2000   M.D., School of Medicine, Tohoku University

2000   Resident, Internal Medicine, Yamagata City Hospital Saiseikan

2007   Ph. D., Graduate School of Medicine, Kyoto University (Advisor: Prof. Shigetada Nakanishi)

2008   Postdoctoral fellow → Research Associate, Department of Anatomy ＆ Neurobiology at the University of Maryland  (PI: Geoffrey Schoenbaum) 

2010   Postdoctoral Associate, Massachusetts Institute of Technology, Media Lab, Synthetic Neurobiology Group (PI: Edward Boyden) 

2013    Project-oriented Assistant Professor, National Institute for Physiological Sciences (PI: Tadashi Isa) 

2016    Assistant Professor, Graduate School of Medicine, Kyoto University (PI: Tadashi Isa) 

2017   Senior Lecturer, Graduate School of Medicine, Kyoto University (PI: Tadashi Isa) 

2018    Project-oriented Associate Professor (PI, group leader), Medical Innovation Center, Graduate School of Medicine, Kyoto University 

2021    Kyoto University FOREST Principal Investigator (supported by FOREST program)

2023    Professor, Department of Systems Physiology, Shiga University of Medical Science