ciochia, Constantineanu, Mustata & Ularu, 1967

Tromatobia ovivora (Boheman, 1821)

Tromatobia pacayae Gauld, Ugalde Gómez & Hanson, 1998

Tromatobia pacifica Momoi, 1966

Tromatobia quadricolor (Kriechbaumer, 1894)

Tromatobia rufopectus (Cresson, 1870)

Tromatobia sejuncta Förster, 1888

Tromatobia sponsa (Haliday, 1836)

Tromatobia sulcata (Benoit, 1953)

Tromatobia taiwana Kusigemati, 1984

Tromatobia tricolor (Brullé, 1846)

Tromatobia ulatei Gauld, Ugalde Gómez & Hanson, 1998

Tromatobia variabilis (Holmgren, 1856)

Tromatobia zonata (Davis, 1895)

Tromatobia zunigai Gauld, 1991

Benoit P.L.G. (1953) Notes Ichneumonologiques Africaines IV. Revue de Zoologie et de Botanique Africaines 47 153-159.

Blanchard E.E. (1942) Nuevos parásitos del bicho de cesto Oeceticus kirbyi Guild. Revista de la Sociedad Entomologica Argentina 11 3-21.

Boheman C.H. (1821) En ny art af Insect-slägtet Pimpla, hvars larv uppehåller sig i Spindelbon. Kungliga Svenska Vetenskaps Akadamiens Handlingar 42 335-337.

Brèthes J. (1913) Hymenópteros de la América meridional. Anales del Museo Nacional de Historia Natural de Buenos Aires 24 35-165.

Brullé M.A. (1846) Tome Quatrième. Des Hyménoptères. Les Ichneumonides. In: A. Lepeletier de Saint Fargeau (eds.). Histoire Naturelle Des Insectes. Hyménoptères. pp. 56-521. Paris, France.

Costa A. (1886) Notizie ed osservazioni sulla Geo-Fauna sarda Memoria Quinta Risultamento delle ricerche fatte in maggio 1885. Atti della Reale Accademia delle Scienze Fisiche e Matematiche di Napoli, ser.2a:vol.II:fasc.7, 24 pp.

Constantineanu M.I., Ciochia V., Constantineanu R.M., Mustata G. & Ularu G. (1967) Contributions a l'etude des Ichneumonides de la montagne de Gutii, région de Maramures. Analele ştiinţifice ale Universităţii “Al. I. Cuza” din Iaşi. Sect. II a 13 47-61.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Davis G.C. (1895) Two new species of Clistopyga. Entomological News 6 198-199.

Fabricius J.C. (1804) Systema Piezatorum: secundum ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. Carolum Reichard, Brunsvigae, 439+32 pp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biologia Tropical 46 (Suppl. 1) 1-189.

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Haliday A.H. (1836) Descriptions, &c. of the Hymenoptera. In: S. Curtis, A.H. Haliday & F. Walker (eds.). Descriptions &c. of the Insects Collected by Captain P.P. King, R.N., F.R.S., in the Survey of the Straits of Magellan. Transactions of the Linnean Society of London. pp. 316-331. London, U.K.

Hellén W. (1915) Beiträge zur Kenntnis der Ichneumoniden Finlands I. Subfamily Pimplinae. Acta Societatis pro Fauna et Flora Fennica 40 1-89.

Holmgren A.E. (1856) Entomologiska anteckningar under en resa i södra Sverige ar 1854. Kongliga Svenska Vetenskapsakademiens Handlingar 75 1-104.

Kriechbaumer J. (1894) Hymenoptera Ichneumonidea a medico nautico Dr. Joh. Brauns in itinere ad oras Africae occidentalis lecta. Berliner Entomologische Zeitschrift 39 43-68.

Kusigemati K. (1984) Some Ephialtinae of South East Asia, with descriptions of eleven new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 5 126-150.

Kusigemati K. (1988) Descriptions of four new ichneumon flies parasitic on pine insect pests in Thailand. Memoirs of the Faculty of Agriculture, Kagoshima University 24 147-155.

Marshall T.A. (1892) Énumerations de quelques Hyménoptères du Venezuela. Annales de la Société Entomologique de France 61 60-76.

Momoi S. (1966) Some new Ichneumonidae (Hymenoptera) from New Guinea and adjacent areas. Pacific Insects 8 152-164.

Momoi S. (1970) Ichneumonidae (Hymenoptera) of the Ryukyu Archipelago. Pacific Insects 12 327-399.

Morley C. (1914) A Revision of the Ichneumonidae Based on the Collection in the British Museum (Natural History) Part III. Tribes Pimplides and Bassides. British Museum, London, U.K. 148 pp.

Obrtel R. (1953) [Notes on some interesting Icheumonidae in Silesia (Hym., Ichn.)]. Přírodovědecký sborník Ostravského kraje 14 192-208 (in Czech with English summary].

Schmiedeknecht O. (1888) Monographische Bearbeitung der Gattung Pimpla. Zoologische Jahrbücher Abteilung für Systematik 3 445-542.

Uchida T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido Imperial University 25 1-115.

Uchida T. (1941) Eine neue Schmarotzerwespe von Spinnen. Insecta Matsumurana 15 161-162.

Uchida T. & Momoi S. (1957) Descriptions of three new species of the tribe Ephialtini from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 21 6-11.

Villers C. de (1789) Caroli Linnaei entomologia, Faunae Suecicae descriptionibus. Tomus tertius. Lugduni, 657 pp.

Wollaston T.V. (1858) Brief diagnostic characters of undescribed Madeiran insects. Annals and Magazine of Natural History ser.3:vol.1 18-28.

3. Genus Zaglyptus Förster, 1869 27 spp. + 7 subspecies

Förster A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 25 135-221.

Zaglyptus ankaratrus Seyrig, 1934

Zaglyptus arizonicus Townes, 1960

Zaglyptus chavesi Gauld, 1991

Zaglyptus cortesi Porter, 1979

Zaglyptus divaricatus Baltazar, 1961

Zaglyptus divaricatus basilanicus Baltazar, 1961

Zaglyptus facifidus Baltazar, 1961

Zaglyptus formosus Cushman, 1933

Zaglyptus glaber Gupta, 1961

Zaglyptus glabrinotus (Girault, 1925)

Zaglyptus grandis Gupta, 1961

Zaglyptus guptai Pham, Broad & Wägele, 2013

Zaglyptus hallowayi Gauld, 1984

Zaglyptus idukkiensis (Manjusha, Sudheer & Ghosh, 2019)

Zaglyptus indicus Gupta, 1961

Zaglyptus indicus nigrithorax Gupta, 1961

Zaglyptus iwatai (Uchida, 1936)

Zaglyptus miarus Gupta, 1961

Zaglyptus multicolor (Gravenhorst, 1829)

Zaglyptus multicolor rufescens (Boyer de Fonscolombe, 1854)

Zaglyptus nigrolineatus Gupta, 1961

Zaglyptus pictilis Townes, 1960

Zaglyptus romeroae Gauld, 1991

Zaglyptus rufus Hellén, 1949

Zaglyptus ruslani Rizki & Idris, 2010

Zaglyptus semirufus Momoi, 1970

Zaglyptus semirufus marginatus Kasparyan, 1981

Zaglyptus simonis (Marshall, 1892)

Zaglyptus trituberculatus (Benoit, 1953)

Zaglyptus varipes (Gravenhorst, 1829)

Zaglyptus varipes albitegulis Constantineanu & Ciochia, 1968

Zaglyptus varipes incompletus (Cresson, 1870)

Zaglyptus varipes meridionator Aubert, 1969

Zaglyptus wuyiensis He, 1984

Aubert J.F. (1969) Ichneumonides non petiolees inédites et révision partielle des genres Phytodietus Grav. et Netelia Gray. Bulletin de la Société Entomologique de Mulhouse 1968(novembre-decembre) 93-103.

Baltazar C.R. (1961) The Philippine Pimplini, Poemeniini, Rhyssini, and Xoridini (Hymenoptera, Ichneumonidæ, Pimplinæ). Monographs of the National Institute of Science and Technology 7 1-130.

Benoit P.L.G. (1953) Notes Ichneumonologiques Africaines IV. Revue de Zoologie et de Botanique Africaines 47 153-159.

Boyer de Fonscolombe M. (1854) Ichneumonologie provençale, ou catalogue des ichneumoisïdes qui se trouvent aux environs d'Aix, et description des espèces inédites. Annales de la Société Entomologique de France ser.3:t.2 479-520.

Constantineanu M.I. & Ciochia V. (1968) Ichneumonides nouveaux de Roumanie. Revue Roumaine de Biologie (Serie de Zoologie) 13 43-47.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Cushman R.A. (1933) H. Sauter's Formosa-collection: Subfamily Ichneumoninae (Pimplinae of Ashmaed). Insecta Matsumurana 8 1-50.

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Girault A.A. (1925) Records and distributions of Australian Ophioninae (Ichneumon-flies). Queensland Agricultural Journal 24 538-541.

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Gupta V.K. (1960) A revision of the Oriental species of the genus Zaglyptus (Hymenoptera: Ichneumonidae). Indian Journal of Entomology 22 244-257.

He J.H. (1984) [Notes on Chinese species of the genus Zaglyptus Förster, with descriptions of a new species and a new species-group (Hymenoptera: Ichneumonidae)]. Wuyi Science Journal 4 199-204 (in Chinese with English summary).

Hellén W. (1949) Zur Kenntnis der Ichneumonidenfauna der Atlantischen Inseln. Commentationes Biologicae Societas Scientiarum Fennica 8 1-23.

Kasparyan D.R. (1981) [New species of the subfamily Pimplinae (Hymenoptera, Ichneumonidae), parasites of spiders in the Far East of the USSR.]. Trudy Zoologicheskogo Instituta 92 108-112 (in Russian).

Manjusha B.M., Sudheer K. & Ghosh S.M. (2019) Polysphincta idukkiensis (Hymenoptera: Ichneumonidae: Pimplinae) a rare new species from the southern Western Ghats. Halteres 10 96-99.

Marshall T.A. (1892) Énumerations de quelques Hyménoptères du Venezuela. Bulletin de la Societe Entomologique de France 61 60-76.

Momoi S. (1970) Ichneumonidae (Hymenoptera) of the Ryukyu Archipelago. Pacific Insects 12 327-399.

Pham N.T., Broad G.R. & Wägele W. (2013) First record of the genus Zaglyptus Förster (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam, with description of one new species. Zootaxa 3702 483-492.

Porter C.C. (1979) Ichneumonidae de Tarapacá. I. Subfamilia Ephialtinae (Hymenoptera). Idesia 5 157-187.

Rizki A. & Idris A.B. (2010) One new species and one new record of the genus Zaglyptus Foerster (Hymenoptera: Ichneumonidae: Pimplinae) from Malaysia. Serangga 15 9-24.

Seyrig A. (1934) Les Ichneumonides de Madagascar. 2. Ichneumonidae Tryphoninae et supplément aux 1. Pimplinae. Mémoires de I'Académie Malgache 19 1-111.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. (1936) Drei neue Gattungen sowie acht neue und fuenf unbeschriebene Arten der Ichneumoniden aus Japan. Insecta Matsumurana 10 111-122.

Taxapad 2016以降に記載あるいは分類学的処理をされた種

Taxonomic changes or newly described species since Taxapad 2016 (Yu et al. 2016) until Takasuka & Broad (2024), reflected in the published global checklist (T&B 2024)

Bordera S., Sääksjärvi I.E., Castillo C., Palacio E. & González-Moreno A. (2016) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part I: the C. chaconi species group, with the description of eleven new species. European Journal of Taxonomy 206 1-37.

Clistopyga amazonica Bordera & Sääksjärvi, 2016 sp. nov.

Clistopyga cinnamoptera Bordera & Sääksjärvi, 2016 sp. nov.

Clistopyga cuscoensis Bordera, Castillo & Sääksjärvi, 2016 sp. nov.

Clistopyga hayesiana Bordera & Sääksjärvi, 2016 sp. nov.

Clistopyga melanoptera Castillo, Sääksjärvi & Bordera, 2016 sp. nov.

Clistopyga misionensis Bordera & Sääksjärvi, 2016 sp. nov.

Clistopyga mocaguae Palacio & Bordera, 2016 sp. nov.

Clistopyga orellanae Bordera & Sääksjärvi, 2016 sp. nov.

Clistopyga porteri Bordera & Sääksjärvi, 2016 sp. nov.

Clistopyga rondoniae Bordera & Sääksjärvi, 2016 sp. nov.

Clistopyga yabuquensis Bordera & Sääksjärvi, 2016  sp. nov.

Matsumoto, R. (2016) Molecular phylogeny and systematics of the Polysphincta group of genera (Hymenoptera, Ichneumonidae, Pimplinae). Systematic Entomology 41 854-864.

Dreisbachia resurrected from Schizopyga

Schizopyga flavifrontalis comb. nov. to Dreisbachia flavifrontalis (Uchida & Momoi, 1959)

Schizopyga punctata comb. nov. to Dreisbachia punctata (Uchida & Momoi, 1959)

Iania Matsumoto, 2016 gen. nov.

Schizopyga pictifrons comb. nov. to Iania pictifrons (Thomson, 1877)

Schizopyga pictifrons rufigaster comb. nov. to Iania pictifrons rufigaster (Constantineanu, Ciochia, Constantineanu & Ularu, 1967)

Acrodactyla inoperta comb. nov. to Megaetaira inoperta (Kusigemati, 1984)

Acrodactyla varicarinata comb. nov. to Megaetaira varicarinata (Uchida & Momoi, 1958)

Palacio E., Bordera S., Sääksjärvi I.E. & Díaz F. (2018) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part II: the C. isayae species group, with the description of seven new species. Zootaxa 4442 101-121.

Clistopyga crassicaudata Sääksjärvi, Bordera & Palacio, 2018 sp. nov.

Clistopyga isayae Díaz, Bordera & Palacio, 2018 sp. nov.

Clistopyga kalima Palacio, 2018 sp. nov.

Clistopyga nigriventris Palacio, 2018 sp. nov.

Clistopyga panchei Palacio, 2018 sp. nov.

Clistopyga splendida Palacio & Bordera, 2018 sp. nov.

Clistopyga taironae Palacio, Bordera & Díaz, 2018 sp. nov.

Varga O. (2018) Afroanomalia pseudoclistopyga, a new genus and species of pimpline parasitoid wasp (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region. Zootaxa 4461 146-150.

Afroanomalia Varga, 2018 gen. nov.

Afroanomalia pseudoclistopyga Varga, 2018 sp. nov.

Varga O. & Reshchikov A. (2018) Schizopyga alinae, a genus of pimpline parasitoid wasps (Hymenoptera: Ichneumonidae: Pimplinae) new to the Afrotropical region. Zootaxa 4422 291-295.

Schizopyga alnae Varga & Reshchikov, 2018 sp. nov.

Varga O., Reshchikov A. & Giovanni F.D. (2018) First record of the genus Brachyzapus Gauld & Dubois, 2006 (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region, with descriptions of two new species. Zootaxa 4377 581-586.

Brachyzapus glaber Varga & Reshchikov, 2018 sp. nov.

Brachyzapus greengoblin Varga & Di Giovanni, 2018 sp. nov.

Palacio E., Bordera S. & Díaz F. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae) Part III: the C. henryi species group, with the description of three new species. Zootaxa 4563 103-118.

Clistopyga carinata Bordera & Palacio, 2019 sp. nov.

Clistopyga declinata Palacio & Bordera, 2019 sp. nov.

Clistopyga teresitae Díaz, Palacio & Bordera, 2019 sp. nov.

Bordera S. & Palacio E. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part IV: the C. eldae species group, with the description of three new species. Zootaxa 4564 327-346.

Clistopyga catarina Bordera & Palacio, 2019 sp. nov.

Clistopyga marcapatensis Bordera & Palacio, 2019 sp. nov.

Clistopyga polita Bordera & Palacio, 2019 sp. nov.

Bordera S., Palacio E. & Martínez J.J. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part V: the C. diazi species group, with the description of three new species. Zootaxa 4661 545-565.

Clistopyga albovittata Bordera & Palacio, 2019 sp. nov.

Clistopyga lapacensis Bordera & Palacio, 2019 sp. nov.

Clistopyga speculata Bordera & Palacio, 2019 sp. nov.

Manjusha B.M., Sudheer K. & Ghosh S.M. (2019a) A new species of Clistopyga Gravenhorst (Hymenoptera: Ichneumonidae: Pimplinae) from the Western Ghats. Records of the Zoological Survey of India 119 60-63.

Clistopyga ghatensis Manjusha, Sudheer & Ghosh, 2019 sp. nov.

Manjusha B.M., Sudheer K. & Ghosh S.M. (2019b) Polysphincta idukkiensis (Hymenoptera: Ichneumonidae: Pimplinae) a rare new species from the southern Western Ghats. Halteres 10 96-99.

Polysphincta idukkiensis Manjusha, Sudheer & Ghosh, 2019 sp. nov. (later transferred to Zaglyptus)

Choi J.K., Kolarov J. & Lee J.W. (2019) Descriptions of two new species of Brachyzapus Gauld & Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from South Korea with key to a Palaearctic species. Zootaxa 4711 293-306.

Brachyzapus convergens Choi & Lee, 2019 sp. nov.

Brachyzapus pyramidalis Choi & Lee, 2019  sp. nov.

Sobczak J.F., Pádua D.G., Villanueva- Bonilla G.A., De Sousa Nóbrega F.A. & Messas Y.F. (2019) Two new species of Zatypota (Hymenoptera: Ichneumonidae, Pimplinae) sharing the same host spider in Northeast Brazil. Zootaxa 4609 169-177.

Zatypota baezae Pádua & Sobczak, 2019 sp. nov.

Zatypota mulunguensis Pádua & Sobczak, 2019 sp. nov.

Higa P.T. & Penteado-Dias A.M. (2020) Altitudinal effects on diversity of Pimplinae (Hymenoptera, Ichneumonidae) from Southeast Brazil and description of new species. Brazilian Journal of Biology 80 377-385.

Acrotaphus japi Higa & Penteado-Dias, 2020 sp. nov.

Polysphincta jundiai Higa & Penteado-Dias, 2020 sp. nov.

Polysphincta soaresi Higa & Penteado-Dias, 2020 sp. nov.

Pádua D.G., Sääksjärvi I.E., Monteiro R.F. & Oliveira M.L. (2020a) Review of the new world genus Acrotaphus Townes, 1960 (Hymenoptera: Ichneumonidae: Pimplinae), with descriptions of fifteen new species. Zootaxa 4719 1-62.

Acrotaphus amajari Pádua, 2020 sp. nov.

Acrotaphus amazonicus Pádua & Sääksjärvi, 2020 sp. nov.

Acrotaphus bodoquenaensis Pádua, 2020 sp. nov.

Acrotaphus cuzconus Pádua & Sääksjärvi, 2020 sp. nov.

Acrotaphus dolichopus Pádua, 2020 sp. nov.

Acrotaphus homeofranklini Pádua, 2020 sp. nov.

Acrotaphus jackiechani Pádua & Sääksjärvi, 2020 sp. nov.

Acrotaphus kourou Pádua & Sääksjärvi, 2020 sp. nov.

Acrotaphus micrus Pádua, 2020 sp. nov.

Acrotaphus monotaenius Pádua, 2020 sp. nov.

Acrotaphus nambilloensis Pádua, 2020 sp. nov.

Acrotaphus pseudoamazonicus Pádua & Sääksjärvi, 2020 sp. nov.

Acrotaphus pseudomexicanus Pádua, 2020 sp. nov.

Acrotaphus venezuelanus Pádua, 2020 sp. nov.

Acrotaphus wagnerianae Pádua, 2020 sp. nov.

Acrotaphus zampieronae Pádua, 2020 sp. nov.

Pádua D.G., Sääksjärvi I.E., Monteiro R.F. & Oliveira M.L. (2020b) Seven new species of spider-attacking Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Ecuador, French Guiana, and Peru, with an identification key to the world species. ZooKeys 935 57-92.

Hymenoepimecis andina Pádua & Sääksjärvi, 2020 sp. nov.

Hymenoepimecis castilloi Pádua & Sääksjärvi, 2020 sp. nov.

Hymenoepimecis dolichocarinata Pádua & Sääksjärvi, 2020 sp. nov.

Hymenoepimecis ecuatoriana Pádua & Sääksjärvi, 2020 sp. nov.

Hymenoepimecis longilobus Pádua & Sääksjärvi, 2020 sp. nov.

Hymenoepimecis pucallpina Pádua & Sääksjärvi, 2020 sp. nov.

Hymenoepimecis rafaelmartinezi Pádua & Sääksjärvi, 2020 sp. nov.

Pádua D.G. & Kloss T.G. (2020) A new species of Acrotaphus Townes (Hymenoptera: Ichneumonidae: Pimplinae) from the Brazilian Amazon, with notes on its host spider behavioral modification. Zootaxa 4810 161-168.

Acrotaphus japi Higa & Penteado-Dias, 2020 sp. nov.

Broad G.R. (2021) Taxonomic changes in Ichneumonoidea (Hymenoptera), and notes on certain type specimens. Zootaxa 4941 511–541.

Acrodactyla lachryma Pham, Broad, Matsumoto & Böhme, 2012 syn. nov. under Acrodactyla iliensis Sheng & Bian, 1996

Sinarachna tropica comb. nov. to Zatypota tropica (Morley 1912)

Pádua D.G., Sääksjärvi I.E., Spasojevic T., Kaunisto K.M., Monteiro R.F. & Oliveira M.L. (2021) A review of the spider-attacking Polysphincta dizardi species-group (Hymenoptera, Ichneumonidae, Pimplinae), with descriptions of seven new species from South America. ZooKeys 1041 137-165.

Polysphincta bonita Pádua & Sääksjärvi, 2021 sp. nov.

Polysphincta cosnipata Pádua & Sääksjärvi, 2021 sp. nov.

Polysphincta inca Pádua, Sääksjärvi & Spasojevic, 2021 sp. nov.

Polysphincta macroepomia Pádua & Sääksjärvi, 2021 sp. nov.

Polysphincta organensis Pádua & Sääksjärvi, 2021 sp. nov.

Polysphincta pichincha Pádua, Sääksjärvi & Spasojevic, 2021 sp. nov.

Polysphincta teresa Pádua & Sääksjärvi, 2021 sp. nov.

Kloss T.G., Pádua D.G., Almeida S.D.S., Penteado-Dias A.M., Mendes-Pereira T., Sobczak J.F., Lacerda F.G. & Gonzaga M.O. (2022) A new Darwin wasp (Hymenoptera: Ichneumonidae) and new records of behavioral manipulation of the host spider Leucauge volupis (Araneae: Tetragnathidae). Neotropical Entomology 51 821-829.

Hymenoepimecis pinheirensis Penteado-Dias & Pádua 2022 sp. nov.

Mohammadi-Khoramabadi A., Talebi A., Broad G.R. & Zwakhals K. (2022) A study of the genus Zabrachypus Cushman, 1920 (Hymenoptera Ichneumonidae Pimplinae) from Iran with descriptions of two new species. Redia 105 29-36.

Zabrachypus rubroabdominalis Mohammadi-Khoramabadi, Talebi & Broad, 2022 sp. nov.

Zabrachypus rugatus Mohammadi-Khoramabadi, Talebi & Broad, 2022 sp. nov.

Takasuka, K. & Broad, G.R. (2024) A bionomic overview of spider parasitoids and pseudo-parasitoids of the ichneumonid wasp subfamily Pimplinae. Contributions to Zoology 93 1-106.

Polysphincta idukkiensis comb. nov. to Zaglyptus idukkiensis (Manjusha, Sudheer & Ghosh, 2019) 

Taxonomic changes or newly described species since the global checklist was published (Takasuka & Broad 2024)

Khalaim A.I., RuÍz-Cancino E. & Coronado-Blanco J.M. (2024) Darwin wasps of the subfamily Pimplinae (Hymenoptera: Ichneumonidae) of Mexico: Polysphincta genus-group. Zootaxa 5458 151-196. 

Schizopyga anseli  comb. nov. to Dreisbachia anseli  (Fernandez-Triana, 2007)

Schizopyga moreirai comb. nov. to Dreisbachia moreirai (Loffredo & Penteado-Dias 2008)

Eruga yehi Gauld, 1991 syn. nov. under Eruga lineata Townes, 1960

Polysphincta sirena Khalaim, 2024 sp. nov.

Polysphincta xena Khalaim, 2024 sp. nov.

Higa P.T., Victorino B.I., Dias M.M. & Dias A.M.P. (2024) New species and new records of Clistopyga (Ichneumonidae: Pimplinae) in Brazil. Zootaxa 5523 354-362.

Clistopyga magninsula Higa, Victorino, Dias & Dias, 2024 sp. nov.

Bhat D.M., Ketelaere A.D. & Maqbool A. (2025) First record of polysphinctine genus Inbioia Gauld & Ugalde Gomez, 2002 (Hymenoptera: Ichneumonidae) from the Palearctic with description of a new species. Zootaxa 5618 424-432. 

Inbioia himalayensis Bhat, Keteraere & Maqbool, 2025 sp. nov.

Varga O. (2025) A new species of the genus Zatypota Förster, 1869 (Hymenoptera: Ichneumonidae: Pimplinae) from South Africa. Journal of Insect Biodiversity 61 58-61. 

Zatypota orbitalis Varga, 2025 sp. nov.

Bordera S., Palacio E. & Herrera-Flórez A.F. (2025) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part VI: the C. calixtoi species group, with the description of twenty-four new species. Zootaxa 5662 1-115.  

Clistopyga andaki Palacio & Bordera, 2025 sp. nov.

Clistopyga andina Bordera & Palacio, 2025 sp. nov.

Clistopyga angustipleura Bordera & Palacio, 2025 sp. nov.

Clistopyga arhuaca Palacio & Bordera, 2025 sp. nov.

Clistopyga aurantia Bordera & Palacio, 2025 sp. nov.

Clistopyga brevicaudata Bordera & Palacio, 2025 sp. nov.

Clistopyga femorator Bordera & Palacio, 2025 sp. nov.

Clistopyga horcoana Bordera & Palacio, 2025 sp. nov.

Clistopyga inca Bordera & Palacio, 2025 sp. nov.

Clistopyga inti Bordera & Palacio, 2025 sp. nov.

Clistopyga isabellae Bordera, 2025 sp. nov.

Clistopyga kogabae Palacio & Bordera, 2025 sp. nov.

Clistopyga laplatense Bordera & Palacio, 2025 sp. nov.

Clistopyga lobulata Palacio & Bordera, 2025 sp. nov.

Clistopyga meridae Bordera & Palacio, 2025 sp. nov.

Clistopyga mochica Bordera & Palacio, 2025 sp. nov.

Clistopyga muisca Palacio & Bordera, 2025 sp. nov.

Clistopyga nasa Palacio & Bordera, 2025 sp. nov.

Clistopyga nubiphila Palacio & Bordera, 2025 sp. nov.

Clistopyga plaumanni Bordera & Palacio, 2025 sp. nov.

Clistopyga quechua Bordera & Palacio, 2025 sp. nov.

Clistopyga subtenella Bordera & Palacio, 2025 sp. nov.

Clistopyga valida Bordera & Palacio, 2025 sp. nov.

Clistopyga wounan Palacio & Bordera, 2025 sp. nov.

Afroanomalia pseudoclistopyga comb. nov. to Clistopyga pseudoclistopyga (Varga, 2018)

Ketelaere A.D. & Hodecek J. (2025) A new species of Dreisbachia (Hymenoptera: Ichneumonidae) from the Dominican Republic. Bulletin de la Société royale belge d'Entomologie 161 70-75.

Dreisbachia delossantosi Ketelaera & Hodecek 2025 sp. nov.

Pádua D.G., Sääksjärvi I.E., Kaunisto K.M. & Monteiro R.F. (2025) Two new species of the Clistopyga henryi species-group (Hymenoptera, Ichneumonidae, Pimplinae) from South America, with a key to species of the group. ZooKeys 1260 23-36. 

Clistopyga peruandina Sääksjärvi & Pádua 2025 sp. nov.

Clistopyga teresopolitana Pádua 2025 sp. nov.

Sushama V., Ranjith A.P., Rajmohana K., Sen S. & Dinesh K.P. (2026) A new species of Clistopyga Gravenhorst (Hymenoptera: Ichneumonidae: Pimplinae) parasitizing Telamonia dimidiata (Simon), with notes on host association and biology. Journal of Natural History 60 165-175. 

Clistopyga telamoniae Ranjith and Sushama 2026 sp. nov.

Bass A., Bennett A.M.R., Spasojevic T. & Schwarzfeld M. (2026) Revision of the world species of Piogaster Perkins (Hymenoptera, Ichneumonidae, Pimplinae), ectoparasitoids of jumping spiders (Araneae, Salticidae) with a description of one new species. Journal of Hymenoptera Research 99 621-701.

Piogaster variegata Bass, Bennett & Schwarzfeld, 2026 sp. nov.

Piogaster pilosator caucasica Kasparyan, 1981 syn. nov. under Piogaster pilosator