Monograph:

1. Härkönen, T. 1986. Guide to the otoliths of the bony fishes of the Northeast Atlantic. Danbiu Aps. Denmark. 256 pp.

Papers in scientific journals:

2. Härkönen, T. J. 1987. Seasonal and regional variations in the feeding habits of harbour seals Phoca vitulina in the Kattegat and the Skagerrak. J. Zool. Lond. 213: 535-543.

3. Härkönen, T. J. 1987. Influence of feeding on haul-out patterns and sizes of sub-populations in harbour seals. Neth. J. Sea Res. 21(4): 331-339.

4. Härkönen, T. J. 1987. Feeding ecology and population dynamics of the harbour seal (Phoca vitulina) in Kattegat-Skagerrak. Ph. D. Thesis. University of Göteborg.

5. Härkönen, T. J. 1988. Food-habitat relationship of harbour seals and black cormorants in Skagerrak and Kattegat. J. Zool. Lond. 214: 673-681.

6. Heide-Jørgensen, M.-P. and T. Härkönen. 1988. Rebuilding seal stocks in the Kattegat-Skagerrak. Marine Mammal Science. 4(3):231-246.

7. Dietz, R., M.-P. Heide-Jørgensen and T. Härkönen 1989. Mass deaths of harbour seals Phoca vitulina in Europe. Ambio 18(5): 258-264.

8. Härkönen, T. and M.-P. Heide-Jørgensen 1990. Density and distribution of the ringed seal in the Bothnian Bay. Holarctic Ecology 13 (2): 122-129.

9. Härkönen, T. and M.-P. Heide-Jørgensen 1990. Short-term effects of the mass dying of harbour seals in the Kattegat-Skagerrak area during 1988. Zeitschrift für Säugetierkunde 55: 233-238.

10. Härkönen, T. and M.-P. Heide-Jørgensen 1990. Comparative life histories of East Atlantic and other harbour seal populations. Ophelia 32 (3): 211-235.

11. Helle, E., O.Stenman, M. Olsson, B. Helander and T. Härkönen. 1990. Baltic Seals. Population size, reproduction and organochlorines. Ambio special report 7:16.

12. Bengtson, J., P. Boveng, U. Franzén, P. Have, M.-P. Heide-Jørgensen and T. Härkönen 1991. Antibodies to canine distemper virus in Antarctic seals. Marine Mammal Science 7(1): 85-87.

13. Dietz, R., M.-P. Heide-Jørgensen, T. Härkönen, J. Teilmann and N. Valentin. 1991. Age determination of European harbour seals. Sarsia 76: 17-21.

14. Härkönen, T. and M.-P. Heide-Jørgensen 1991. The harbour seal Phoca vitulina as a predator in the Skagerrak. Ophelia 34(3):191-207.

15. Heide-Jørgensen, M.-P. and T. Härkönen 1992. Epizootiology of seal disease. J. Appl. Ecol. 29: 99-107.

16. Heide-Jørgensen, M.-P., T. Härkönen, R. Dietz and P. Thompson 1992. Retrospective of the 1988 European seal epizootic. Diseases of Aquatic Organisms. 13: 37-62.

17. Heide-Jørgensen, M.-P., T. Härkönen and P. Åberg. 1992. Long-term effects of the seal epizootic. Ambio 21: 511-516.

18. Härkönen, T and S. G. Lunneryd 1992. Estimating abundance of ringed seals in the Bothnian Bay. Ambio 21:497-510

19. Mortensen, P., Bergman, A., Bignert, A., Hansen, H.J., Härkönen, T., Olsson, M. 1992. Prevalence of skull lesions in harbour seals Phoca vitulina in Swedish and Danish museum collections during the period 1835-1988. Ambio 21:520-524.

20. Roos, A., G. Blomkvist, S. Jensen, Mats Olsson, A. Bergman and T. Härkönen. 1992.Sample selection and preparation procedures for analyses of Metals and Organohalogen compounds in Swedish seals. Ambio 21:525-528.

21. Hårding K.C. and T. Härkönen. 1995. Estimating age at sexual maturity in crabeater seals Lobodon carcinophagus. Can. J. Fish. Aquat. Sci. 52: 2347-2352.

22. Harding, K.C. and T.J. Härkönen 1999. Development in the Baltic grey seal (Halichoerus grypus) and ringed seal (Phoca hispida) populations during the 20th century. Ambio. 28: 619-627.

23. Härkönen, T, K.C. Harding and S.G. Lunneryd 1999. Age and sex specific behaviour in harbour seals leads to biased estimates of vital population parameters. J. Appl. Ecol. 36: 824-840.

24. Härkönen, T, and K.C. Harding. (2001). Spatial structure of harbour seal populations and the implications thereof. Can. J. Zool. 79: 2115-2127.

25. Härkönen, T., K.C Harding, and M.-P. Heide-Jørgensen (2002). Rates of increase in age structured populations: A lesson from the European harbour seals. Can. J. Zool. (in press).

26. Harding, K.C. Härkönen, T. and H, Caswell. The 2002 European seal plague: Epidemiology and population consequences. Ecology Letters (In press).

32. Harding K.C., Härkönen, T. and J. Pineda. (2003). Estimating quasi-extinction risk of European harbour seals.: a reply to Lonergan and Harwood. Ecology Letters 6: 894-897.

33. Härkönen, T. 2003. Development of populations of harbour seals and grey seals in the Wadden Sea and the North Sea since 1988. Wadden Sea Ecosystems, 17: 13-18.

34. Harding, K., M. Fujiwara, Y. Axberg and T. Härkönen (2005). Mass dependent energetics and survival in harbour seal pups. Functional Ecology, 19: 129-135.

35. Härkönen, T, K.C. Harding, S. Goodman, and K. Johannesson (2005) Colonization history of the Baltic harbor seals: Integrating archaeological, behavioural and genetic data. Marine Mammal Science 21: 695-716.

36. Härkönen, T., R. Dietz, P. Reijnders, J. Teilmann, K. Harding, A. Hall, S. Brasseur, U. Siebert, S. Goodman, P. Jepson, T. Dau Rasmussen, P. Thompson (2006). A review of the 1988 and 2002 phocine distemper virus epidemics in European harbour seals. Diseases of Aquatic Organisms, 68: 115-130.

37. Hall, A., P. Jepson, S. Goodman and T. Härkönen (2006). Phocine Distemper Virus – Data and models, nature and nurture. Biological Conservation. 131: 221-229.

38. Harding, K.C. Härkönen, T., B. Helander and O. Karlsson (2007). Status of Baltic grey seals: Population assessment and risk analysis. NAMMCO Scientific Publications, 6: 33-56.

39. Härkönen, T., S. Brasseur, J. Teilmann, C. Vincent, R. Dietz, P. Reijnders, K. Abt (2007) Status of grey seals along mainland Europe, from the Baltic to France. NAMMCO Scientific Publications, 6: 57-68.

40. Leidenberger, S., K. C. Harding and T. Härkönen (2007). Phocid seals, seal lice and heartworms: a terrestrial host-parasite system conveyed to the marine environment. Diseases of Aquatic Organisms, 77: 235-253.

41. Kreutzer, M. , Kreutzer, R. , Siebert, U . Müller G., Reijnders, P., Brasseur, S., Härkönen, T., Dietz, R., Sonne, C., Born, E.W. & Baumgärtner, W. (2007). In search for virus carriers of the 1988 and 2002 phocine distemper virus outbreaks in European harbour seals. Archives of Virology, DOI 10.1007/s00705-007-1070-6.

42. Härkönen, T., Harding, K., Rasmussen, T.D., Teilmann, J., Dietz, R. (2007). Age- and Sex-specific Mortality Patterns in an Emerging Wildlife Epidemic: the Phocine Distemper in European Harbour Seals. PLoS ONE, 2(9): e887. doi: 10.1371/journal.pone.0000887

43. Jüssi, M., Härkönen, T., Jüssi, I. Helle, E. (2008). Decreasing ice coverage will reduce the reproductive success of Baltic grey seal (Halichoerus grypus) females. Ambio, 37: 80–85.

44. Harkonen, T., M. Jüssi, M. Baimukanov, A. Bignert, L. Dmitrieva, Y. Kasimbekov, M. Verevkin, S. Wilson & S. J. Goodman (2008). Pup production and breeding distribution of the Caspian seal (Phoca caspica) in relation to human impacts. Ambio 37, 356-361.

45. Harkonen, T., Bäcklin, B-M., Barrett, T., Anders Bergman, A., Corteyn, M., Dietz, R., Harding, K., Malmsten, J., Roos, A., Teilmann, T. (2008). Mass mortality in harbour seals and harbour porpoises caused by an unknown pathogen. The Veterinary Record, 162: 555-556.

46. Harkonen, T., M. Jüssi, I. Jüssi, M. Verevkin, L. Dmitrieva, E. Helle, R. Sagitov, K.C. Harding. (2008). Seasonal activity budget of adult Baltic ringed seals (Phoca hispida botnica). PLoS ONE 3(4): e2006.

doi:10.1371/journal.pone.0002006

47 Teilmann, J., F. Riget, T. Harkonen. 2010. Optimising survey design in Scandinavian harbour seals: Population trend as an ecological quality element. ICES Journal of Marine Science, 67: 952–958

48. Anna Arcalís-Planas, Signe Sveegaard, Olle Karlsson, Karin C Harding, Anna Wåhlin, Tero Harkonen, Jonas Teilmann (2010Limited use of sea ice by the Ross seal (Ommatophoca rossii), in Amundsen Sea, Antarctica, using telemetry and remote sensing data.Polar Biology 38 (4), 445-461

49. Lockyer, C., Mackay, B., Härkönen, T., Read, F. and Hasselmeier, I. 2011. Age Determination in Harbour Seals. . NAMMCO Sci. Publ. 8: 245-264

50. Nilssen, K.T., Skavberg, N.E., Poltermann, M., Haug, T., Härkönen, T. and Henriksen, G. 2011. Status of harbour seals (Phoca vitulina) in Norway. NAMMCO Sci. Publ. 8: 61-70.

51. Olsen, M. T., L, Andersen, S.M., Teilmann, J. Dietz, R., Harkonen, T. 2011. Status of the harbour seal in Southern Scandinavia. NAMMCO Sci. Publ. 8: 77-94

52. Harkonen, T., Harding, K.C. 2011. Predicting recurrent PDV epidemics in European harbour seals. NAMMCO Sci. Publ. 8: 275-284

53. Harkonen, T. and Isakson, E. 2011. Historical and current status of harbour seals in the Baltic proper. NAMMCO Sci. Publ. 8: 71-76

54. Svensson, C.J., Hansson, A. Harkonen, T. Harding, K .2011. Detecting density dependence in growing seal populations. AMBIO (2011) 40:52–59, DOI 10.1007/s13280-010-0091-7

55. Kovacs, K.M., Aguilar, A., Aurioles, D., Burkanov, V., Campagna, C., Gales, N., Gelatt, T., Goldsworthy, S., Goodman, S, Hofmeyr, G., Härkönen, T., Lowry, L., Lydersen, C., Schipper, J., Sipilä, T., Southwell, C., Stuart, S., Thompson, D., Trillmich, F. 2011. Global threats to Pinnipeds. Marine Mammal Science. DOI: 10.1111/j.1748-7692.2011.00479.x

56. Aubail A., Teilmann J., Dietz R., Rigét F., Harkonen T., Karlsson O., Caurant, F. 2011. Investigation of mercury concentrations in fur of phocid seals using stable isotopes as tracers of trophic levels and geographical regions. Polar Biology. 34:1411-1420, DOI: 10.1007/s00300-011-0996-z

57. Sundqvist, L., T Harkonen, C J Svensson, K C. Harding. 2012. Linking climate trends to population dynamics in the Baltic ringed seal - Impacts of historical and future winter temperatures. Ambio. DOI 10.1007/s13280-012-0334-x

58. Harkonen, T., K. Harding, S. Wilson, C. J. Svensson, L. Dmitrieva, S. J. Goodman. 2012. Collapse of a marine mammal species driven by human impacts. PlosOne. PLOS ONE PONE-D-12-07084R1 10.1371/journal.pone.0043130

59. Olsen, M. T., L. Wesley Andersen, R. Dietz, J. Teilmann, T. Harkonen and H. R. Siegismund 2014. 'Integrating genetic data and population viability analyses for the identification of harbour seal (Phoca vitulina) populations and management units. Molecular Ecology. 23: 815-831.

60. Arcalís-Planas, A., J. Teilmann, S. Sveegaard, O. Karlsson, K. C. Harding, A. Wåhlin, A and T. Harkonen 2014. Limited use of sea ice by the Ross seal (Ommatophoca rossii), in Amundsen Sea, Antarctica, using telemetry and remote sensing data. Polar Biology. DOI: 10.1007/s00300-014-1602-y

61. Zohari S, Neimanis A, Härkönen T, Moraeus C, Valarcher JF. Avian influenza A(H10N7) virus involvement in mass mortality of harbour seals (Phoca vitulina) in Sweden, March through October 2014. Euro Surveill. 2014;19(46):pii=20967. Available online: http://www.eurosurveillance.org/ViewArticle.aspx?ArticleId=20967

62. Dmitrieva, L., Härkönen, T., Baimukanov,M., Anders Bignert, A., Jüssi, I., Jüssi, M., Kasimbekov, Y., Verevkin, M, VysotskiyV., Wilson, S., Goodman, S.J.. 2015. Inter-year variation in pup production of Caspian

seals Pusa caspica 2005–2012 determined from aerial surveys. Endangered Species Research 28: 209–223

63. Bodewes, R., Zohari, S., Krog, JS., Hall, M.B., Harder, T.C., Bestebroer, T.M., van de Bildt, MWG, Spronken, M.I., Larsen, L.E., Siebert, U., Wohlsein, P., Puff, C., Seehusen, F., Baumgärtner, W., Härkönen, T., Smits, S.L., Herfst, S., Osterhaus, ADME., Fouchier, RAM., Koopmans, M.P., Kuiken, T. 2016. Spatiotemporal analysis of the genetic diversity of seal influenza A (H10N7) virus, Northwestern Europe. Journal of virology. 96: 4269-4277.

64. K.Lehnerta, L.Weirupa, KC Harding, T. Härkönen, O. Karlsson, J. Teilmann 2017. Antarctic seals: Molecular biomarkers as indicators for pollutant exposure, health effects and diet. Sci. Tot. Env.1693-1704

65. Hansson S., Bergström U., Bonsdorff E., Härkönen T., Jepsen N., Kautsky L., Lundström K., Lunneryd SG., Ovegård M., Salmi J., Sendek D., Vetemaa V. 2017. Competition for the fish - fish extraction from the Baltic Sea by humans, aquatic mammals and birds. ICES J. Marine Sci. . doi:10.1093/icesjms/fsx207

66. Olsen. MT., Galatius A., Härkönen T. (2018). The history and effects of seal-fisheries conflicts in Denmark. Marine Ecology Progress Series. 595: 233–243, 2018

67. Sonne C., Andersen-Ranberg E., Rajala EL., Agerholm JS., Bonefeldt-Jörgensen E. Desforges JP., Eulers I., Jenssen BM., Koch A. Rosing-Asvid A., Siebert U., Tryland M., Mulvad G., Härkönen T., Dietz R., Magnusson U. (2018). Seroprevalence for Brucella spp. in Baltic ringed seals (Phoca hispida), East Greenland harp (Pagophilus groenlandicus) and hooded seals (Cystophora cristata). Veterinary Immunology and Immunopathology

Volume 198, April 2018, Pages 14-18

68. Harding, K.C., Salmon, M., Teilmann, J., Dietz, R & T. Harkonen. 2018. Population Wide Decline in Somatic Growth in Harbor Seals—Early Signs of Density Dependence. Frontiers in ecology and evolution. doi: 10.3389/fevo.2018.00059

69. EK Brault, PL Koch, DP Costa, MD McCarthy, LA Hückstädt, KT Goetz, ... (2019), Trophic position and foraging ecology of Ross, Weddell, and crabeater seals revealed by compound-specific isotope analysis.

Marine Ecology Progress Series 611, 1-18

70. Stokholm, I. Tero Härkönen, Karin C Harding, Ursula Siebert, Kristina Lehnert, Rune Dietz, Jonas Teilmann, Anders Galatius, Linnea Worsøe Havmøller, Emma L Carroll, Ailsa Hall, Morten Tange Olsen (2019). Phylogenomic insights to the origin and spread of phocine distemper virus in European harbour seals in 1988 and 2002. Diseases of aquatic organisms. 133:47-56

71. Cervin, L., Harkonen, T., Harding. K. (2020). Multiple stressors and data deficient populations; a comparative life-history approach sheds new light on the extinction risk of the highly vulnerable Baltic harbour porpoises (Phocoena phocoena). Environment International. 144, 106076

72. Harbour seal

MANY MORE TO ADD HERE, under construction

Other international publications, reports etc. (recent)

Härkönen, T., M. Jüssi, K. Harding, I. Jüssi.& E. Helle. 2000. Distribution and diving behaviour of satellite tagged ringed seals (Phoca hispida botnica) in Estonian coastal waters of the Baltic Sea. ICES WGMMHA Helsinki 28 Feb- 3 March 2000 WP 9. 17 pp.

Härkönen, T., K.C. Harding & M.P. Heide-Jørgensen 2000. Impact of initial conditions on estimates of growth rates: recovery of European harbor seals after the 1988 epizootic. ICES WGMMHA Helsinki 28 Feb- 3 March 2000.WP5. 43 pp.

Härkönen, T., M. Olsson & K. Harding. 2000. Population effects of decreased fertility rates in Baltic ringed seals after 1960. ICES WGMMHA Helsinki 28 Feb- 3 March 2000.WP8. 7 pp.

Härkönen, T. & K.C. Harding. 2000. History of colonisation, past and recent trends in Baltic seal populations . ICES WGMMHA Helsinki 28 Feb- 3 March 2000. WP7. 14 pp.

Härkönen, T., K.C. Harding & M.P. Heide-Jørgensen 1999. Recovery of seal stocks in the Kattegat-Skagerrak and the Limfjord after the seal epizootic. ICES WGMMHA Copenhagen 8-12 March 1999. Working Paper 15 -99. 10 pp.

Härkönen T. and K.C. Harding 1999. Why is the East Baltic harbour seal (Phoca vitulina) genetically divergent from all other harbour seals? In: Habitat Loss: Ecological, Evolutionary and Genetic Consequences. Helsinki 7-12 September 1999. p. 126.

Harding K.C. and T. Härkönen 1999. Developments of the Baltic grey seal (Halichoerus grypus) and ringed seal (Phoca hispida) populations during the 20th century. In European Mammology 3rd European Congress of Mammology May 29-June 3 1999, Jyväskylä, Finland. Abstract p. 120.

Härkönen, T. & K. Harding 1999. Population development and reproduction of the Baltic harbour seal. Abstract. Symposium of Baltic Seals in Pärnu, Estonia Nov. 1999.

Härkönen, T., M. Jüssi, K. Harding & I. Jüssi.1999. Seasonal changes in diving behaviour of ringed seals in the Baltic. Abstract. Symposium of Baltic Seals in Pärnu, Estonia Nov. 1999.

Härkönen, T. & K. Harding . 1999. Long-term studies of freeze branded seals give detailed insight into seal biology. Abstract. Symposium of Baltic Seals in Pärnu, Estonia Nov. 1999.

Härkönen, T., M. Jüssi, K. Harding, I. Jüssi and Eero Helle. 1999. Seasonal activity patterns and area use of Baltic ringed seals (Phoca hispida botnica). ICES WGMMHA Copenhagen 8-12 March 1999. Working Paper 9 -99. 15 pp.

Harding, K and Tero Härkönen. 1999. Site fidelity, a mechanism causing spatially structured populations of harbour seals (Phoca vitulina). ICES WGMMHA Copenhagen 8-12 March 1999. Working Paper 8 -99. 15 pp.

Härkönen, Harding POSTER Biennal Congress ofMammalogy. Vancouver Canada. Nov. 2001

Härkönen, T. 2002. Faktablad: Phoca vitulina-knubbsäl. Artdatabanken.

Härkönen, T. 2002. Faktablad: Phoca hispida hispida, vikare. Artdatabanken.

Harding, K.C. Härkönen, T. och B. Helander 2003. Populationsstatus och ekologisk riskanalys för gråsälen i Östersjön. Naturvårdsverket. 30 pp.

Härkönen T. 2003. Redovisning av insatserna vid säldöden 2002.Naturvårdsverket. 11pp.

Härkönen T. 2003. Vikaren – en ishavsrelikt. Havsutsikt 2003

Härkönen T. et al. 2003. Status of the Baltic ringed seal. ICES WGMME, Hel, Poland. 10pp

Härkönen, T. and K.C. Harding 2003. Historical and present status of the Baltic harbour seal. ICES WGMME, Hel, Poland. 10pp

Härkönen, T. 2003. Massmortalitet hos knubbsäl. Havsmiljön 2003.

Härkönen, T. 2004. Åtgärdsprogram för vikaresäl. Naturvårdsverket. 18 pp,

Karlsson, O., Härkönen, T. & Bäcklin, B.-M. 2007. Sälar på uppgång. Havet

2007, pp 84-89.

An effective survey design for harbour seals in the Wadden Sea: tuning Trilateral Seal Agreement and EU-Habitat Directive requirement

E Meesters, PJH Reijnders, S Brasseur, S Tougaard, M Stede, U Siebert, ...

The meeting of the Trilateral Working Group TWG 7 (1)

Sälpopulationer och sälhälsa

T Härkönen, O Karlsson, BM Bäcklin, C Moraeus

Havet 2011, 95-96

Sälpopulationer och sälhälsa

T Härkönen, O Karlsson, BM Bäcklin, C Moraeus

Havet 2011, 95-96

Guidelines for seal abundance monitoring in the HELCOM area

A Galatius, M Ahola, T Härkönen, I Jüssi, M Jüssi, O Karlsson, M Verevkin

HELCOM, 1-8

Book chapters in edited books

Härkönen, T., O. Stenman, M. Jüssi, I. Jüssi, R. Sagitov, M. Verevkin. 1998. Population size and distribution of the Baltic ringed seal (Phoca hispida botnica). In: Ringed Seals (Phoca hispida) in the North Atlantic. Edited by C.Lydersen and M.P. Heide-Jørgensen. NAMMCO Scientific Publications, Vol. 1, 167-180.

Kapel, F.O., Christiansen, J., Heide-Jørgensen, M.P., Härkönen, T., Born, E.W., Knutsen, L.Ø., Riget, F. and Teilman, J. 1998. Netting and conventional tagging used for studying movements of ringed seals (Phoca hispida) in Greenland. In: Ringed Seals (Phoca hispida) in the North Atlantic. Edited by C.Lydersen and M.P. Heide-Jørgensen. NAMMCO Scientific Publications, Vol. 1:211-228.

Siegstad, H., Neve, P.B., Heide-Jørgensen, M.P.and Härkönen, T. 1998. Diet of the ringed seal in Greenland. In: Ringed Seals (Phoca hispida) in the North Atlantic. Edited by C. Lydersen and M.P. Heide-Jørgensen. NAMMCO Scientific Publications, Vol. 1, 229-241.

Bengtson, J.L., L.M. Ferm, T. J, Härkönen and B. S. Stewart. 1990 Abundance of Antarctic Fur seals in the South Shetland Islands, Antarctica, During the 1986/87 Austral summer. In: Antarctic Ecosystems. Editors KR Kerry and G. Hempel. 265-270.

Examples of reports etc. in Swedish (this is not the full list)

Härkönen T.J. & K.C. Harding 2001. Sälbestånden i Östersjön: Vad kan vi lära oss av historien för att lösa dagens problem? Kungliga Skogs- och Lantbruksakademiens Tidskrift. 140:1, 9-36.

Harding K & T. Härkönen 1998. Gråsäl och vikare i Östersjön, en rekonstruktion av populationernas utveckling under 1900-talet. Slutrapport till Projekt Sälar och Fiske. 36 pp.

Härkönen, T. & K. Harding 1996. Sälbestånden hämtar sig. Havsmiljön, Aktuell rapport om miljötillståndet i Kattegatt, Skagerrak och Öresund. Mars 1996. p 18.

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