2000   M.D., School of Medicine, Tohoku University

2000   Resident, Internal Medicine, Yamagata City Hospital Saiseikan

2007   Ph. D., Graduate School of Medicine, Kyoto University (Advisor: Prof. Shigetada Nakanishi)

2008   Postdoctoral fellow → Research Associate, Department of Anatomy ＆ Neurobiology at the University of Maryland  (PI: Geoffrey Schoenbaum) 

2010   Postdoctoral Associate,  MIT Media Lab Synthetic Neurobiology Group (PI: Edward Boyden) 

2013    Project-oriented Assistant Professor, National Institute for Physiological Sciences (PI: Tadashi Isa) 

2016    Assistant Professor, Graduate School of Medicine, Kyoto University (PI: Tadashi Isa) 

2017   Senior Lecturer, Graduate School of Medicine, Kyoto University (PI: Tadashi Isa) 

2018    Project-oriented Associate Professor (PI, group leader), Medical Innovation Center, Graduate School of Medicine, Kyoto University 

2021    Kyoto University FOREST Principal Investigator (supported by FOREST program)

2023    Project-oriented Associate Professor & FOREST PI, Graduate School of Medicine, Kyoto University 

2023    Professor, Department of Systems Physiology, Shiga University of Medical Science