クモヒメバチ属群 & クモフシヒメバチ チェックリスト

クモヒメバチ属群 the Polysphincta group of genera

Currently, 25 genera and 296 extant species plus 5 fossil species (Yu et al. 2016 +alpha)


Schizopyga subgroup sensu Matsumoto (2016)

  1. Genus Brachyzapus Gauld & Dubois, 2006 16 spp. (see 4 spp. nov. by Choi et al. 2019, Varga et al. 2018)

  2. Genus Dreisbachia Townes, 1962 15 spp. (see genus resurrection by Matsumoto, 2016 and 4 comb. nov. from Schizopyga by Varga & Reshchikov 2018, Khalaim et al. 2024 )

  3. Genus Iania Matsumoto, 2016 1 species + 1 subspecies (Matsumoto, 2016)

  4. Genus Inbioia Gauld & Ugalde Gόmez, 2002 1 species

  5. Genus Piogaster Perkins, 1958 7 spp. +1 subspecies

  6a. Genus Schizopyga (Schizopyga) Gravenhorst, 1829 8 spp. + 3 subspecies (see 4 comb. nov. to Dreisbachia by Varga & Reshchikov 2018, Khalaim et al. 2024 )

  6b. Genus Schizopyga (Schizopygoides) Kasparyan, 1976 1 species

  7. Genus Zabrachypus Cushman, 1920 5 spp. + 1 fossil species (see 2 spp. nov. by Mohammadi-Khoramabadi et al. 2022)


Acrodactyla subgroup sensu Matsumoto (2016)

  8. Genus Acrodactyla Haliday, 1838 29 spp. +1 subspecies (see 2 comb. nov. to Megaetaira by Matsumoto 2016 and 1 syn. nov. by Broad 2021)

  9. Genus Eruga Townes, 1960 14 spp. (see 1 sp. nov. by Sobczak et al. 2018 and 1 syn. nov. by  Khalaim et al. 2024)

10. Genus Lamnatibia Palacio & Sääksjärvi, 2007 1 species

11. Genus Longitibia He & Yu, 1999 1 species

12.  Genus Megaetaira Gauld & Dubois, 2006 3 spp. (see 2 comb. nov. from Acrodactyla by Matsumoto 2016)

13. Genus Pterinopus Townes, 1969 1 species


Polysphincta subgroup sensu Matsumoto (2016)

14. Genus Acrotaphus Townes, 1960 27 spp. (see 17 spp. nov. by Higa & Penteado-Dias 2020, Pádua et al. 2020a, Pádua & Kloss 2020)

15. Genus Aravenator Momoi, 1973 1 species

16. Genus Chablisea Gauld & Dubois, 2006 7 spp.

17. Genus Eriostethus Morley, 1914 18 spp.

18. Genus Flacopimpla Gauld, 1991 7 spp.

19. Genus Hymenoepimecis Viereck, 1912 29 spp. (see 8 spp. nov. by Pádua et al.  2020b, Pádua 2022, Kloss et al. 2022)

20. Genus Oxyrrhexis Förster, 1869 4 spp. + 1 subspecies

21. Genus Polysphincta Gravenhorst, 1829 33 spp. + 4 fossil species + 2 subspecies (see 11 spp. nov. by Kloss et al. 2018, Higa & Penteado-Dias 2020, Pádua et al.  2021, Khalaim et al. 2024 and 1 comb. nov. to Zaglyptus by Takasuka & Broad 2024)

22. Genus Reclinervellus He & Ye, 1998 4 spp.

23. Genus Sinarachna Townes, 1960 4 spp. + 1 subspecies (see 1 comb. nov. to Zatypota by Broad 2021)

24. Genus Ticapimpla Gauld, 1991 5 spp.

25. Genus Zatypota Förster, 1869 53 spp. + 8 subspecies (see 2 spp. nov. by Sobczak et al. 2019 and 1 comb. nov. from Sinarachna by Broad 2021)



祖先群(クモ卵塊寄生バチ Pseudo-parasitoids of spider egg masses)


1. Genus Afroanomalia Varga, 2018  1 sp. (Varga 2018)

2. Genus Clistopyga Gravenhorst, 1829 66 spp. +1 subspecies (see 28 spp. nov. by Bordera et al. 2016, 2019, Bordera & Palacio 2019, Manjusha et al. 2019a, Palacio et al. 2018, 2019)

3. Genus Tromatobia Förster, 1869 33 spp. + 3 subspecies

4. Genus Zaglyptus Förster, 1869 27 spp. +7 subspecies (see 1 sp. nov.  by Manjusha et al. 2019b transferred from Polysphincta by Takasuka & Broad 2024)


Bordera S., Sääksjärvi I.E., Castillo C., Palacio E. & González-Moreno A. (2016) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part I: the C. chaconi species group, with the description of eleven new species. European Journal of Taxonomy 206 1-37.

Bordera S. & Palacio E. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part IV: the C. eldae species group, with the description of three new species. Zootaxa 4564 327-346.

Bordera S., Palacio E. & Martínez J.J. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part V: the C. diazi species group, with the description of three new species. Zootaxa 4661 545-565.

Broad G.R. (2021) Taxonomic changes in Ichneumonoidea (Hymenoptera), and notes on certain type specimens. Zootaxa 4941 511–541.

Choi J.K., Kolarov J. & Lee J.W. (2019) Descriptions of two new species of Brachyzapus Gauld & Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from South Korea with key to a Palaearctic species. Zootaxa 4711 293-306.

Higa P.T. & Penteado-Dias A.M. (2019) Altitudinal effects on diversity of Pimplinae (Hymenoptera, Ichneumonidae) from Southeast Brazil and description of new species. Brazilian Journal of Biology 80 377-385.

Khalaim A.I., RuÍz-Cancino E. & Coronado-Blanco J.M. (2024) Darwin wasps of the subfamily Pimplinae (Hymenoptera: Ichneumonidae) of Mexico: Polysphincta genus-group. Zootaxa 5458 151-196. 

Kloss T.G., Pádua D.G., Almeida S.D.S., Penteado-Dias A.M., Mendes-Pereira T., Sobczak J.F., Lacerda F.G. & Gonzaga M.O. (2022) A new Darwin wasp (Hymenoptera: Ichneumonidae) and new records of behavioral manipulation of the host spider Leucauge volupis (Araneae: Tetragnathidae). Neotropical Entomology Online first

Manjusha B.M., Sudheer K. & Ghosh S.M. (2019a) A new species of Clistopyga Gravenhorst (Hymenoptera: Ichneumonidae: Pimplinae) from the Western Ghats. Records of the Zoological Survey of India 119 60-63.

Manjusha B.M., Sudheer K. & Ghosh S.M. (2019b) Polysphincta idukkiensis (Hymenoptera: Ichneumonidae: Pimplinae) a rare new species from the southern Western Ghats. Halteres 10 96-99.

Matsumoto, R. (2016) Molecular phylogeny and systematics of the Polysphincta group of genera (Hymenoptera, Ichneumonidae, Pimplinae). Systematic Entomology 41 854-864.

Mohammadi-Khoramabadi A., Talebi A., Broad G.R. & Zwakhals K. (2022) A study of the genus Zabrachypus Cushman, 1920 (Hymenoptera Ichneumonidae Pimplinae) from Iran with descriptions of two new species. Redia 105 29-36.

Pádua D.G., Sääksjärvi I.E., Monteiro R.F. & Oliveira M.L. (2020a) Review of the new world genus Acrotaphus Townes, 1960 (Hymenoptera: Ichneumonidae: Pimplinae), with descriptions of fifteen new species. Zootaxa 4719 1-62.

Pádua D.G., Sääksjärvi I.E., Monteiro R.F. & Oliveira M.L. (2020b) Seven new species of spider-attacking Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Ecuador, French Guiana, and Peru, with an identification key to the world species. ZooKeys 935 57-92.

Pádua D.G. & Kloss T.G. (2020) A new species of Acrotaphus Townes (Hymenoptera: Ichneumonidae: Pimplinae) from the Brazilian Amazon, with notes on its host spider behavioral modification. Zootaxa 4810 161-168.

Pádua D.G., Sääksjärvi I.E., Spasojevic T., Kaunisto K.M., Monteiro R.F. & Oliveira M.L. (2021) A review of the spider-attacking Polysphincta dizardi species-group (Hymenoptera, Ichneumonidae, Pimplinae), with descriptions of seven new species from South America. ZooKeys 1041 137-165.

Palacio E., Bordera S., Sääksjärvi I.E. & Díaz F. (2018) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part II: the C. isayae species group, with the description of seven new species. Zootaxa 4442 101-121.

Palacio E., Bordera S. & Díaz F. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae) Part III: the C. henryi species group, with the description of three new species. Zootaxa 4563 103-118.

Sobczak J.F., Pádua D.G., Villanueva- Bonilla G.A., De Sousa Nóbrega F.A. & Messas Y.F. (2019) Two new species of Zatypota (Hymenoptera: Ichneumonidae, Pimplinae) sharing the same host spider in Northeast Brazil. Zootaxa 4609 169-177.

Takasuka, K. & Broad, G.R. (2024) A bionomic overview of spider parasitoids and pseudo-parasitoids of the ichneumonid wasp subfamily Pimplinae. Contributions to Zoology in press.

Varga O. (2018) Afroanomalia pseudoclistopyga, a new genus and species of pimpline parasitoid wasp (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region. Zootaxa 4461 146-150.

Varga O. & Reshchikov A. (2018) Schizopyga alinae, a genus of pimpline parasitoid wasps (Hymenoptera: Ichneumonidae: Pimplinae) new to the Afrotropical region. Zootaxa 4422 291-295.

Varga O., Reshchikov A. & Giovanni F.D. (2018) First record of the genus Brachyzapus Gauld & Dubois, 2006 (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region, with descriptions of two new species. Zootaxa 4377 581-586.

Yu D.S., Achterberg K. van & Horstmann K. (2016) Taxapad 2016, World Ichneumonoidea 2015. Database on flash-drive. http://www.taxapad.com, Ottawa, Ontario, Canada.



クモヒメバチ属群 the Polysphincta group of genera


Schizopyga subgroup sensu Matsumoto (2016)

 

1. Genus Brachyzapus Gauld & Dubois, 2006 16 spp.

Gauld I.D. & Dubois J. (2006) Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae): a taxonomic revision of spider ectoparasitoids. Systematic Entomology 31 529-564.


atripedalis (Sheng, 2001)

carinatus Pham, Broad, Matsumoto & Wägele, 2012

convergens Choi & Lee, 2019

convexus Pham, Broad, Matsumoto & Wägele, 2012

duboisi Pham, Broad, Matsumoto & Wägele, 2012

fansipanensis Pham, Broad, Matsumoto & Wägele, 2012

glaber Varga & Reshchikov, 2018

greengoblin Varga & Di Giovanni, 2018

hoanglienensis Pham, Broad, Matsumoto & Wägele, 2012

nikkoensis (Uchida, 1928)

nitidus (Hao & Sheng, 2002)

nonareaeidos (Wang, 1997)

politus Pham, Broad, Matsumoto & Wägele, 2012

pyramidalis Choi & Lee, 2019 

tenuiabdominalis (Uchida, 1941)

unicarinatus (Uchida & Momoi, 1958)

Choi J.K., Kolarov J. & Lee J.W. (2019) Descriptions of two new species of Brachyzapus Gauld & Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from South Korea with key to a Palaearctic species. Zootaxa 4711 293-306.

Hao D. & Sheng M.L. (2002) A new specie of genus Zabrachypus Cushman (Hymenoptera: Ichneumonidae). Journal of Northeast Forestry University 30 95-96 (in Chinese with English summary).

Pham N.T., Broad G.R., Matsumoto R. & Wägele W.J. (2012) First record of the genus Brachyzapus Gauld and Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam, with descriptions of six new species. Journal of Natural History 46 1639-1661.

Sheng M.L. & Pan H.Y. (2001) Two new species of family Ichneumonidae (Hymenoptera). Entomologia Sinica 8 25-29.

Uchida T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido Imperial University 25 1-115.

Uchida T. (1941) Baiträge zur Systematik der Tribus Polysphinctini Japans. Insecta Matsumurana 15 112-122.

Uchida T. & Momoi S. (1958) On the species of Polysphincta Gravenhorst and Zatypota Förster from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 22-30.

Varga O., Reshchikov A. & Giovanni F.D. (2018) First record of the genus Brachyzapus Gauld & Dubois, 2006 (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region, with descriptions of two new species. Zootaxa 4377 581-586.

Wang S.F., Yao J. & Wang G.G. (1997) Hymenoptera: Ichneumonidae. In: X.K. Yang (eds.). Insects of the Three Gorge Reservoir area of Yangtze River. pp. 1617-1646 (in Chinese). Chongqing Publishing Company, Chongqing, China.



2. Genus Dreisbachia Townes, 1962 15 spp. 

Townes H. (1962) A generic name in the Polysphinctine Ichneumonids (Hymenoptera). Proceedings of the Entomological Society of Washington 64 38.


amurensis Kasparyan, 1976

anseli Fernandez-Triana, 2007

aperta Momoi, 1966

avivae Gauld, 1991

congica (Benoit, 1953)

curvicauda (Seyrig, 1935)

dotae Gauld, Ugalde Gómez & Hanson, 1998

eberhardi Gauld, Ugalde Gómez & Hanson, 1998

flavifrontalis (Uchida & Momoi, 1959)

lutea Gauld, 1984

mira (Tosquinet, 1903)

moreirai Loffredo & Penteado-Dias 2008

navaja (Townes, 1960)

punctata (Uchida & Momoi, 1959)

slossonae (Davis, 1898)

Benoit P.L.G. (1953) Sur des Polysphinctini du Congo belge (Ichneum. - Pimpl.). Revue de Zoologie et de Botanique Africaines 47 137-141.

Davis G.C. (1897) Descriptions of new species of Trigonalidæ, Stephanidæ and Ichneumonidæ. Transactions of the American Entomological Society 24 349-372 

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biología Tropical 46 (Suppl. 1) 1-189.

Kasparyan D.R. (1976) [Review of Ichneumonids of the tribe Polysphinctini and Poemeniini (Hymenoptera, Ichneumonidae) of the Far East.] Trudy Zoologicheskogo Instituta 67 68-89 (in Russian).

Momoi S. (1966) Some new Ichneumonidae (Hymenoptera) from New Guinea and adjacent areas. Pacific Insects 8 152-164.

Seyrig A. (1935) Mission scientifique de l'Omo. Tome III. Fascicule 18. Hymenoptera, II. Ichneumonidae: Cryptinae, Pimplinae, Tryphoninae et Ophioninae. Mémoires du Muséum National d'Histoire Naturelle 4 1-100.

Tosquinet D. (1903) Ichneumonides nouveaux. Mémoires de la Société entomologique de Belgique 10 1-403.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. & Momoi S. (1959) On the species of the genera Laufeia and Lycorina occurring in Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 82-87.



 3. Genus Iania Matsumoto, 2016 1 species + 1 subspecies (Matsumoto, 2016)

Matsumoto R. (2016) Molecular phylogeny and systematics of the Polysphincta group of genera (Hymenoptera, Ichneumonidae, Pimplinae). Systematic Entomology 41 854-864.


pictifrons (Thomson, 1877)

pictifrons rufigaster (Constantineanu, Ciochia, Constantineanu & Ularu, 1967)

Constantineanu M.I., Ciochia V., Constantineanu R.M., Mustata G. & Ularu G. (1967) Contributions a l'etude des Ichneumonides de la montagne de Gutii, région de Maramures. Analele ştiinţifice ale Universităţii “Al. I. Cuza” din Iaşi. Sect. II a 13 47-61.

Thomson C.G. (1877) XXVII. Bidrag till kännedom om Sveriges Pimpler. Opuscula Entomologica, Lund 8 732-777.



 4. Genus Inbioia Gauld & Ugalde Gόmez, 2002 1 species

pivai Gauld & Ugalde Gómez, 2002

Gauld I.D. & Ugalde Gόmez J.A. (2002) Appendix 1. Subfamily Pimplinae. Further supplement to volume I. Memoirs of the American Entomological Institute 66 744-746.



 5. Genus Piogaster Perkins, 1958 7 spp. +1 subspecies

Perkins J.F. (1958) A new genus and three new species of Polysphinctini from Europe (Hym., Ichneumonidae). Entomologist 91 263-267.


albina Perkins, 1958

daisetsuzana Kusigemati, 1985

lucida Constantineanu & Constantineanu, 1969

maculata Townes, 1960

pilosator (Aubert, 1958)

pilosator caucasica Kasparyan, 1981

punctulata Perkins, 1958

ussuriensis Kasparyan & Khalaim, 2007

Aubert J.F. (1958) Ichneumonide Pimpline d'un genre nouveau, espèce nouvelle, rèpandue sur le rivage mèditerranéen. Bulletin de la Société Entomologique de Mulhouse 1958 79-80.

Constantineanu M.I. & Constantineanu R.M. (1969) Ichneumonides nouveaux pour la science et pour la faune de la Roumanie. Comunicari de Zoologie, Bucaresti 8 155-165 (in Romanian with French & Russian summary).

Kasparyan D.R. (1981) [New species of the subfamily Pimplinae (Hymenoptera, Ichneumonidae), parasites of spiders in the Far East of the USSR.]. Trudy Zoologicheskogo Instituta 92 108-112 (in Russian).

Kasparyan D.R. & Khalaim A.I. (2007) [1. Subfamily Pimplinae]. In: A.S. Lelej (eds.). [Key to the insects of the Russian Far East. Vol. IV. Neuropteroidea, Mecoptera, Hymenoptera. Pt. 5]. pp. 279-333 (in Russian). Vladivostok, Russia.

Kusigemati K. (1985) A new species of the genus Piogaster Perkins from Japan (Hymenoptera, Icuneumonidae). Kontyû 53 583-586.

Perkins J.F. (1958) A new genus and three new species of Polysphinctini from Europe (Hym., Ichneumonidae). Entomologist 91 263-267.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.



 6a. Genus Schizopyga (Schizopyga) Gravenhorst, 1829 8 spp. + 3 subspecies

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.


alnae Varga & Reshchikov, 2018

circulator (Panzer, 1800)

circulator meridionator Aubert, 1963

circulator pulchra Walley, 1936

coxator Constantineanu, 1973

flavifrons Holmgren, 1856

frigida Cresson, 1870

podagrica Gravenhorst, 1829

podagrica nigrifacies Constantineanu & Ciochia, 1967

quannanica Sun & Sheng, 2011

varipes Holmgren, 1856

Aubert J.F. (1963) Les Ichneumonides du rivage Méditerranéen Français (5e série, Département du Var). Vie et Milieu 14 847-878.

Constantineanu M.I. (1973) Ichneumonidae (Hymenoptera) din Delta Dunarii (Grindul Caraorman). Buletin Muzeul Delta Dunarii, Tulcea. Peuce 3 131-163.

Constantineanu M.I. & Ciochia V. (1967) Ichneumonidae in der Burzenländer Senke und ihrer Umgebung. Muzeul Regional Brasov. Culegere de Studii si Cercetari 1 33-52.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Fernández-Triana J.L. (2007) Notes on Cuban Pimplinae (Hymenoptera: Ichneumonidae), with the description of two new species. Boletin de la Sea 40 211-214.

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Holmgren A.E. (1856) Om slägtet Schizopyga. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar 13 69-72.

Loffredo A.P.S. & Penteado-Dias A.M. (2008) First record of Schizopyga Gravenhorst (Hymenoptera, Ichneumonidae, Pimplinae) from Brazil and a description of a new species. Brazilian Journal of Biology 68 457-458.

Panzer G.W.F. (1800) Faunae Insectorum Germanicae. Heft 79 8-14.

Sun S.P. & Sheng M.L. (2011) A new species of the genus Schizopyga Gravenhorst (Hymenoptera, Ichneumonidae) with a key to species known from China. Acta Zootaxonomica Sinica 36 970-972.

Varga O. & Reshchikov A. (2018) Schizopyga alinae, a genus of pimpline parasitoid wasps (Hymenoptera: Ichneumonidae: Pimplinae) new to the Afrotropical region. Zootaxa 4422 291-295.

Walley G.S. (1936) A new Schizopyga from Canada (Hymenop.; Ichneumonidae). The Canadian Entomologist 68 20-21.



 6b. Genus Schizopyga (Schizopygoides) Kasparyan, 1976 1 species

nitida Kasparyan, 1976

Kasparyan D.R. (1976) [Review of Ichneumonids of the tribe Polysphinctini and Poemeniini (Hymenoptera, Ichneumonidae) of the Far East.] Trudy Zoologicheskogo Instituta 67 68-89 (in Russian).



 7. Genus Zabrachypus Cushman, 1920 5 spp. + 1 fossil species

Cushman R.A. (1920) The North American ichneumon-flies of the tribes Lycorini, Polysphinctini, and Theroniini. Proceedings of the United States National Museum 58 7-48.


cazorlensis Rey del Castillo, 1996

moldavicus Constantineanu & Constantineanu, 1969

primus Cushman, 1920

rubroabdominalis Mohammadi-Khoramabadi, Talebi & Broad, 2022

rugatus Mohammadi-Khoramabadi, Talebi & Broad, 2022

(tumidus Khalaim, 2008 [fossil])

Constantineanu M.I. & Constantineanu R.M. (1969) Ichneumonides nouveaux pour la science et pour la faune de la Roumanie. Comunicari de Zoologie, Bucaresti 8 155-165 (in Romanian with French & Russian summary).

Cushman R.A. (1920) The North American ichneumon-flies of the tribes Lycorini, Polysphinctini, and Theroniini. Proceedings of the United States National Museum 58 7-48.

Khalaim A.I. (2008) Fossil ichneumon wasps (Hymenoptera: Ichneumonidae) from Biamo (Russia), Oligocene. Alavesia 2 101-112.

Mohammadi-Khoramabadi A., Talebi A., Broad G.R. & Zwakhals K. (2022) A study of the genus Zabrachypus Cushman, 1920 (Hymenoptera Ichneumonidae Pimplinae) from Iran with descriptions of two new species. Redia 105 29-36.

Rey del Castillo C. (1996) Addition to the knowledge of Spanish Polysphinctini with description of Zabrachypus cazorlensis sp. n. (Hymenoptera: Ichneumonidae). Boletín de la Asociación Española de Entomología 20 141-147 (in Spanish).



Acrodactyla subgroup sensu Matsumoto (2016)


 8. Genus Acrodactyla Haliday, 1838 29 spp. +1 subspecies

Haliday A.H. (1838) Descriptions of new British insects, indicated in Mr. Curtis's guide. Annals of Natural History 2 112-121.


aequaria Momoi, 1966

atricoxa Kusigemati, 1984

carinator (Aubert, 1965)

carinator meridionator (Aubert, 1965)

cursor Gauld, 1984

degener (Haliday, 1838)

elongata Pham, Broad, Matsumoto & Böhme, 2012

gressitti Momoi, 1966

iliensis Sheng & Bian, 1996

indica Momoi, 1966

jubata Townes, 1960

lydia Kasparyan, 1979

maiphuquyi Pham, Broad, Matsumoto & Böhme, 2012

micans Gauld, 1984

mitis Kasparyan, 1976

mixta Kasparyan, 1976

ocellata Townes, 1960

orientalis (Cushman, 1933)

phuthoensis Pham, Broad, Matsumoto & Böhme, 2012

quadrisculpta (Gravenhorst, 1820)

rufithorax (Györfi, 1944)

rufotibiator Aubert, 1964

shawi Pham, Broad, Matsumoto & Böhme, 2012

similis Horstmann, 2011

syndromosa Kusigemati, 1984

taiwanensis (Cushman, 1933)

takewakii (Uchida, 1927)

tami Pham, Broad, Matsumoto & Böhme, 2012

trochanterata Kasparyan, 1976

zekhem Gauld, 1984

Aubert J.F. (1964) Ichneumonides de France et du Bassin méditerranéen appartenant à un genre nouveau et neuf espèces nouvelles. Bulletin de la Société Entomologique de Mulhouse 1964 35-40.

Aubert J.F. (1965) Ichneumonides d'Europe appartenant à dix espèces nouvelles et plusieurs genres nouveaux. Bulletin de la Société Entomologique de Mulhouse 1965 15-23.

Cushman R.A. (1933) H. Sauter's Formosa-collection: Subfamily Ichneumoninae (Pimplinae of Ashmaed). Insecta Matsumurana 8 1-50.

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

Gravenhorst J.L.C. (1820) Monographia Ichneumonum Pedemontanae Regionis. Memorie della Reale Academia dell Scienze di Torino 24 275-388.

Györfi J. (1944) Vier neue Ichneumoniden-Arten aus Ungarn. Fragmenta Faunistica Hungarica 7 42-45.

Haliday A.H. (1838) Descriptions of new British insects, indicated in Mr. Curtis's guide. Annals of Natural History 2 112-121.

Horstmann K. (2011) Revisionen von Schlupfwespen-Arten XV (Hymenoptera: Ichneumonidae). Mitteilungen der Münchner Entomologischen Gesellschaft 101 5-13.

Kasparyan D.R. (1976) [Review of Ichneumonids of the tribe Polysphinctini and Poemeniini (Hymenoptera, Ichneumonidae) of the Far East.] Trudy Zoologicheskogo Instituta 67 68-89 (in Russian).

Kasparyan D.R. (1979) [A new species of Acrodactyla Haliday (Hymenoptera, Ichneumonidae).] Trudy Vsesoyuznogo Entomologicheskogo Obshchestva 61 132-134 (in Russian).

Kusigemati K. (1984) Some Ephialtinae of South East Asia, with descriptions of eleven new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 5 126-150.

Momoi S. (1966) Some new Ichneumonidae (Hymenoptera) from New Guinea and adjacent areas. Pacific Insects 8 152-164.

Pham N.T., Broad G.R., Matsumoto R. & Böhme W. (2012) First record of the genus Acrodactyla Haliday (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam, with descriptions of six new species. Zootaxa 3207 40-53.

Sheng M.L. & Bian X. (1996) A new species of genus Acrodactyla (Hymenoptera: Ichneumonidae) from China. Entomotaxonomia 18 216-218 (in Chinese).

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. (1927) Zwei neue Schmarotzerhymenopteren der Spinnen. Insecta Matsumurana 1 171-174.

Uchida T. & Momoi S. (1958) On the species of Polysphincta Gravenhorst and Zatypota Förster from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 22-30.



 9. Genus Eruga Townes, 1960 14 spp.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.


albanoi Gauld, 1991

atrata Townes, 1960

draperi Gauld, 1991

erratica (Seyrig, 1932)

gutfreundi Gauld, 1991

haseltinei Gauld, 1991

kauli Gauld, 1991

lineata Townes, 1960

paulisiana (Benoit, 1953)

rufa Townes, 1960

straussi Gauld, 1991

telljohanni Gauld, 1991

unilabiana Pádua & Sobczak, 2018

viatrix (Seyrig, 1932)

Benoit P.L.G. (1953) Notes Ichneumonologiques Africaines IV. Revue de Zoologie et de Botanique Africaines 47 153-159.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Seyrig A. (1932) Les Ichneumonides de Madagascar. I Ichneumonidae Pimplinae. Mémoires de l'Académie Malgache. Fascicule 11 1-183.

Sobczak J.F., Pádua D.G., Costa L.F.A., Carvalho J.L.V.R., Ferreira J.P.S., Sobczak J.C.M.S.M. & Messas Y.F. (2018) The parasitoid wasp Eruga unilabiana Pádua & Sobczak, sp. nov. (Hymenoptera: Ichneumonidae) induces behavioral modification in its spider host. Entomological Science 21 59-65.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.



10. Genus Lamnatibia Palacio & Sääksjärvi, 2007 1 species

andina Palacio & Sääksjärvi, 2007

Palacio E., Sääksjärvi I.E. & Vahtera V. (2007) Lamnatibia, a new genus of the Polysphincta group of genera from Colombia (Hymenoptera: Ichneumonidae; Pimplinae). Zootaxa 1431 55-63.



11. Genus Longitibia He & Yu, 1999 1 species

sinica He & Ye, 1999

He J.H. & Ye S.F. (1999) A new genus of Polysphinctini (Hymenoptera: Ichneumonidae) from China. Entomologia Sinica 6 8-10.



12. Genus Megaetaira Gauld & Dubois, 2006 3 spp.

Gauld I.D. & Dubois J. (2006) Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae): a taxonomic revision of spider ectoparasitoids. Systematic Entomology 31 529-564.


inoperta (Kusigemati, 1984)

madida (Haliday, 1838)

varicarinata (Uchida & Momoi, 1958)

Haliday A.H. (1838) Descriptions of new British insects, indicated in Mr. Curtis's guide. Annals of Natural History 2 112-121.

Kusigemati K. (1984) Some Ephialtinae of South East Asia, with descriptions of eleven new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 5 126-150.

Uchida T. & Momoi S. (1958) On the species of Polysphincta Gravenhorst and Zatypota Förster from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 22-30.



13. Genus Pterinopus Townes, 1969 1 species

scambus Townes, 1969

Townes H. (1969) The genera of Ichneumonidae, part 1. Memoirs of the American Entomological Institute 11 1-300.



Polysphincta subgroup sensu Matsumoto (2016)


14. Genus Acrotaphus Townes, 1960 27 spp.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.


amajari Pádua, 2020

amazonicus Pádua & Sääksjärvi, 2020

bodoquenaensis Pádua, 2020

chedelae Gauld, 1991

cuzconus Pádua & Sääksjärvi, 2020

dolichopus Pádua, 2020

fasciatus (Brullé, 1846)

fascipennis (Cresson, 1865)

ferruginosus (Cresson, 1865)

franklini Gauld, 1991

fuscipennis (Cresson, 1865)

homeofranklini Pádua, 2020

jackiechani Pádua & Sääksjärvi, 2020

japi Higa & Penteado-Dias, 2020

kourou Pádua & Sääksjärvi, 2020

latifasciatus (Cameron, 1911)

mexicanus (Cameron, 1886)

micrus Pádua, 2020

monotaenius Pádua, 2020

nambilloensis Pádua, 2020

pseudoamazonicus Pádua & Sääksjärvi, 2020

pseudomexicanus Pádua, 2020

tibialis (Cameron, 1886)

venezuelanus Pádua, 2020

wagnerianae Pádua, 2020

wiltii (Cresson, 1870)

zampieronae Pádua, 2020

Brullé M.A. (1846) Tome Quatrième. Des Hyménoptères. Les Ichneumonides. In: A. Lepeletier de Saint Fargeau (eds.). Histoire Naturelle Des Insectes. Hyménoptères. pp. 56-521. Paris, France.

Cameron P. (1911) On the Hymenoptera of the Georgetown Museum, Britich Guiana. Part I. Timehri 1 152-186.

Cameron P. (1886) Hymenoptera. In: F.D. Godman & O. Salvin (eds.). Biologia Centrali-Americana, or, Contributions to the knowledge of the fauna and flora of Mexico and Central America. Zoology.). pp. 241-328. 

Cresson E.T. (1865) On the Hymenoptera of Cuba. Proceedings of the Entomological Society of Philadelphia 4 1-200.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Higa P.T. & Penteado-Dias A.M. (2020) Altitudinal effects on diversity of Pimplinae (Hymenoptera, Ichneumonidae) from Southeast Brazil and description of new species. Brazilian Journal of Biology 80 377-385.

Pádua D.G., Sääksjärvi I.E., Monteiro R.F. & Oliveira M.L. (2020) Review of the new world genus Acrotaphus Townes, 1960 (Hymenoptera: Ichneumonidae: Pimplinae), with descriptions of fifteen new species. Zootaxa 4719 1-62.

Pádua D.G. & Kloss T.G. (2020) A new species of Acrotaphus Townes (Hymenoptera: Ichneumonidae: Pimplinae) from the Brazilian Amazon, with notes on its host spider behavioral modification. Zootaxa 4810 161-168.



15. Genus Aravenator Momoi, 1973 1 species

kamijoi Momoi, 1973

Momoi S. (1973) Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mongolei. 331. Einige mongolischen Arten der Unterfamilien Ephialtinae und Xoridinae (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica 26 (Suppl.) 219-239.



16. Genus Chablisea Gauld & Dubois, 2006 7 spp.

Gauld I.D. & Dubois J. (2006) Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae): a taxonomic revision of spider ectoparasitoids. Systematic Entomology 31 529-564.


albifacialis (Kusigemati, 1984)

concava Liu, He & Chen, 2009

condimenta Pham, Broad, Matsumoto & Wägele, 2011

imbiba Gauld & Dubois,2006

khuatdanglongi Pham, Broad, Matsumoto & Wägele, 2011

shaanxiensis (Liu, He & Chen, 2009)

varicolor Liu, He & Chen, 2009

Gauld I.D. & Dubois J. (2006) Phylogeny of the Polysphincta group of genera (Hymenoptera: Ichneumonidae; Pimplinae): a taxonomic revision of spider ectoparasitoids. Systematic Entomology 31 529-564.

Kusigemati K. (1984) Some Ephialtinae of South East Asia, with descriptions of eleven new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 5 126-150.

Liu J.X., He J.H. & Chen X.X. (2009) Two new species of genus Chablisea Gauld et Dubois, 2006 (Hymenoptera: Ichneumonidae: Pimplinae) from China. Biologia 64 1165-1169.

Pham N.T., Broad G., Matsumoto R. & Wägele W. (2011) Two new species of the genus Chablisea Gauld et Dubois (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam. Biologia 66 1134-1139.



17. Genus Eriostethus Morley, 1914 18 spp.

Morley C. (1914) A Revision of the Ichneumonidae Based on the Collection in the British Museum (Natural History) Part III. Tribes Pimplides and Bassides. British Museum, London, U.K. 148 pp.


babai (Kusigemati, 1987)

bifasciatus (Baltazar, 1964)

carinatus Baltazar, 1964

chinensis (He, 1985)

fasciatus (Momoi, 1966)

infumatus (Momoi, 1966)

krombeini (Baltazar, 1964)

maximus Gauld, 1984

mesorufus Momoi, 1966

minimus Gauld, 1984

nigropictus Baltazar, 1964

nodatus (Baltazar, 1964) 

perkinsi (Baltazar, 1964)

pulcherrimus Morley, 1914

rufus (Uchida, 1932)

samoanus (Fullaway, 1938)

sinuatus (Momoi, 1966)

trifasciatus (Baltazar, 1964)

Baltazar C.R. (1964) Eriostethus Morley and a new polysphinctine genus (Ichneumonidae, Hymenoptera). Pacific Insects 6 388-401.

Fullaway D.T. (1938) New species from the Bishop Museum collection of Samoan parasitic Hymenoptera. Proceedings of the Hawaiian Entomological Society 10 49-52.

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

He J.H. (1985) [A new species of Millironia Baltazar (Hymenoptera: Ichneumonidae).] Wuyi Science Journal 5 63-65 (in Chinese with English summary).

Kusigemati K. (1987) Some Ephialtinae, Banchinae, Porizontinae, Mesochorinae, Metopiinae and Acaenitinae of Formosa (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 8 21-33.

Momoi S. (1966) Some new Ichneumonidae (Hymenoptera) from New Guinea and adjacent areas. Pacific Insects 8 152-164.

Morley C. (1914) A Revision of the Ichneumonidae Based on the Collection in the British Museum (Natural History) Part III. Tribes Pimplides and Bassides. British Museum, London, U.K. 148 pp.



18. Genus Flacopimpla Gauld, 1991 7 spp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.


barathrica Fritzén, 2014

kasparyani Khalaim & Ruíz-Cancino, 2011

gerardoi (Gauld, Ugalde Gómez & Hanson, 1998)

nigriceps (Walsh, 1873)

parva (Cresson, 1870)

sulina Graf & Kumagai, 1998

varelae Gauld, 1991

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Fritzén N.R. (2014) Two new species of the Polysphincta genus-group (Hymenoptera: Ichneumonidae: Pimplinae) reared from their spider (Araneae) hosts in Europe. Zootaxa 3894 117-130.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biologia Tropical 46 (Suppl. 1) 1-189.

Graf V. & Kumagai A.F. (1997) The occurrence of Flacopimpla Gauld in Brazil (Hymenoptera, Ichneumonidae, Pimplinae). Revista Brasileira de Zoologia 14 737-777.

Khalaim A.I. & Ruíz-Cancino E. (2011) North and Central American species of the genus Flacopimpla Gauld (Hymenoptera: Ichneumonidae: Pimplinae), with description of a new species from Mexico. Proceedings of the Zoological Institute RAS 315 70-74.

Walsh B.D. (1873) Descriptions of North American Hymenoptera. Transactions of the Academy of Sciences of St. Louis 3 65-166.



19. Genus Hymenoepimecis Viereck, 1912 29 spp.

Viereck H.L. (1912) Descriptions of five new genera and twenty six new species of Ichneumon-flies. Proceedings of the United States National Museum 42 139-153.


amazonensis Pádua & Oliveira, 2015

andina Pádua & Sääksjärvi, 2020

argyraphaga Gauld, 2000

atriceps (Cresson, 1865)

atlantica Pádua, 2022

bicolor (Brullé, 1846)

cameroni Townes, 1966

castilloi Pádua & Sääksjärvi, 2020

dolichocarinata Pádua & Sääksjärvi, 2020

duckensis Pádua & Onody, 2015

ecuatoriana Pádua & Sääksjärvi, 2020

heidyae Gauld, 1991

heteropus (Kriechbaumer, 1890)

japi Sobczak, Loffredo, Penteado-Dias & Gonzaga, 2009

jordanensis Loffredo & Penteado-Dias 2009

kleini Pádua & Sobczak, 2015

longilobus Pádua & Sääksjärvi, 2020

manauara Pádua & Oliveira, 2015

neotropica (Brues & Richardson, 1913)

pinheirensis Penteado-Dias & Pádua 2022

pucallpina Pádua & Sääksjärvi, 2020

rafaelmartinezi Pádua & Sääksjärvi, 2020

ribeiroi Pádua & Sobczak, 2015

robertsae Gauld, 1991

silvanae Loffredo & Penteado-Dias 2009

sooretama Sobczak, Loffredo, Penteado-Dias & Gonzaga, 2009

tedfordi Gauld, 1991

uberensis Pádua & Onody, 2015

veranii Loffredo & Penteado-Dias 2009

Brues C.T. & Richardson C.H. (1913) Descriptions of new parasitic Hymenoptera from British Guiana. Bulletin of the American Museum of Natural History 32 485-503.

Brullé M.A. (1846) Tome Quatrième. Des Hyménoptères. Les Ichneumonides. In: A. Lepeletier de Saint Fargeau (eds.). Histoire Naturelle Des Insectes. Hyménoptères. pp. 56-521. Paris, France.

Cresson E.T. (1865) On the Hymenoptera of Cuba. Proceedings of the Entomological Society of Philadelphia 4 1-200.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D. (2000) The re-definition of pimpline genus Hymenoepimecis (Hymenoptera: Ichneumonidae) with a description of a plesiomorphic new Costa Rican species. Journal of Hymenoptera Research 9 213-219.

Kloss T.G., Pádua D.G., Almeida S.D.S., Penteado-Dias A.M., Mendes-Pereira T., Sobczak J.F., Lacerda F.G. & Gonzaga M.O. (2022) A new Darwin wasp (Hymenoptera: Ichneumonidae) and new records of behavioral manipulation of the host spider Leucauge volupis (Araneae: Tetragnathidae). Neotropical Entomology Online first

Kriechbaumer J. (1890) Ichneumoniden-Studien. Neue Ichneumoniden des Wiener Museums. II. Annalen des Naturhistorischen Hofmuseums Wien 5 479-491.

Loffredo A.P.S. & Penteado-Dias A.M. (2009) New species of Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Brazilian Atlantic forest. Revista Brasileira de Entomologia 53 11-14.

Pádua D.G. (2022) First record of the Darwin wasp Hymenoepimecis Viereck, 1912 (Hymenoptera: Ichneumonidae: Pimplinae) from Argentina and Bolivia, with description of a new species. Zootaxa 5169 49-60.

Pádua D.G., Oliveira M.L., Onody H.C., Sobczak J.F., Sääksjärvi I.E. & Gómez I.C. (2015) The Brazilian Amazonian species of Hymenoepimecis Viereck, 1912 (Hymenoptera: Ichneumonidae: Pimplinae). Zootaxa 4058 175-194.

Pádua D.G., Sääksjärvi I.E., Monteiro R.F. & Oliveira M.L. (2020) Seven new species of spider-attacking Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Ecuador, French Guiana, and Peru, with an identification key to the world species. ZooKeys 935 57-92.

Sobczak J.F., Loffredo A.P.S., Penteado-Dias A.M. & Gonzaga M.O. (2009) Two new species of Hymenoepimecis (Hymenoptera: Ichneumonidae: Pimplinae) with notes on their spider hosts and behaviour manipulation. Journal of Natural History 43 2691-2699.

Townes H. & Townes M. (1966) A catalogue and reclassification of the Neotropic Ichneumonidae. Memoirs of the American Entomological Institute 8 1-367.



20. Genus Oxyrrhexis Förster, 1869 4 spp. +1 subspecies

Förster A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 25 135-221.


carbonator (Gravenhorst, 1807)

carbonator texana (Cresson, 1870)

eurus Kasparyan, 1977

rugosa Liu, He & Chen, 2009

zephyrus Fritzén, 2014

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Fritzén N.R. & Fjellberg A. (2014) Natural history of Oxyrrhexis zephyrus sp.n. (Hymenoptera, Ichneumonidae), a parasitoid of Enoplognatha serratosignata (Araneae, Theridiidae), with notes on taxonomy and other host species of Oxyrrhexis. Arthropoda Selecta 23 135-144.

Gravenhorst J.L.C. (1807) Vergleichende Übersicht des Linneischen und einiger neurern zoologischen Systeme, nebst dem eingeschalteten Verzeichnisse der zoologischen Sammlung des Verfasser und den Beschreibungen neuer Thierarten, die in derselben vorhanden sind. Göttingen. 476 pp.

Kasparyan D.R. (1977) [Ichneumonids of the subfamilies Pimplinae and Tryphoninae (Hymenoptera, Ichneumonidae) new for Mogolia and Transbaikalia.] Nasekomye Mongolii 5 456-469 (in Russian).

Liu J.X., He J.H. & Chen X.X. (2009) Revision of genus Oxyrrhexis Foerster, 1869 (Hymenoptera: Ichneumonidae: Pimplinae) from China. Annales Zoologici 59 171-177.



21. Genus Polysphincta Gravenhorst, 1829 33 spp. + 4 fossil species + 2 subspecies

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.


albipes Cresson, 1880

areolaris Ratzeburg, 1844

(atavina Cockerell, 1921 [fossil])

bonita Pádua & Sääksjärvi, 2021

boops Tschek, 1869

burgessii Cresson, 1870

cosnipata Pádua & Sääksjärvi, 2021

dizardi Gauld, 1991

gutfreundi Gauld, 1991

inca Pádua, Sääksjärvi & Spasojevic, 2021

(P.? inundata Brues, 1910 [fossil], see Spasojevic et al. (2022))

janzeni Gauld, 1991

jundiai Higa & Penteado-Dias, 2019

koebelei Howard, 1892

latistriata Ratzeburg, 1844

limata Cresson, 1870

longa Kasparyan, 1976

macroepomia Pádua & Sääksjärvi, 2021 

mascoi Gauld, 1991

naranjae Gauld, Ugalde Gómez & Hanson, 1998

organensis Pádua & Sääksjärvi, 2021

pichincha Pádua, Sääksjärvi & Spasojevic, 2021

punctigaster Varga & Reshchikov, 2015

purcelli Gauld, 1991

rufipes Gravenhorst, 1829

(saxea (Scudder, 1878) [fossil])

shabui Gauld, 1991

sinearanea Pádua, 2018

sirena Khalaim, 2024

soaresi Higa & Penteado-Dias, 2020

(statzi (Meunier, 1923) [fossil])

teresa Pádua & Sääksjärvi, 2021 

thoracica (Cresson, 1874)

townesorum De Santis & Millan, 1968

tuberosa Gravenhorst, 1829

tuberosa bruneti Provancher, 1873

tuberosa meridionator Aubert, 1963

vexator Fitton, Shaw & Gauld, 1988

xena Khalaim, 2024

Aubert J.F. (1963) Les Ichneumonides du rivage Méditerranéen Français (5e série, Département du Var). Vie et Milieu 14 847-878.

Brues C.T. (1910) The parasitic Hymenoptera of the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology at Harvard University 54 1-125.

Cockerell T.D.A. (1921) Fossil Arthropods in the British Museum. V. Oligocene Hymenoptera from the Isle of Wight. Annals and Magazine of Natural History 7 1-25.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Cresson E.T. (1874) Descriptions of Mexican Ichneumonidae. Proceedings of the Academy of Natural Sciences of Philadelphia 1873 374-413.

Cresson E.T. (1880) Report of the commissioner of agriculture. Report of the Entomologist of the United States Department of Agriculture 1879 208, 235, 238.

De Santis L. & Millan E. (1968) Adicones a la fauna Argentina de Icneumonidos (Hymenoptera). Notas de la Comision de Investigacion Cientificas de la Provincia de Buenos Aires 5 1-7.

Fitton M.G., Shaw M.R. & Gauld I.D. (1988) Pimpline ichneumon-flies Hymenoptera, Ichneumonidae (Pimplinae). Handbook for the Identification of British Insects, Royal Entomological Sciety of London, London, U.K. 110 pp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Higa P.T. & Penteado-Dias A.M. (2020) Altitudinal effects on diversity of Pimplinae (Hymenoptera, Ichneumonidae) from Southeast Brazil and description of new species. Brazilian Journal of Biology 80 377-385.

Howard L.O. (1892) The hymenopterous parasites of spiders. Proceedings of the Entomological Society of Washington 2 290-302.

Khalaim A.I., RuÍz-Cancino E. & Coronado-Blanco J.M. (2024) Darwin wasps of the subfamily Pimplinae (Hymenoptera: Ichneumonidae) of Mexico: Polysphincta genus-group. Zootaxa 5458 151-196. 

Kloss T.G., Pádua D.G., Lacerda F.G., Oliveira L.S., Cossolin J.F.S., Serrão J.E. & Gonzaga M.O. (2018) Suppression of orb-web building behavior of the spider Metazygia laticeps (O. Pickard-Cambridge, 1889) (Araneae: Araneidae) by a new parasitoid wasp. Zoologischer Anzeiger 276 100-106.

Meunier F. (1923) Sur quelques insectes de l'Aquitanien de Rott (Sept-Monts, Rhénanie). Miscellanea Entomologica, Revue Entomologique Internationale 26 82-88.

Pádua D.G., Sääksjärvi I.E., Spasojevic T., Kaunisto K.M., Monteiro R.F. & Oliveira M.L. (2021) A review of the spider-attacking Polysphincta dizardi species-group (Hymenoptera, Ichneumonidae, Pimplinae), with descriptions of seven new species from South America. ZooKeys 1041 137-165.

Provancher L. (1873) Les Ichneumonides de Québec avec description de plusieurs especès nouvelles. Le Naturaliste Canadien 5 470-477.

Ratzeburg J.T.C. (1844) Die Ichneumonen der Forstinsecten in forstlicher und entomologischer Beziehung. Berlin, Germany. 224 pp.

Scudder S.H. (1878) The insects of the tertiary beds at Quesnel. Report of Progress. Geological Survey Canada 1876-1877 266-280.

Spasojevic T., Broad G.R. & Klopfstein S. (2022) Revision of 18 ichneumonid fossil species (Hymenoptera, Ichneumonidae) highlights the need for open nomenclature in palaeontology. Fossil Record 25 187-212.

Tschek C. (1869) Beiträge zur Kenntniss der österreichischen Pimplarien. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 18 269-280.

Varga O. & Reshchikov A. (2015) New records of the genus Polysphincta Gravenhorst, 1829 (Hymenoptera: Ichneumonidae: Pimplinae) from the Oriental region. Zootaxa 3955 435-443.



22. Genus Reclinervellus He & Ye, 1998 4 spp.

He J.H. & Ye S.F. (1998) Reclinervellus, a new genus of Polysphinctini (Hymenoptera: Ichneumonidae: Pimplinae) from China. Entomotaxonomia 20 153-156 (in Chinese with English summary).


dorsiconcavus He & Ye, 1998

masumotoi Matsumoto & Konishi, 2007

nielseni (Roman, 1923)

tuberculatus (Uchida, 1932)

He J.H. & Ye S.F. (1998) Reclinervellus, a new genus of Polysphinctini (Hymenoptera: Ichneumonidae: Pimplinae) from China. Entomotaxonomia 20 153-156 (in Chinese with English summary).

Matsumoto R. & Konishi K. (2007) Life histories of two ichneumonid parasitoids of Cyclosa octotuberculata (Araneae): Reclinervellus tuberculatus (Uchida) and its new sympatric congener (Hymenoptera: Ichneumonidae: Pimplinae). Entomological Science 10 267-278.

Roman A. (1923) Nya Polysphincta-former ur de Nielsen'ska kläckningarne. Entomologiske Meddelelser 14 206-210.

Uchida T. (1932) Beiträge zur Kenntnis der japanischen Ichneumoniden. Insecta Matsumurana 6 145-168.



23. Genus Sinarachna Townes, 1960 4 spp. + 1 subspecies

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.


ceylonica (Ashmead, 1896)

maculata He & Ye, 2006

nigricornis (Holmgren, 1860)

pallipes (Holmgren, 1860)

pallipes strigis (Howard, 1892)

He J.H., Chen X.X., Ma Y. & Ye S.F. (2006) Ichneumonidae. In: Z. Li & D. Jin (eds.). Insects from Fanjingshan landscape (Insect Fauna from National Nature Reserve in Guizhow Province VI). pp. 571-575. Guizhow Science and Technology Publishing House, Guiyang, Guizhou, China.

Holmgren A.E. (1860) Försök till uppställning och beskrifning af Sveriges Ichneumonider. Tredje Serien. Fam. Pimplariae. (Monographia Pimplariarum Sueciae). Vetenskapsakademiens Handlingar, (B) 3 1-76.

Howard L.O. (1892) The hymenopterous parasites of spiders. Proceedings of the Entomological Society of Washington 2 290-302.

Howard L.O. & Ashmead W.H. (1896) On some reared parasitic hymenopterous insects from Ceylon. Proceedings of the United States National Museum 18 633-648.



24. Genus Ticapimpla Gauld, 1991 5 spp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.


amazonica Palacio, Broad, Sääksjärvi & Veijalainen, 2010

carinata Palacio, Broad, Sääksjärvi & Veijalainen, 2010

matamatae Palacio, Broad, Sääksjärvi & Veijalainen, 2010

soinii Palacio, Broad, Sääksjärvi & Veijalainen, 2010

vilmae Gauld, 1991

Palacio E., Broad G.R., Sääksjärvi I.E. & Veijalainen A. (2010) Western Amazonian Ticapimpla (Hymenoptera: Ichneumonidae: Pimplinae): four new species from Colombia, Ecuador, and Peru, with a key to species of the genus. Psyche: A Journal of Entomology 2010 Article ID 161595.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.



25. Genus Zatypota Förster, 1869 53 spp. + 8 subspecies

Förster A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 25 135-221.


albicoxa (Walker, 1874)

albicoxa meridicolor Aubert, 1989

alborhombarta (Davis, 1895)

anomala (Holmgren, 1860)

arizonica Townes, 1960

baezae Pádua & Sobczak, 2019

baragi Matsumoto, 2010

bayamensis Fernández-Triana, 2007

bingili Gauld, 1984

bohemani (Holmgren, 1860)

brachycera Matsumoto, 2010

capicola Benoit, 1959

celer Gauld, 1984

chryssophaga Matsumoto, 2010

cingulata Townes, 1960

crassipes Townes, 1960

dandiensis Gauld, 1984

dendrobia Matsumoto, 2010

dichroa (Marshall, 1892)

discolor (Holmgren, 1860)

discolor helveticator Aubert, 1989

discolor nigrithorax Constantineanu, Ciochia, Constantineanu, Mustata & Ularu, 1967

discolor obscura (Roman, 1923)

elegans Matsumoto, 2010

exilis Townes, 1960

favosa Townes, 1960

flamma Fritzén, 2014

fonsecai Gauld, 1991

gracilipes Uchida & Momoi, 1958

inexpectata (Seyrig, 1932)

kauros Gauld, 1984

kerstinae Fritzén, 2010

lagiralda Gauld Menjivar, Monro & Gonzalez, 2002

luteipes Townes, 1960

maculata Matsumoto & Takasuka, 2010

medranoi Gauld, 1991

mongolica Sedivy, 1971

morsei Gauld, 1991

mulunguensis Pádua & Sobczak, 2019

pallipes Schmiedeknecht, 1888

patellata Townes, 1960

percontatoria (Müller, 1776)

percontatoria celticator Aubert, 1989

percontatoria meridianator Aubert, 1989

petronae Gauld, 1991

phraxos Gauld, 1984

picticollis (Thomson, 1888)

picticollis gallicator Aubert, 1989

picticollis meridiator Aubert, 1989

prima Benoit, 1953

rennefer Gauld, 1984

riverai Gauld, 1991

solanoi Gauld, 1991

stellata Gauld, 1984

sulcata Matsumoto, 2010

takayu Matsumoto & Takasuka, 2010

talamancae Gauld, Ugalde Gómez & Hanson, 1998

tropica (Morley, 1912)

velata Gauld, 1984

walleyi Townes, 1960

yambar Matsumoto, 2010

Aubert J.F. (1989) Ichneumonides non pétiolées inédites et quatrième suppl. aux Scolobatinae (Ctenopelmatinae): les Homaspis Foerst. Bulletin de la Société Entomologique de Mulhouse 1989 1-11.

Benoit P.L.G. (1953) Notes Ichneumonologiques Africaines IV. Revue de Zoologie et de Botanique Africaines 47 153-159.

Benoit P.L.G. (1959) Hymenoptera: Ichneumonidae (Banchinae, Bassinae, Pimplinae et Tryphoninae). South African Animal Life 6 441-482.

Constantineanu M.I., Ciochia V., Constantineanu R.M., Mustata G. & Ularu G. (1967) Contributions a l'etude des Ichneumonides de la montagne de Gutii, région de Maramures. Analele ştiinţifice ale Universităţii “Al. I. Cuza” din Iaşi. Sect. II a 13 47-61.

Davis G.C. (1895) Two new species of Clistopyga. Entomological News 6 198-199.

Fernández-Triana J.L. (2007) Notes on Cuban Pimplinae (Hymenoptera: Ichneumonidae), with the description of two new species. Boletin de la Sea 40 211-214.

Fritzén N.R. (2010) Natural history and description of Zatypota kerstinae sp.nov (Hymenoptera: Ichneumonidae) reared from Theridion palmgreni Marusik et Tsellarius (Araneae: Theridiidae) in Finland. Zootaxa 2487 52-60.

Fritzén N.R. (2014) Two new species of the Polysphincta genus-group (Hymenoptera: Ichneumonidae: Pimplinae) reared from their spider (Araneae) hosts in Europe. Zootaxa 3894 117-130.

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biologia Tropical 46 (Suppl. 1) 1-189.

Gauld I.D., Menjivar R.U.H., Monro A.K. & González M.O. (2002) Guía para la identificación de los Pimplinae de cafetales bajo sombra de El Salvador (Hymenoptera: Ichneumonidae). Natural History Museum, London, U.K. 76 pp.

Holmgren A.E. (1860) Försök till uppställning och beskrifning af Sveriges Ichneumonider. Tredje Serien. Fam. Pimplariae. (Monographia Pimplariarum Sueciae). Vetenskapsakademiens Handlingar, (B) 3 1-76.

Marshall T.A. (1892) Énumerations de quelques Hyménoptères du Venezuela. Annales de la Société Entomologique de France 61 60-76.

Matsumoto R. & Takasuka K. (2010) A revision of the genus Zatypota Förster of Japan, with descriptions of nine new species and notes on their hosts (Hymenoptera: Ichneumonidae: Pimplinae). Zootaxa 2522 1-43.

Morley C. (1912) No. XII. Hymenoptera, Ichneumonidae. In: J.S. Gardiner (eds.). Transactions of the Linnean Society of London 15. pp. 169-179. Linnean Society of London, London, U.K.

Müller O.F. (1776) Zoologiae Danicae prodromus, seu anumalium Daniae et Norvegiae indigenarum characteres, nomina et synomyma imprimis popularium. Hafniae, 282 pp.

Roman A. (1923) Nya Polysphincta-former ur de Nielsen'ska kläckningarne. Entomologiske Meddelelser 14 206-210.

Schmiedeknecht O. (1888) Die europäischen Gattungen der Schlupfwespen Familie Pimplariae. Zoologische Jahrbücher Abteilung für Systematik 3 419-444.

Sedivy J. (1971) Ergebnisse der mongolisch-tschechoslowakischen entomologisch-botanischen Expeditionen in der Mongolei: 24. Hymenoptera, Ichneumonidae. Acta Faunistica Entomologica Musei Nationalis Pragae 14 73-91.

Seyrig A. (1932) Les Ichneumonides de Madagascar. I Ichneumonidae Pimplinae. Mémoires de l'Académie Malgache. Fascicule 11 1-183.

Sobczak J.F., Pádua D.G., Villanueva- Bonilla G.A., De Sousa Nóbrega F.A. & Messas Y.F. (2019) Two new species of Zatypota (Hymenoptera: Ichneumonidae, Pimplinae) sharing the same host spider in Northeast Brazil. Zootaxa 4609 169-177.

Thomson C.G. (1888) XXXVII. Bidrag till Sveriges insectfauna. Opuscula Entomologica. Lund. XII 1202-1265.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. & Momoi S. (1958) On the species of Polysphincta Gravenhorst and Zatypota Förster from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 22 22-30.

Walker F. (1874) Descriptions of some Japanese Hymenoptera. Cistula Entomologica 1 301-310.



祖先群(クモ卵塊寄生バチ Pseudo-parasitoids of spider egg masses)


 1. Genus Afroanomalia Varga, 2018  1 sp. (Varga 2018)

pseudoclistopyga Varga, 2018

Varga O. (2018) Afroanomalia pseudoclistopyga, a new genus and species of pimpline parasitoid wasp (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region. Zootaxa 4461 146-150.



2. Genus Clistopyga Gravenhorst, 1829 66 spp. +1 subspecies

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.


aenigmatica Bordera & González-Moreno 2014

africana Benoit, 1956

albovittata Bordera & Palacio, 2019

alutaria Townes, 1960

amazonica Bordera & Sääksjärvi, 2016

arctica Kusigemati, 1985

arizonica Bordera & González-Moreno 2014

atrata Cushman, 1922

californica Khalaim & Hernández, 2008

calixtoi Gauld, 1991

canadensis Provancher, 1880

caramba Castillo & Sääksjärvi, 2015

carinata Bordera & Palacio, 2019

carvajali Gauld, Ugalde Gómez & Hanson, 1998

catarina Bordera & Palacio, 2019

chaconi Gauld, 1991

cinnamoptera Bordera & Sääksjärvi, 2016

crassicaudata Sääksjärvi, Bordera & Palacio, 2018

cuscoensis Bordera, Castillo & Sääksjärvi, 2016

declinata Palacio & Bordera, 2019

diazi Porter, 1979

eldae Gauld, 1991

emphera Kusigemati, 1985

fernandezi Gauld, 1991

ghatensis Manjusha, Sudheer & Ghosh, 2019

hayesiana Bordera & Sääksjärvi, 2016

henryi Gauld, 1991

incitator (Fabricius, 1793)

incitator mongolica Kasparyan, 2007

isayae Díaz, Bordera & Palacio, 2018

jakobii Graf, 1985

juliana Bordera & González-Moreno 2014

kalima Palacio, 2018

laevis Kasparyan, 1981

lapacensis Bordera & Palacio, 2019

latifrontalis (Uchida, 1941)

linearis (Wollaston, 1858)

longifemoralis Varga & Reshchikov, 2015

lopezrichinii (Blanchard, 1941)

maculifrons Cushman, 1922

manni Cushman, 1922

marcapatensis Bordera & Palacio, 2019

maya González-Moreno & Bordera 2014

melanoptera Castillo, Sääksjärvi & Bordera, 2016

misionensis Bordera & Sääksjärvi, 2016

mocaguae Palacio & Bordera, 2016

moraviae Gauld, Ugalde Gómez & Hanson, 1998

nigrifrons Cushman, 1922

nigriventris Palacio, 2018

oaxacana González-Moreno & Bordera 2014

orellanae Bordera & Sääksjärvi, 2016

panchei Palacio, 2018

plana Morley, 1914

polita Bordera & Palacio, 2019

porteri Bordera & Sääksjärvi, 2016

recurva (Say, 1835)

rondoniae Bordera & Sääksjärvi, 2016

rufator Holmgren, 1856

rugulosa Kusigemati, 1985

serricauda Khalaim & Hernández, 2008

speculata Bordera & Palacio, 2019

splendida Palacio & Bordera, 2018

stanfordi Gauld, 1991

sziladyi Kiss, 1959

taironae Palacio, Bordera & Díaz, 2018

teresitae Díaz, Palacio & Bordera, 2019

yabuquensis Bordera & Sääksjärvi, 2016

Bajáry N.E. (1959) Revision der Ichneumoniden-Typen von Kiss und Szépligeti II (Hymenoptera). Annales Historico-Naturales Musei Nationalis Hungarici 51 415-418.

Benoit P.L.G. (1956) Ichneumonidae nouveaux ou interessants de l'Afrique du Sud. Annals of the South African Museum 43 123-135.

Bordera S., González-Moreno A., Khalaim A.I. & Sääksjärvi I.E. (2014) Revision of North American species of Clistopyga (Hymenoptera: Ichneumonidae: Pimplinae). The Canadian Entomologist 146 355-414.

Bordera S., Sääksjärvi I.E., Castillo C., Palacio E. & González-Moreno A. (2016) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part I: the C. chaconi species group, with the description of eleven new species. European Journal of Taxonomy 206 1-37.

Bordera S. & Palacio E. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part IV: the C. eldae species group, with the description of three new species. Zootaxa 4564 327-346.

Bordera S., Palacio E. & Martínez J.J. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part V: the C. diazi species group, with the description of three new species. Zootaxa 4661 545-565.

Cushman R.A. (1922) North American Ichneumon-flies of the genera Clistopyga and Schizopyga. Proceedings of the United States National Museum 60 1-14.

Fabricius J.C. (1793) Entomologia systematica emendata et aucta. Tom. II. Hafniae, 519 pp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biologia Tropical 46 (Suppl. 1) 1-189.

Graf V. (1985) Ichneumofauna do sudeste e sul do Brasil. VI. Nova espécie de Clistopyga (Ephialtinae, Hymenoptera). Revista Brasileira de Entomologia 29 349-350.

Holmgren A.E. (1856) Entomologiska anteckningar under en resa i södra Sverige ar 1854. Kongliga Svenska Vetenskapsakademiens Handlingar 75 1-104.

Kasparyan D.R. & Khalaim A.I. (2007) [1. Subfamily Pimplinae]. In: A.S. Lelej (eds.). [Key to the insects of the Russian Far East. Vol. IV. Neuropteroidea, Mecoptera, Hymenoptera. Pt. 5]. pp. 279-333 (in Russian). Vladivostok, Russia.

Khalaim A.I. & Hernández S.A. (2008) Review of the genus Clistopyga Grav., 1829 of Mexico (Hymenoptera: Ichneumonidae: Pimplinae). Russian Entomological Journal 17 311-315.

Kusigemati K. (1985) Three new species of Clistopyga from Japan (Hymenoptera: Ichneumonidae). Memoirs of the Faculty of Agriculture, Kagoshima University 21 189-194.

Manjusha B.M., Sudheer K. & Ghosh S.M. (2019) A new species of Clistopyga Gravenhorst (Hymenoptera: Ichneumonidae: Pimplinae) from the Western Ghats. Records of the Zoological Survey of India 119 60-63.

Morley C. (1914) A Revision of the Ichneumonidae Based on the Collection in the British Museum (Natural History) Part III. Tribes Pimplides and Bassides. British Museum, London, U.K. 148 pp.

Palacio E., Bordera S., Sääksjärvi I.E. & Díaz F. (2018) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part II: the C. isayae species group, with the description of seven new species. Zootaxa 4442 101-121.

Palacio E., Bordera S. & Díaz F. (2019) The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae) Part III: the C. henryi species group, with the description of three new species. Zootaxa 4563 103-118.

Porter C.C. (1979) Ichneumonidae de Tarapacá. I. Subfamilia Ephialtinae (Hymenoptera). Idesia 5 157-187.

Provancher L. (1880) Faune canadienne. Les insectes - Hyménoptères. Naturaliste Canadien 12 33-48.

Sääksjärvi I.E., Castillo C., Bordera S. & Broad G.R. (2015) Clistopyga caramba sp. nov. (Hymenoptera: Ichneumonidae; Pimplinae), an astonishing example of mimicry in spider-attacking parasitoid wasps. Zootaxa 4013 287-292.

Say T. (1835) Descriptions of new North American Hymenoptera, and observations on some already described. Boston Journal of Natural History 1 210-305.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. (1941) Beiträge zur Systematik der Tribus Polysphinctini Japans. Insecta Matsumurana 15 112-122.

Varga O. & Reshchikov A. (2015) New records of the genus Clistopyga Gravenhorst, 1829 (Hymenoptera: Ichneumonidae: Pimplinae) from the Oriental region, with description of a new species. Zootaxa 3964 561-568.



 3. Genus Tromatobia Förster, 1869 33 spp. + 3 subspecies

Förster A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 25 135-221.


argiopei Uchida, 1941

argiopes (Marshall, 1892)

blancoi Gauld, 1991

carballoi Gauld, Ugalde Gómez & Hanson, 1998

commutabilis Förster, 1888

dilutula (Benoit, 1953)

flavistellata Uchida & Momoi, 1957

forsiusi (Hellén, 1915)

huebrichi (Brèthes, 1913)

hutacharerni Kusigemati, 1988

koehleri (Blanchard, 1941)

lineata (Wollaston, 1858)

lineatoria (Villers, 1789)

lineatoria decorata Obrtel, 1953

lineatoria rubella (Costa, 1886)

lineiger Morley, 1914

maculata Momoi, 1970

nipponica Uchida, 1928

notator (Fabricius, 1804)

onerosa Förster, 1888

ornata (Gravenhorst, 1829)

ornata nigrescens Constantineanu, Ciochia, Constantineanu, Mustata & Ularu, 1967

ovivora (Boheman, 1821)

pacayae Gauld, Ugalde Gómez & Hanson, 1998

pacifica Momoi, 1966

quadricolor (Kriechbaumer, 1894)

rufopectus (Cresson, 1870)

sejuncta Förster, 1888

sponsa (Haliday, 1836)

sulcata (Benoit, 1953)

taiwana Kusigemati, 1984

tricolor (Brullé, 1846)

ulatei Gauld, Ugalde Gómez & Hanson, 1998

variabilis (Holmgren, 1856)

zonata (Davis, 1895)

zunigai Gauld, 1991

Benoit P.L.G. (1953) Notes Ichneumonologiques Africaines IV. Revue de Zoologie et de Botanique Africaines 47 153-159.

Blanchard E.E. (1942) Nuevos parásitos del bicho de cesto Oeceticus kirbyi Guild. Revista de la Sociedad Entomologica Argentina 11 3-21.

Boheman C.H. (1821) En ny art af Insect-slägtet Pimpla, hvars larv uppehåller sig i Spindelbon. Kungliga Svenska Vetenskaps Akadamiens Handlingar 42 335-337.

Brèthes J. (1913) Hymenópteros de la América meridional. Anales del Museo Nacional de Historia Natural de Buenos Aires 24 35-165.

Brullé M.A. (1846) Tome Quatrième. Des Hyménoptères. Les Ichneumonides. In: A. Lepeletier de Saint Fargeau (eds.). Histoire Naturelle Des Insectes. Hyménoptères. pp. 56-521. Paris, France.

Costa A. (1886) Notizie ed osservazioni sulla Geo-Fauna sarda Memoria Quinta Risultamento delle ricerche fatte in maggio 1885. Atti della Reale Accademia delle Scienze Fisiche e Matematiche di Napoli, ser.2a:vol.II:fasc.7, 24 pp.

Constantineanu M.I., Ciochia V., Constantineanu R.M., Mustata G. & Ularu G. (1967) Contributions a l'etude des Ichneumonides de la montagne de Gutii, région de Maramures. Analele ştiinţifice ale Universităţii “Al. I. Cuza” din Iaşi. Sect. II a 13 47-61.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Davis G.C. (1895) Two new species of Clistopyga. Entomological News 6 198-199.

Fabricius J.C. (1804) Systema Piezatorum: secundum ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. Carolum Reichard, Brunsvigae, 439+32 pp.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Gauld I.D., Ugalde Gómez J.A. & Hanson P.E. (1998) Guía de los Pimplinae de Costa Rica (Hymenoptera: Ichneumonidae). Revista de Biologia Tropical 46 (Suppl. 1) 1-189.

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Haliday A.H. (1836) Descriptions, &c. of the Hymenoptera. In: S. Curtis, A.H. Haliday & F. Walker (eds.). Descriptions &c. of the Insects Collected by Captain P.P. King, R.N., F.R.S., in the Survey of the Straits of Magellan. Transactions of the Linnean Society of London. pp. 316-331. London, U.K.

Hellén W. (1915) Beiträge zur Kenntnis der Ichneumoniden Finlands I. Subfamily Pimplinae. Acta Societatis pro Fauna et Flora Fennica 40 1-89.

Holmgren A.E. (1856) Entomologiska anteckningar under en resa i södra Sverige ar 1854. Kongliga Svenska Vetenskapsakademiens Handlingar 75 1-104.

Kriechbaumer J. (1894) Hymenoptera Ichneumonidea a medico nautico Dr. Joh. Brauns in itinere ad oras Africae occidentalis lecta. Berliner Entomologische Zeitschrift 39 43-68.

Kusigemati K. (1984) Some Ephialtinae of South East Asia, with descriptions of eleven new species (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific 5 126-150.

Kusigemati K. (1988) Descriptions of four new ichneumon flies parasitic on pine insect pests in Thailand. Memoirs of the Faculty of Agriculture, Kagoshima University 24 147-155.

Marshall T.A. (1892) Énumerations de quelques Hyménoptères du Venezuela. Annales de la Société Entomologique de France 61 60-76.

Momoi S. (1966) Some new Ichneumonidae (Hymenoptera) from New Guinea and adjacent areas. Pacific Insects 8 152-164.

Momoi S. (1970) Ichneumonidae (Hymenoptera) of the Ryukyu Archipelago. Pacific Insects 12 327-399.

Morley C. (1914) A Revision of the Ichneumonidae Based on the Collection in the British Museum (Natural History) Part III. Tribes Pimplides and Bassides. British Museum, London, U.K. 148 pp.

Obrtel R. (1953) [Notes on some interesting Icheumonidae in Silesia (Hym., Ichn.)]. Přírodovědecký sborník Ostravského kraje 14 192-208 (in Czech with English summary].

Schmiedeknecht O. (1888) Monographische Bearbeitung der Gattung Pimpla. Zoologische Jahrbücher Abteilung für Systematik 3 445-542.

Uchida T. (1928) Dritter Beitrag zur Ichneumoniden-Fauna Japans. Journal of the Faculty of Agriculture, Hokkaido Imperial University 25 1-115.

Uchida T. (1941) Eine neue Schmarotzerwespe von Spinnen. Insecta Matsumurana 15 161-162.

Uchida T. & Momoi S. (1957) Descriptions of three new species of the tribe Ephialtini from Japan (Hymenoptera, Ichneumonidae). Insecta Matsumurana 21 6-11.

Villers C. de (1789) Caroli Linnaei entomologia, Faunae Suecicae descriptionibus. Tomus tertius. Lugduni, 657 pp.

Wollaston T.V. (1858) Brief diagnostic characters of undescribed Madeiran insects. Annals and Magazine of Natural History ser.3:vol.1 18-28.



 4. Genus Zaglyptus Förster, 1869 27 spp. +7 subspecies

Förster A. (1869) Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 25 135-221.


ankaratrus Seyrig, 1934

arizonicus Townes, 1960

chavesi Gauld, 1991

cortesi Porter, 1979

divaricatus Baltazar, 1961

divaricatus basilanicus Baltazar, 1961

facifidus Baltazar, 1961

formosus Cushman, 1933

glaber Gupta, 1961

glabrinotus (Girault, 1925)

grandis Gupta, 1961

guptai Pham, Broad & Wägele, 2013

hallowayi Gauld, 1984

idukkiensis (Manjusha, Sudheer & Ghosh, 2019)

indicus Gupta, 1961

indicus nigrithorax Gupta, 1961

iwatai (Uchida, 1936)

miarus Gupta, 1961

multicolor (Gravenhorst, 1829)

multicolor rufescens (Boyer de Fonscolombe, 1854)

nigrolineatus Gupta, 1961

pictilis Townes, 1960

romeroae Gauld, 1991

rufus Hellén, 1949

ruslani Rizki & Idris, 2010

semirufus Momoi, 1970

semirufus marginatus Kasparyan, 1981

simonis (Marshall, 1892)

trituberculatus (Benoit, 1953)

varipes (Gravenhorst, 1829)

varipes albitegulis Constantineanu & Ciochia, 1968

varipes incompletus (Cresson, 1870)

varipes meridionator Aubert, 1969

wuyiensis He, 1984

Aubert J.F. (1969) Ichneumonides non petiolees inédites et révision partielle des genres Phytodietus Grav. et Netelia Gray. Bulletin de la Société Entomologique de Mulhouse 1968(novembre-decembre) 93-103.

Baltazar C.R. (1961) The Philippine Pimplini, Poemeniini, Rhyssini, and Xoridini (Hymenoptera, Ichneumonidæ, Pimplinæ). Monographs of the National Institute of Science and Technology 7 1-130.

Benoit P.L.G. (1953) Notes Ichneumonologiques Africaines IV. Revue de Zoologie et de Botanique Africaines 47 153-159.

Boyer de Fonscolombe M. (1854) Ichneumonologie provençale, ou catalogue des ichneumoisïdes qui se trouvent aux environs d'Aix, et description des espèces inédites. Annales de la Société Entomologique de France ser.3:t.2 479-520.

Constantineanu M.I. & Ciochia V. (1968) Ichneumonides nouveaux de Roumanie. Revue Roumaine de Biologie (Serie de Zoologie) 13 43-47.

Cresson E.T. (1870) Descriptions of new species belonging to the subfamily Pimplariae found in America north of Mexico. Transactions of the American Entomological Society (Philadelphia) 3 143-172.

Cushman R.A. (1933) H. Sauter's Formosa-collection: Subfamily Ichneumoninae (Pimplinae of Ashmaed). Insecta Matsumurana 8 1-50.

Gauld I.D. (1984) The Pimplinae, Xoridinae, Acaenitinae and Lycorininae of Australia. Bulletin of the British Museum (Natural History) Entomology 49 235-339.

Gauld I.D. (1991) The Ichneumonidae of Costa Rica, 1. Introduction, keys to subfamilies, and keys to the species of the lower pimpliform subfamilies Rhyssinae, Pimplinae, Poemeniinae, Acaenitinae and Cylloceriinae. Memoirs of the American Entomological Institute 47 1-589.

Girault A.A. (1925) Records and distributions of Australian Ophioninae (Ichneumon-flies). Queensland Agricultural Journal 24 538-541.

Gravenhorst J.L.C. (1829) Ichneumonologia Europaea. Pars III. Vratislaviae. 1097 pp.

Gupta V.K. (1960) A revision of the Oriental species of the genus Zaglyptus (Hymenoptera: Ichneumonidae). Indian Journal of Entomology 22 244-257.

He J.H. (1984) [Notes on Chinese species of the genus Zaglyptus Förster, with descriptions of a new species and a new species-group (Hymenoptera: Ichneumonidae)]. Wuyi Science Journal 4 199-204 (in Chinese with English summary).

Hellén W. (1949) Zur Kenntnis der Ichneumonidenfauna der Atlantischen Inseln. Commentationes Biologicae Societas Scientiarum Fennica 8 1-23.

Kasparyan D.R. (1981) [New species of the subfamily Pimplinae (Hymenoptera, Ichneumonidae), parasites of spiders in the Far East of the USSR.]. Trudy Zoologicheskogo Instituta 92 108-112 (in Russian).

Manjusha B.M., Sudheer K. & Ghosh S.M. (2019) Polysphincta idukkiensis (Hymenoptera: Ichneumonidae: Pimplinae) a rare new species from the southern Western Ghats. Halteres 10 96-99.

Marshall T.A. (1892) Énumerations de quelques Hyménoptères du Venezuela. Bulletin de la Societe Entomologique de France 61 60-76.

Momoi S. (1970) Ichneumonidae (Hymenoptera) of the Ryukyu Archipelago. Pacific Insects 12 327-399.

Pham N.T., Broad G.R. & Wägele W. (2013) First record of the genus Zaglyptus Förster (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam, with description of one new species. Zootaxa 3702 483-492.

Porter C.C. (1979) Ichneumonidae de Tarapacá. I. Subfamilia Ephialtinae (Hymenoptera). Idesia 5 157-187.

Rizki A. & Idris A.B. (2010) One new species and one new record of the genus Zaglyptus Foerster (Hymenoptera: Ichneumonidae: Pimplinae) from Malaysia. Serangga 15 9-24.

Seyrig A. (1934) Les Ichneumonides de Madagascar. 2. Ichneumonidae Tryphoninae et supplément aux 1. Pimplinae. Mémoires de I'Académie Malgache 19 1-111.

Townes H. & Townes M. (1960) Ichneumon-flies of American North of Mexico 2. Subfamily Ephialtinae, Xoridinae, Acaenitinae. United States National Museum Bulletin 216 1-676.

Uchida T. (1936) Drei neue Gattungen sowie acht neue und fuenf unbeschriebene Arten der Ichneumoniden aus Japan. Insecta Matsumurana 10 111-122.