Nothofagus
Southern Beech
Southern Beech
Wikipedia links: Angiosperms > Eudicots > Rosids > Fabids > Fagales > Nothofagus
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Common name: Southern Beech
Conservation status: ...
Etymology:
Nothofagus - from the Greek nothos, meaning false and fagus, referring to the original belief that this genus was related to Northern Hemisphere Beech (Fagus) trees
Nothofagus means “false beech” although there is some suggestion that this is due to a misspelling of the intent NOTOfagus, meaning “Southern beech”
There is a suggestion that the genus was changed due to a spelling error
Flowers:
Fruit:
The fruit is a small, flattened or triangular nut
Borne in cupules containing 1 to 7 nuts
Leaves:
The leaves are toothed or entire, evergreen or deciduous
Stem & branches:
Roots:
Habit:
Trees and shrubs
Habitat:
The species are ecological dominants in many temperate forests
Distribution:
Native to the Southern Hemisphere:
southern South America (Chile, Argentina)
east and southeast Australia
New Zealand
New Guinea
New Caledonia
Species:
World: 43
Australia: 3
Additional notes:
Some species are reportedly naturalised in Germany and Great Britain
The genus has a rich fossil record of leaves, cupules, and pollen,
Fossils extendinto the late Cretaceous period and occurring in Australia, New Zealand, Antarctica, and South America
Reproduction
Many individual trees are extremely old, and at one time, some populations were thought to be unable to reproduce in present-day conditions where they were growing, except by suckering (clonal reproduction), being remnant forest from a cooler time
Sexual reproduction has since been shown to be possible
Taxonomy
The genus Nothofagus was first formally described in 1850 by Carl Ludwig Blume
He published the description in his book Museum botanicum Lugduno-Batavum, sive, Stirpium exoticarum novarum vel minus cognitarum ex vivis aut siccis brevis expositio et descriptio
In the past, they were included in the family Fagaceae, but genetic tests revealed them to be genetically distinct, and they are now included in their own family, Nothofagaceae
Species list
The following is a list of species, hybrids and varieties (44) of accepted by the Plants of the World Online as of April 2023:
Australia
Nothofagus cunninghamii (Tasmania, Victoria)
Nothofagus gunnii (Tasmania)
Nothofagus moorei (NSW, Queensland)
New Caledonnia
Nothofagus aequilateralis (New Caledonia)
Nothofagus balansae (New Caledonia)
Nothofagus baumanniae (New Caledonia)
Nothofagus codonandra (New Caledonia)
Nothofagus discoidea (New Caledonia)
New Guinea
Nothofagus brassii (New Guinea)
Nothofagus carrii (New Guinea)
Nothofagus crenata (New Guinea)
Nothofagus flaviramea (New Guinea)
Nothofagus grandis (New Guinea)
Nothofagus nuda (New Guinea)
Nothofagus perryi (New Guinea)
Nothofagus pseudoresinosa (New Guinea)
Nothofagus pullei (New Guinea)
Nothofagus resinosa (New Guinea)
Nothofagus rubra (New Guinea)
Nothofagus starkenborghiorum (Bismarck Archipelago, NG)
Nothofagus stylosa (New Guinea)
Nothofagus womersleyi (New Guinea)
New Zealand
Nothofagus cliffortioides (NZ North, NZ South)
Nothofagus fusca (NZ North, NZ South)
Nothofagus menziesii (NZ North, NZ South)
Nothofagus solandri (NZ North, NZSouth)
Nothofagus × apiculata (NZ North, NZ South)
Nothofagus × blairii (NZ North, NZ South)
Nothofagus × solfusca (New Zealand North)
Nothofagus truncata (NZ North, NZ South)
South America
Nothofagus alessandrii (Central Chile)
Nothofagus alpina (Argentina South, Chile)
CNothofagus antarctica (Argentina South, Chile)
Nothofagus betuloides (Argentina South, Chile South)
Nothofagus dombeyi (Argentina South, Chile)
Nothofagus glauca (Chile Central)
Nothofagus macrocarpa (Chile Central)
Nothofagus nitida (Chile South)
Nothofagus obliqua (Argentina South, Chile)
Nothofagus pumilio (Argentina South, Chile)
Nothofagus rutila (Chile Central)
Nothofagus × leoni (Chile Central)
Other
Nothofagus × dodecaphleps (artificial hybrid)
Nothofagus × eugenananus (artificial hybrid)
Subgenera
Four subgenera are recognized, based on morphology and DNA analysis:
Subgenus Fuscospora, six species (N. alessandri, N. cliffortioides, N. fusca, N. gunnii, N. solandri, and N. truncata) in New Zealand, Tasmania, and southern South America
Subgenus Lophozonia, seven species (N. alpina, N. cunninghamii, N. glauca, N. macrocarpa, N. menziesii, N. moorei, and N. obliqua) in New Zealand, Australia, and southern South America.
Subgenus Nothofagus, five species (N. antarctica, N. betuloides, N. dombeyi, N. nitida, and N. pumilio) in southern South America
Subgenus Brassospora (or Trisyngyne), 20 accepted species (N. aequilateralis, N. balansae, N. baumanniae, N. brassii, N. carrii, N. codonandra, N. crenata, N. discoidea, N. flaviramea, N. grandis, N. nuda, N. perryi, N. pseudoresinosa, N, pullei, N. recurva, N. resinosa, N. rubra, N. starkenborghiorum, N. stylosa, and N. womersleyi) in New Guinea and New Caledonia
In 2013, Peter Brian Heenan and Rob D. Smissen proposed splitting the genus into four, turning the four recognized subgenera into the new genera Fuscospora, Lophozonia and Trisyngyne, with the five South American species of subgenus Nothofagus remaining in genus Nothofagus
The proposed new genera are not accepted at the World Checklist of Selected Plant Families
Sources of information:
https://en.wikipedia.org/wiki/Nothofagus (Jan 2024)
Sources of information: