Aulonochares lingulatus Girón and Short, 2019: 119
Distribution: French Guiana; Suriname
Aulonochares novoairensis Girón and Short, 2019: 119
Distribution: Brazil (Amazonas)
Aulonochares tubulus Girón and Short, 2019: 120
Distribution: Brazil (Roraima); Guyana; Suriname; Venezuela
Girón, J. C. & A. E. Z. Short. 2019. Three additional new genera of acidocerine water scavenger beetles from the Guiana and Brazilian Shield regions of South America (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys, 855: 109–154. https://doi.org/10.3897/zookeys.855.33013
Syn. Crephelochares Kuwert, 1890
Crepidelochares Ganglbauer, 1904
Dieroxenus Spangler, 1979: 753 [by Girón & Short 2018: 154]
Chasmogenus acuminatus Smith & Short, 2020: 32
Distribution: Brazil (Amapá, Pará), French Guiana, Guyana, Suriname
Chasmogenus amplius Smith & Short, 2020: 35
Distribution: Venezuela (Amazonas)
Chasmogenus australis García, 2000: 52
Distribution: Brazil (Roraima), French Guiana, Guyana, Venezuela
Chasmogenus bariorum García, 2000: 49
Syn. Chasmogenus occidentalis García, 2000: 49 [by Smith & Short 2020: 40]
Syn. Chasmogenus yukparum García, 2000: 50 [by Smith & Short 2020: 40]
Distribution: Venezuela (Zulia)
Chasmogenus barrae Short, 2005: 194
Distribution: Costa Rica (Guanacaste)
Chasmogenus berbicensis Smith & Short, 2020: 47
Distribution: Guyana
Chasmogenus brownsbergensis Smith & Short, 2020: 48
Distribution: Suriname
Chasmogenus cajuina Alves, Clarkson & Lima, 2020: 580
Distribution: Brazil (Piauí)
Chasmogenus castaneus Smith & Short, 2020: 50
Distribution: Venezuela (Zulia)
Chasmogenus clavijoi Smith & Short, 2020: 53
Distribution: Venezuela (Guárico)
Chasmogenus clinatus Glynn & Short, 2021: 436
Distribution: Brazil (Amazonas); Venezuela (Amazonas)
Chasmogenus cremnobates (Spangler, 1979: 754)
Syn. Dieroxenus cremnobates Spangler, 1979: 754 [by Girón & Short 2018: 155]
Distribution: Ecuador (Napo)
Chasmogenus cuspifer Smith & Short, 2020: 54
Distribution: Venezuela (Zulia)
Chasmogenus flavomarginatus Smith & Short, 2020: 55
Distribution: Venezuela (Barinas)
Chasmogenus fluminensis Clarkson & Ferreira, 2014
Distribution: Brazil: Rio de Janeiro
Chasmogenus fragilis Sharp, 1882: 73
Distribution: Guatemala; Panama
Chasmogenus gato Smith & Short, 2020: 56
Distribution: Venezuela (Amazonas)
Chasmogenus gironae Glynn & Short, 2021: 440
Distribution: Venezuela (Amazonas)
Chasmogenus guianensis Smith & Short, 2020: 58
Distribution: Guyana, Suriname
Chasmogenus ignotus Smith & Short, 2020: 60
Distribution: Brazil (Amazonas)
Chasmogenus inpa Glynn & Short, 2021: 440
Distribution: Brazil (Amazonas)
Chasmogenus itatiaia Clarkson & Ferreira, 2014
Distribution: Brazil (Minas Gerais, Rio de Janeiro)
Chasmogenus ligulatus Smith & Short, 2020: 61
Distribution: Suriname
Chasmogenus lilianae Clarkson & Ferreira, 2014
Distribution: Brazil (Rio de Janeiro)
Chasmogenus lineatus Smith & Short, 2020: 64
Distribution: Venezuela (Guárico)
Chasmogenus lorenzo Short, 2005: 195
Distribution: Costa Rica (Alajuela)
Chasmogenus pandus Smith & Short, 2020: 68
Distribution: Brazil (Amapá), French Guiana, Suriname
Chasmogenus rufinasus (Knisch, 1924: 124)
Distribution: Ecuador (Guayaquil)
Chasmogenus ruidus Short 2005: 196
Distribution: Costa Rica (Limón)
Chasmogenus sapucay Fernández, 1986: 192
Distribution: Argentina; Paraguay
Chasmogenus schmits Smith & Short, 2020: 69
Distribution: Suriname
Chasmogenus schoedli Short 2005: 197
Distribution: Costa Rica (Guanacaste)
Chasmogenus sinnamarensis Smith & Short, 2020: 70
Distribution: French Guyana
Chasmogenus tafelbergensis Smith & Short, 2020: 71
Distribution: Suriname
Chasmogenus ubatuba Clarkson & Ferreira, 2014: 491
Distribution: Brazil (São Paulo)
Chasmogenus undulatus Smith & Short, 2020: 73
Distribution: Guyana
Clarkson, B., & Ferreira-Jr., N. 2014. Four new species and first nominal record of Chasmogenus Sharp, 1882 (Coleoptera: Hydrophilidae) from Brazil. Zootaxa, 3765(5): 481–494. https://biotaxa.org/Zootaxa/article/view/zootaxa.3765.5.6
Fernández, L.A. 1986. Consideraciones sobre el género Chasmogenus Sharp y descripción de Chasmogenus sapucay sp. nov. (Coleoptera: Hydrophilidae). Neotropica, 32, 189–193.
García, M. 2000. Four new species of Chasmogenus Sharp, 1882 (Coleoptera: Hydrophilidae: Hydrophilinae) from Venezuela. Boletín del Centro de Investigaciones Biológicas Universidad del Zulia, 34(1): 45–58. http://www.produccioncientificaluz.org/index.php/boletin/article/view/218
Girón J.C. & Short A.E.Z. 2018. Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 768: 113–158. https://doi.org/10.3897/zookeys.768.24423
Glynn, R.D., Short, A.E.Z. (2021) New species and records of Chasmogenus Sharp, 1882 from the southwestern margin of the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 5048: 435–443. https://doi.org/10.11646/zootaxa.5048.3.8
Hebauer, F. 1992. The species of the genus Chasmogenus Sharp, 1882 (Coleoptera, Hydrophilidae). Acta Coleopterologica, 8(2), 61–92. https://web.archive.org/web/20160304194744/http://www.hydrophiloidea.org/pdfs/Hebauer1992_Chasmogenus.pdf
Sharp, D. 1882. Fam. Hydrophilidae. Biologia Centrali-Americana Insecta. Coleoptera, 1(2): 53–80. https://www.biodiversitylibrary.org/item/14601#page/75/mode/1up
Short, A.E.Z. 2005. A review of the subtribe Acidocerina of Central America with special reference to Costa Rica (Coleoptera: Hydrophilidae). Koleopterologische Rundschau, 75: 191–226. http://www.zobodat.at/pdf/KOR_75_2005_0191-0226.pdf
Smith, R.R. & Short, A.E.Z. (2020) Review of the genus Chasmogenus Sharp, 1882 of northeastern South America with an emphasis on Venezuela, Suriname, and Guyana (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 934: 25–79. https://doi.org/10.3897/zookeys.934.49359
Spangler, P.J. 1979. A new genus of madicolous beetles from Ecuador (Coleoptera: Hydrophilidae: Hydrobiinae). Proceedings of the Biological Society of Washington 92(4), 753–761. https://www.biodiversitylibrary.org/item/110033#page/787/mode/1up
Crucisternum escalera Girón & Short 2018: 120
Distribution: Venezuela (Bolívar)
Crucisternum ouboteri Girón & Short 2018: 121
Distribution: French Guiana; Guyana; Suriname; Venezuela (Amazonas)
Crucisternum queneyi Girón & Short 2018: 123
Distribution: French Guiana
Crucisternum sinuatus Girón & Short 2018: 124
Distribution: Brazil (Minas Gerais, Pará)
Crucisternum toboganensis Girón & Short 2018: 126
Distribution: Venezuela (Amazonas)
Crucisternum vanessae Girón & Short 2018: 127
Distribution: Suriname
Crucisternum xingu Girón & Short 2018: 131
Distribution: Brazil (Pará)
Girón J.C. & Short A.E.Z. 2018. Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 768: 113–158. https://doi.org/10.3897/zookeys.768.24423
Ephydrolithus hamadae Girón and Short, 2019: 130
Distribution: Brazil (Minas Gerais)
Ephydrolithus minor Girón and Short, 2019: 130 - Brazil, Bahia
Distribution: Brazil (Bahia)
Ephydrolithus ogmos Girón and Short, 2019: 131
Distribution: Brazil (Bahia)
Ephydrolithus spiculatus Girón and Short, 2019: 132
Distribution: Brazil (Minas Gerais)
Ephydrolithus teli Girón and Short, 2019: 132
Distribution: Brazil (Bahia, Minas Gerais)
Girón, J. C. & A. E. Z. Short. 2019. Three additional new genera of acidocerine water scavenger beetles from the Guiana and Brazilian Shield regions of South America (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys, 855: 109–154. https://doi.org/10.3897/zookeys.855.33013
Globulosis hemisphericus García, 2001: 156
Distribution: Venezuela: Bolívar; Guyana, Suriname, Brazil: Amazonas, Pará
Globulosis flavus Short, García, & Girón, 2017
Distribution: Venezuela: Amazonas, Apure, Bolívar
García, M. 2001. Nueva subtribu, género y especie de Hydrophilini (Coleoptera: Hydrophilidae) del extremo suroriental de Venezuela. Boletín del Centro de Investigaciones Biológicas Universidad del Zulia, 35(2), 151–160. http://produccioncientificaluz.org/index.php/boletin/article/view/3978
Short, A.E.Z., García, M., Girón, J.C. 2017. Revision of the Neotropical water scavenger beetle genus Globulosis García, 2001 (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa 4232(2): 271–281. https://biotaxa.org/Zootaxa/article/view/zootaxa.4232.2.10
Helobata amazonensis Clarkson, Dias Dos Santos & Ferreira, 2016: 550
Distribution: Brazil (Amazonas)
Helobata aschnakiranae Makhan, 2007: 1
Distribution: Suriname
Helobata bitriangulata García, 2000: 244
Distribution: Venezuela (Apure)
Helobata confusa Fernández & Bachmann, 1987: 155
Distribution: Paraguay (Asunción)
Helobata corumbaensis Fernández & Bachmann, 1987: 155
Distribution: Brazil (Mato Grosso do Sul)
Helobata cossyphoides (Bruch, 1915: 458)
Distribution: Argentina (Buenos Aires, La Plata)
Helobata cuivaum García, 2000: 242
Distribution: Venezuela (Apure)
Helobata larvalis (Horn, 1873) [reinstated as valid name by Hansen 1991: 923]
Syn. Helopeltis larvalis Horn, 1873: 137
Helopeltina larvalis Horn,
Helobata striata (Brullé)
Hydrophilus (Philhydrus) striatus Brullé, 1841: 58 [primary homonym of Hydrophilus striatus Turton, 1802 and Hydrophilus striatus Say, 1825]
Philhydrus striatus (Brullé)
Helopeltis striatus (Brullé)
Enochrus (Lumetus) striatus (Brullé)
Distribution: Argentina (Misiones, Chaco, Corrientes, Entre Ríos, Santa Fé, Buenos Aires, Santiago del Estero, Salta); Bolivia; Brazil; Paraguay; Venezuela; Guatemala, Mexico; Cuba; USA (Florida, Louisiana, Mississippi, North Carolina, South Carolina, Texas)
Helobata lilianae García, 2000: 239
Distribution: Venezuela (Apure)
Helobata pantaneira Clarkson, Dias Dos Santos & Ferreira, 2016: 553
Distribution: Brazil (Mato Grosso)
Helobata perpunctata Fernández & Bachmann, 1987: 156
Distribution: Argentina (Chaco)
Helobata quatipuru Fernández & Bachmann, 1987: 158
Distribution: Brazil (Pará)
Helobata soesilae Makhan, 2007: 3
Distribution: Suriname
Clarkson, B., Dias Dos Santos, A. & Ferreira-Jr., N. 2016. On Brazilian Helobata Bergroth, 1888 (Coleoptera: Hydrophilidae): description of two new species, new records, and key to species. Zootaxa, 4126(4): 548-562. http://www.mapress.com/j/zt/article/view/zootaxa.4126.4.6
Fernández, L. A., & Bachmann, A. O. 1987. Revisión del género Helobata Bergroth (Coleoptera: Hydrophilidae). Revista de la Sociedad Entomológica Argentina, 44(2), 1985.
García, M. 2000. Tres nuevas especies de Helobata Bergroth 1888 (Hydrophilidae: Hydrophilinae), de Venezuela. Boletín del Centro de Investigaciones Biológicas, 34(2): 237-246. http://www.produccioncientificaluz.org/index.php/boletin/article/view/201
Horn, G. H. 1873. Revision of the genera and species of the tribe Hydrobiini. Proceedings of the American Philosophical Society, 13(90): 118-137. https://www.jstor.org/stable/981607
Makhan, D. 2007. Helobata soesilae sp. nov. and Helobata aschnakiranae sp. nov. from Suriname (Coleoptera: Hydrophilidae). Calodema Supplementary Paper, 14, 1-3. https://www.researchgate.net/publication/321107032_Helobata_soesilae_sp_nov_and_Helobata_aschnakiranae_sp_nov_from_Suriname_Coleoptera_Hydrophilidae
Helochares championi Sharp, 1882: 75
Syn. Helochares (Hydrobaticus) championi Sharp, 1882: 75
Distribution: Costa Rica (Cartago, Puntarenas); Guatemala; Nicaragua
Helochares laevis Short & Girón, 2018: 36
Syn. Helochares (Hydrobaticus) laevis Short & Girón, 2018: 36
Distribution: Mexico (Chiapas)
Helochares maculicollis Mulsant, 1844: 379
Syn. Philhydrus maculicollis (Mulsant)
Philhydrus (Helochares) maculicollis (Mulsant)
Helochares (Grapidelochares) maculicollis Mulsant
Helochares (Hydrobaticus) maculicollis Mulsant, 1844: 379
? Helochares bipunctatus Sharp, 1882: 76 [Mexico; Guatemala]
Distribution: USA: Alabama, Arkansas, District of Columbia, Florida, Georgia, Illinois, Indiana, Iowa, Kansas, Kentucky, Louisiana, Maryland, Mississippi, Missouri, North Carolina, Ohio, Oklahoma, Pennsylvania, South Carolina, Tennessee, Texas, Virginia; ?Guatemala; ?Mexico
Helochares nexus Short & Girón, 2018: 39
Syn. Helochares (Hydrobaticus) nexus Short & Girón, 2018: 39
Distribution: Panama (Coclé, Colón, Panama); Ecuador (Esmeraldas, Guayas, Manabí), Venezuela (Zulia)
Helochares normatus (LeConte, 1861: 341)
Syn. Philhydrus normatus LeConte 1861: 341
Helochares normatus (LeConte, 1861)
Chasmogenus normatus (LeConte, 1861)
Helochares seriatus Sharp, 1882: 76 [Guatemala]
Helochares (Grapidelochares) seriatus Sharp
? Helochares regularis Sharp, 1882: 76 [Mexico]
? Helochares (Grapidelochares) regularis Sharp
Helochares (Hydrobaticus) normatus (LeConte, 1861: 341)
Distribution: USA (Arizona, California, Nevada, Oregon, Texas); Guatemala; Costa Rica (Guanacaste); El Salvador, Honduras, Mexico (Baja California, Baja California del Sur, Colima, Morelos, Nayarit, Puebla, Sonora, Veracruz), Nicaragua (Madriz, Managua, Matagalpa, Rivas)
Helochares politus Short & Girón, 2018: 45
Syn. Helochares (Hydrobaticus) politus Short & Girón, 2018: 45
Distribution: Guatemala (Huehuetenango)
Helochares trujillo Short & Girón, 2018: 45
Syn. Helochares (Hydrobaticus) trujillo Short & Girón, 2018: 45
Distribution: Venezuela (Mérida, Trujillo)
Helochares zamora Short & Girón, 2018: 46
Syn. Helochares (Hydrobaticus) zamora Short & Girón, 2018: 46
Distribution: Ecuador (Pastaza, Zamora-Chinchipe)
Bruch, C. 1915. Nuevas especies de coleópteros hidrofílidos. Revista del Museo de la Plata, 19(2): 447–470.
Clarkson, B., & Ferreira-Jr, N. 2014. A new species and records of Helochares (Insecta: Coleoptera: Hydrophilidae) from Southeastern Brazil. Zoologia, 31(4): 400–404. http://www.scielo.br/pdf/zool/v31n4/12.pdf
Fernández, L. A. 1981. Dos especies nuevas del género Helochares (Coleoptera: Hydrophilidae). Limnobios, 2(3): 189–192.
Fernández, L. A. 1982a. Notas sobre el género Helochares (Insecta, Coleoptera: Hydrophilidae). Neotropica, 8(79): 31-40.
Fernández, L.A. 1982b. Cinco especies nuevas del género Helochares (Coleoptera: Hydrophilidae). Physis (Buenos Aires), secc. b, 40(99): 85-90.
Fernández, L.A. 1983. Helochares (Sindolus) talarum sp. nov., y redescripción de Helochares (Helochares) pallipes (Brullé), con descripción de los estados preimaginales (Coleoptera: Hydrophilidae). Limnobios, 2(6): 439-449.
Fernández, L.A. 1989. Notas sobre el género Helochares. II (Coleoptera: Hydrophilidae). Descripción de dos especies nuevas neotropicales. Clave para determinar las especies argentinas y de áreas vecinas. Revista de la Sociedad Entomológica Argentina, 45(1-4): 145-151.
LeConte, J.L. 1861. New species of Coleoptera inhabiting the Pacific district of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 13, 338–359. https://www.biodiversitylibrary.org/item/17713#page/350/mode/1up
MacLeay, W.S. 1871. Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales, 2, 79–205. https://www.biodiversitylibrary.org/item/54870#page/91/mode/1up
Mulsant, E. (1844) Description de quelques Palpicornes inédits. Annales de la Société d’ Agriculture de Lyon, 7, 372–382. https://books.google.com/books?id=NIg9AQAAMAAJ&lpg=PT3&ots=ip_COff4dp&dq=Description%20de%20quelques%20Palpicornes%20in%C3%A9dits.&pg=PA372#v=onepage&q&f=false
Sharp, D. 1882. Fam. Hydrophilidae. Biologia Centrali-Americana Insecta. Coleoptera, 1(2): 53–80. https://www.biodiversitylibrary.org/item/14601#page/75/mode/1up
Short, A. E. Z. & J. C. Girón. (2018) Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa, 4407(1): 29–50. http://www.mapress.com/j/zt/article/view/zootaxa.4407.1.2
Katasophistes charynae Girón & Short, 2018: 136
Distribution: Peru (Madre de Dios)
Katasophistes cuzco Girón & Short, 2018: 136
Distribution: Peru (Cuzco)
Katasophistes merida Girón & Short, 2018: 136
Distribution: Venezuela (Mérida)
Katasophistes superficialis Girón & Short, 2018: 136
Distribution: Ecuador (Pastaza)
Girón J.C. & Short A.E.Z. 2018. Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 768: 113–158. https://doi.org/10.3897/zookeys.768.24423
Nanosaphes castaneus Girón & Short, 2018: 146
Distribution: Brazil (Pará)
Nanosaphes hesperus Girón & Short 2018: 148
Distribution: Guyana, Suriname
Nanosaphes punctatus Girón & Short, 2018: 151
Distribution: Guyana, Suriname
Nanosaphes tricolor Girón & Short, 2018: 151
Distribution: Guyana, Suriname
Girón J.C. & Short A.E.Z. 2018. Three new genera of acidocerine water scavenger beetles from tropical South America (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 768: 113–158. https://doi.org/10.3897/zookeys.768.24423
Novochares abbreviatus (Fabricius, 1801: 251)
Syn. Hydrophilus abbreviatus Fabricius, 1801: 251 ("America Meridionali")
Philydrus pallidus Castelnau, 1840: 53 [secondary homonym of Hydrophilus pallidus Rossi, 1792 (Brazil)]
Philhydrus pallidus Castelnau
Helochares pallidus (Castelnau)
Enochrus (Lumetus) pallidus (Castelnau)
Helochares (Hydrobaticus) rufobrunneus Balfour-Browne, 1939: 293 (Lesser Antilles, Grenada)
Helochares (s. str) abbreviatus (Fabricius, 1801: 251)
Distribution: Argentina; Bolivia; Brazil; Colombia; French Guiana; Paraguay; Suriname; Venezuela; Panama; Cuba; Lesser Antilles
Novochares atlanticus (Clarkson & Ferreira-Jr. 2014: 401)
Syn. Helochares (s. str.) atlanticus Clarkson & Ferreira-Jr. 2014: 401
Distribution: Brazil
Novochares atratus (Bruch, 1915: 451)
Syn. Helochares atratus Bruch
Helochares (s. str.) parhedrus d'Orchymont, 1939: 259
Helochares (s. str) atratus Bruch, 1915: 451
Distribution: Argentina; Brazil; ?Ecuador; Paraguay
Novochares bolivianus (Fernández, 1989: 146)
Syn. Helochares (s. str) bolivianus Fernández, 1989: 146
Distribution: Bolivia
Novochares carmona (Short, 2005: 215)
Syn. Helochares (s. str.) carmona Short, 2005: 215
Distribution: Costa Rica (Guanacaste)
Novochares chaquensis (Fernández, 1982: 87)
Syn. Helochares (s. str) chaquensis Fernández, 1982: 87
Distribution: Argentina; Brazil
Novochares cochlearis (Fernández, 1982: 89)
Syn. Helochares (s. str) cochlearis Fernández, 1982: 89
Distribution: Argentina; Paraguay
Novochares coya (Fernández, 1982: 87)
Syn. Helochares (s. str) coya Fernández, 1982: 87
Distribution: Bolivia
Novochares guadelupensis (d'Orchymont, 1926: 233)
Syn. Helochares (s. str) guadelupensis d'Orchymont, 1926: 233
Distribution: Lesser Antilles: Guadeloupe
Novochares inornatus (d'Orchymont, 1926: 235)
Syn. Helochares (s. str) inornatus d'Orchymont, 1926: 235
Distribution: Brazil; French Guiana
Novochares oculatus (Sharp, 1882: 74)
Syn. Helochares (s. str) oculatus Sharp, 1882: 74
Distribution: Argentina; Brazil; Guatemala; Panama; ?Mexico; ?Antilles
Novochares pallipes (Brullé, 1841: 58)
Syn. Hydrophilus (Philydrus) pallipes Brullé, 1841: 58 (Uruguay)
Philhydrus pallipes (Brullé)
Helochares pallipes (Brullé)
Helochares (s. str) pallipes (Brullé, 1841: 58)
Distribution: Argentina; Brazil; Paraguay; Uruguay
Novochares pichilingue (Fernández, 1989: 147)
Syn. Helochares (s. str) pichilingue Fernández, 1989: 147
Distribution: Ecuador
Novochares sallaei (Sharp, 1882: 75)
Syn. Helochares sallaei Sharp, 1882: 75
Helochares (s. str) Sellae Sharp
Philhydrus estriatus Blatchley, 1917: 139 (USA: Florida)
Enochrus (Lumetus) estriatus (Blatchley)
Helochares (s. str) sallaei Sharp, 1882: 75
Distribution: Mexico; USA: Florida
Novochares tectiformis (Fernández, 1982: 88)
Syn. Helochares (s. str) tectiformis Fernández, 1982: 88
Distribution: Argentina; Brazil; Paraguay; Venezuela
Bruch, C. 1915. Nuevas especies de coleópteros hidrofílidos. Revista del Museo de la Plata, 19(2): 447–470.
Brullé A (1841) Famille des Hydrophiliens. In: d’Orbigny A. Voyage dans l’Amérique méridionale, tome sexiéme, 2e partie, Insectes. P. Bertrand, Paris, France, 52–59. https://biodiversitylibrary.org/page/2531016
Clarkson, B., & Ferreira-Jr, N. 2014. A new species and records of Helochares (Insecta: Coleoptera: Hydrophilidae) from Southeastern Brazil. Zoologia, 31(4): 400–404. http://www.scielo.br/pdf/zool/v31n4/12.pdf
d’Orchymont, A. (1926) Contribution à l’étude des Hydrophilides VI. Bulletin et Annales de la Société entomologique de Belgique 66: 201–248.
Fabricius, J.C. (1801) Systema Eleutheratorum: secundum ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. Bibliopolii Academici Novi, Kiliae, 24 + 506 pp.
Fernández, L. A. 1981. Dos especies nuevas del género Helochares (Coleoptera: Hydrophilidae). Limnobios, 2(3): 189–192.
Fernández, L. A. 1982a. Notas sobre el género Helochares (Insecta, Coleoptera: Hydrophilidae). Neotropica, 8(79): 31-40.
Fernández, L.A. 1982b. Cinco especies nuevas del género Helochares (Coleoptera: Hydrophilidae). Physis (Buenos Aires), secc. b, 40(99): 85-90.
Fernández, L.A. 1989. Notas sobre el género Helochares. II (Coleoptera: Hydrophilidae). Descripción de dos especies nuevas neotropicales. Clave para determinar las especies argentinas y de áreas vecinas. Revista de la Sociedad Entomológica Argentina, 45(1-4): 145-151.
Sharp, D. 1882. Fam. Hydrophilidae. Biologia Centrali-Americana Insecta. Coleoptera, 1(2): 53–80. https://www.biodiversitylibrary.org/item/14601#page/75/mode/1up
Short, A.E.Z. (2005) A review of the subtribe Acidocerina of Central America with special reference to Costa Rica (Coleoptera: Hydrophilidae). Koleopterologische Rundschau: 191–226. http://www.zobodat.at/pdf/KOR_75_2005_0191-0226.pdf
Primocerus cuspidis Girón and Short, 2019: 144
Distribution: Venezuela (Amazonas)
Primocerus gigas Girón and Short, 2019: 145
Distribution: Venezuela (Amazonas)
Primocerus maipure Girón and Short, 2019: 146
Distribution: Venezuela (Amazonas)
Primocerus neutrum Girón and Short, 2019: 147
Distribution: Guyana; Suriname; Venezuela (Bolívar)
Primocerus ocellatus Girón and Short, 2019: 148
Distribution: Venezuela (Amazonas)
Primocerus petilus Girón and Short, 2019: 148
Distribution: Brazil (Pará)
Primocerus pijiguaense Girón and Short, 2019: 149
Distribution: Venezuela (Bolívar)
Primocerus semipubescens Girón and Short, 2019: 150
Distribution: Guyana
Primocerus striatolatus Girón and Short, 2019: 151
Distribution: Suriname
Girón, J. C. & A. E. Z. Short. 2019. Three additional new genera of acidocerine water scavenger beetles from the Guiana and Brazilian Shield regions of South America (Coleoptera: Hydrophilidae: Acidocerinae). ZooKeys, 855: 109–154. https://doi.org/10.3897/zookeys.855.33013
Quadriops acroreius Girón & Short, 2017: 123
Distribution: Suriname; French Guiana
Quadriops clusia Girón & Short, 2017: 125
Distribution: Guyana; Suriname; Brazil (Amazonas)
Quadriops dentatus Hansen, 1999: 134
Distribution: Venezuela (Bolívar); French Guiana; Suriname
Quadriops depressus Hansen, 1999: 136
Syn. Quadriops amazonensis García, 2000: 59
Quadriops politus Hansen, 1999: 135
Distribution: Peru; Ecuador; Venezuela (Amazonas)
Quadriops reticulatus Hansen, 1999: 135
Distribution: Costa Rica, Panama
Quadriops similaris Hansen, 1999: 136
Distribution: Venezuela (Bolívar); Guyana; Suriname; French Guiana
Hansen, M. 1999. Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family. Insect Systematics & Evolution 30(2): 121–172. https://doi.org/10.1163/187631200X00228
García, M. 2000. Una nueva especie de Quadriops Hansen, 1999 (Coleoptera: Hydrophilidae: Hydrophilinae) de Venezuela. Boletín del Centro de Investigaciones Biológicas 34(1): 59–65. http://produccioncientificaluz.org/index.php/boletin/article/view/219
Girón, J.C., Short A.E.Z. 2017. Revision of the Neotropical water scavenger beetle genus Quadriops Hansen, 1999 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 705: 115–141. https://doi.org/10.3897/zookeys.705.19815
Radicitus ayacucho Short & García, 2014: 252
Distribution: Venezuela (Amazonas, Bolívar)
Radicitus granitum Short & García, 2014: 254
Distribution: Venezuela (Amazonas, Bolívar)
Radicitus surinamensis Short & García, 2014: 257
Distribution: Suriname
Short A.E.Z. & García, M. 2014. A new genus of egg case-carrying water scavenger beetle from the Guiana Shield (Coleoptera: Hydrophilidae: Acidocerinae). Zootaxa, 3835(2): 251–262. https://biotaxa.org/Zootaxa/article/view/zootaxa.3835.2.5
Sindolus femoratus (Brullé, 1841: 59)
Syn. Hydrophilus (Philydrus) femoratus Brullé, 1841: 59 [Argentina]
Hydrobius femoratus (Brullé)
Helochares femoratus Brullé
Helochares (Sindolus) femoratus Brullé
? Philhydrus spadiceus Mulsant, 1844: 380 [Colombia; French Guiana]
? Enochrus (Lumetus) spadiceus (Mulsant)
Helochares gravidus Bruch, 1915: 452
Helochares (Sindolus) gravidus Bruch
Helochares (Sindolus) femoratus (Brullé, 1841: 59)
Distribution: Argentina; Brazil; ? Colombia; ? French Guiana; Lesser Antilles: Antigua
Sindolus mesostitialis (Fernández, 1981: 189)
Syn. Helochares (Sindolus) mesostitialis Fernández, 1981: 189
Distribution: Argentina; Brazil
Sindolus mini (Fernández, 1982: 89)
Syn. Helochares (Sindolus) mini Fernández, 1982: 89
Distribution: Argentina; Paraguay
Sindolus mundus Sharp, 1882: 73
Syn. Helochares (Sindolus) mundus Sharp, 1882: 73 [Mexico]
Distribution: Mexico
Sindolus optatus Sharp, 1882: 72
Syn. Helochares (s. str.) guatemalensis Knisch, 1921: 68 [Guatemala]
Helochares (Sindolus) guatemalensis Knisch
Helochares (Sindolus) optatus (Sharp, 1882: 72) [Guatemala]
Distribution: Guatemala
Sindolus spatulatus (Fernández, 1981: 191)
Syn. Helochares (Sindolus) spatulatus Fernández, 1981: 191
Distribution: Argentina
Sindolus talarum (Fernández, 1983: 440)
Syn. Helochares (Sindolus) talarum Fernández, 1983: 440
Distribution: Argentina
Sindolus ventricosus (Bruch, 1915: 452)
Syn. Hydrophilus (Philydrus) gibbus Brullé, 1841: 58 (primary homonym of Hydrophilus gibbus Illiger, 1801 and Hydrophilus gibbus Thunberg, 1820) [Argentina]
Philhydrus gibbus (Brullé)
Helochares gibbus (Brullé)
Helochares (Sindolus) gibbus (Brullé)
Helochares (Sindolus) ventricosus Bruch, 1915: 452
Distribution: Argentina; Bolivia; Brazil; Paraguay; Uruguay
Bruch, C. 1915. Nuevas especies de coleópteros hidrofílidos. Revista del Museo de la Plata, 19(2): 447–470.
Brullé, A. (1841) Famille des Hydrophiliens. d’Orbigny A. Voyage dans l’Amérique méridionale, tome sexiéme, 2e partie, Insectes. P. Bertrand, Paris, France, 52–59. https://biodiversitylibrary.org/page/2531016
Fernández, L. A. 1981. Dos especies nuevas del género Helochares (Coleoptera: Hydrophilidae). Limnobios, 2(3): 189–192.
Fernández, L.A. 1982b. Cinco especies nuevas del género Helochares (Coleoptera: Hydrophilidae). Physis (Buenos Aires), secc. b, 40(99): 85-90.
Fernández, L.A. 1983. Helochares (Sindolus) talarum sp. nov., y redescripción de Helochares (Helochares) pallipes (Brullé), con descripción de los estados preimaginales (Coleoptera: Hydrophilidae). Limnobios, 2(6): 439-449.
Sharp, D. 1882. Fam. Hydrophilidae. Biologia Centrali-Americana Insecta. Coleoptera, 1(2): 53–80. https://www.biodiversitylibrary.org/item/14601#page/75/mode/1up
Tobochares akoerio Girón & Short, 2021: 120
Distribution: Suriname
Tobochares arawak Girón & Short, 2021: 122
Distribution: Guyana
Tobochares anthonyae Girón & Short, 2021: 125
Distribution: Venezuela (Bolívar)
Tobochares atures Girón & Short, 2021: 126
Distribution: Venezuela (Amazonas)
Tobochares benettii Girón & Short, 2021: 106
Distribution: Brazil (Amazonas)
Tobochares canaima Girón & Short, 2021: 128
Distribution: Venezuela (Bolívar)
Tobochares canaliculatus Kohlenberg & Short, 2017: 119
Distribution: Venezuela (Amazonas)
Tobochares canthus Kohlenberg & Short, 2017: 122
Distribution: Venezuela: Amazonas
Tobochares communis Girón & Short, 2021: 129
Distribution: Suriname
Tobochares emarginatus Kohlenberg & Short, 2017: 123
Distribution: Suriname
Tobochares fusus Girón & Short, 2021: 117
Distribution: Brazil (Amapá)
Tobochares goias Girón & Short, 2021: 109
Distribution: Brazil (Goiás)
Tobochares kappel Girón & Short, 2021: 133
Distribution: Suriname
Tobochares kasikasima Short, 2013: 83
Distribution: Suriname
Tobochares kolokoe Girón & Short, 2021: 134
Distribution: Suriname
Tobochares kusad Kohlenberg & Short, 2017: 126
Distribution: Guyana
Tobochares luteomargo Girón & Short, 2021: 115
Distribution: Venezuela (Bolívar)
Tobochares microps Girón & Short, 2021: 135
Distribution: Suriname
Tobochares pallidus Kohlenberg & Short, 2017: 130
Distribution: Venezuela (Amazonas, Bolívar)
Tobochares pemon Girón & Short, 2021: 136
Distribution: Venezuela (Bolívar)
Tobochares romanoae Girón & Short, 2021: 137
Distribution: Brazil (Roraima)
Tobochares sipaliwini Short & Kadosoe, 2011: 85
Distribution: Suriname, Guyana
Tobochares striatus Short, 2013: 83
Distribution: Suriname
Tobochares sulcatus Short & García, 2007: 4
Distribution: Venezuela: Amazonas, Bolívar
Girón, J.C., Short, A.E.Z. (2021) Review of the Neotropical water scavenger beetle genus Tobochares Short & García, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records. ZooKeys 1019: 93–140. https://doi.org/10.3897/zookeys.1019.59881
Kohlenberg, A.T. & Short, A.E.Z. 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short & García, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113–146. https://doi.org/10.3897/zookeys.669.11773
Short, A.E.Z. 2013. Chapter 4 – Aquatic Beetles of the Grensgebergte and Kasikasima Regions, Suriname (Insecta: Coleoptera). In: Alonso, L.E., Larsen, T.H. (Eds) A Rapid Biological Assessment of the Upper Palumeu River Watershed (Grensgebergte and Kasikasima) of Southeastern Suriname. RAP Bulletin of Biological Assessment, 67. Conservation International, Arlington, VA, 79–89. http://www.bioone.org/doi/full/10.1896/054.067.0112
Short, A.E.Z. & García, M. 2007. Tobochares sulcatus, a new genus and species of water scavenger beetle from Amazonas State, Venezuela (Coleoptera: Hydrophilidae). Aquatic Insects 29: 1–7. https://www.tandfonline.com/doi/abs/10.1080/01650420701247869
Short, A.E.Z. & Kadosoe, V. 2011. Chapter 4. Aquatic Beetles of the Kwamalasamutu Region, Suriname (Insecta: Coleoptera). In: O’Shea, B.J., Alonso, L.E. & Larsen, T.H., (Eds) A Rapid Biological Assessment of the Kwamalasamutu region, Southwestern Suriname. RAP Bulletin of Biological Assessment, 63. Conservation International, Arlington, VA, 79–90. http://www.bioone.org/doi/abs/10.1896/054.063.0107
Troglochares ashmolei Spangler, 1981: 318
Distribution: Ecuador: Morona-Santiago
Spangler, P. J., 1981. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington, 83(2): 316–323. https://www.biodiversitylibrary.org/item/55215#page/338/mode/1up