Overview of Study

After a long-distance migration from the East, Avar people of Eastern Asian ancestry arrived in Eastern Central Europe in 567/568 CE and encountered groups with very different, European ancestry. We used ancient genome-wide data of 722 individuals and fine-grained interdisciplinary analysis of large 7th-8th c. CE neighboring cemeteries south of Vienna (Austria) to address the centuries-long impact of this encounter. We found that the ancestry at one site (Leobersdorf) was and remained dominantly East-Asian-like, even 200 years after the Avar immigration, while the other site (Mödling) displays local, European-like ancestry. These two nearby sites exhibit very few links of direct biological relatedness, despite sharing a distinctive late-Avar culture. We reconstructed 6-generation pedigrees at both sites linking together up to 450 closely-related individuals, allowing per-generation demographic profiling of the communities. Despite different ancestry, these pedigrees together with large networks of distant relatedness display an absence of consanguinity, a strong patrilineal pattern with female exogamy, multiple reproductive partnerships (e.g. levirate) and direct correlation of social status to biological connectivity through markers of high social status in the archaeological material. The generation-long genetic barrier was maintained by systematically choosing partners with similar ancestry from other sites in the Avar realm. Leobersdorf had more biological connections with the Avar heartlands than Mödling, which is instead linked to another site from the Vienna Basin with European-like ancestry. Mobility between sites was mostly due to female exogamy pointing to different marriage networks as the main driver of the maintenance of the genetic barrier.

At three archaeological sites from the seventh and eighth centuries ce at the western periphery of the Avar empire in the Vienna Basin, we revealed the strong contrast between very diverse genetic ancestry, a shared cultural habitus and similar social structure. Whereas Leobersdorf displays East Asian ancestry with minor Pontic steppe and European admixture that occurred within the last 200 years, individuals at Mödling-An der Goldenen Stiege showed a mixture of European-like ancestries that were acquired several times in the past, with only minor elements of more recent East Asian admixture. 

https://www.ebi.ac.uk/ena/browser/view/PRJEB76548

Supplemental Information

Mödling-An der Goldenen Stiege Cemetery
About 18 km north of Leobersdorf, the cemetery of Mödling-An der Goldenen Stiege is situated on a terrace on the eastern slope of the Vienna Woods. A total of 488 graves with 536 burials of the 7th to 9th centuries were excavated between 1967 and 197324–29. The burial site was built on the remains of a Neolithic settlement and a Hallstatt period cremation burial ground and was surrounded by an undated rampart30. About 900 m away and 20 m deeper in the flatland of the Vienna Basin, evidence of earlier burials from the end of the 4th (Mödling-Lerchengasse) and the 6th centuries (Mödling-Leinerinnen) was found31,32.

Excerpt from Vienna Making of a Capital

MIlitary Camp
Like many other cities of Continental Europe, Vienna originated in ancient Roman times. In the first century AD, the Romans set up a military camp, called Vindobona, which formed part of the large number of similar facilities along the Limes frontier. The camp was situated in what is today the core of the city. The course of the wall is reflected, to this day, in a series of very striking streets in the inner city. From the third century, there is evidence for a civilian settlement in the southeast, a little outside the camp. Much less is known about its layout and buildings, however, than about the camp. Neither of these facilities on the site of the present city had any significance within the ancient "cityscape" of Austria. The late 4th and 5th centuries began to see the dramatic decline not only of the Roman Empire in general but also of Vindobona in particular. Archaeological evidence provides information of a devastating fire in the storage area at the beginning of the 5th century.

No written records have survived from the period between the 5th to 6th centuries and the 9th century to tell about the fate of this small settlement on the Danube. But research on the basis of archaeological and onomastic evidence suggests that we must assume that at least some "remainder settlement" continued to exist. The protection provided by the remnants of the ancient camp fortifications, which continued well into the 13th century, was made further use of. It may have been as early as the Carolingian period that the first churches were established in the area. Of course, it would be impossible to speak of a city or even urban structures, particularly in the light of the fact that the settlement, whose present name was first mentioned in 881, was to see turbulent times in the wake of the Magyar expansion, in the 9th and 10th centuries.

Early Middle Ages
It was only after the Magyar threat had been contained, following the Battle of the Lechfeld in 955, and the establishment of the margravate of Austria under the Babenbergs (976) had brought forth a "quasi-state" that the basis for a general increase in settlement in the fledgling country was laid. The effects did not become fully palpable for Vienna. The city continued to sit virtually on the Magyar frontier well into the 12th century, until the Austrian sovereigns from the Babenberg dynasty had finally consolidated their position. Until the first half of the 12th century, the centre of Babenberg power lay west of Vienna. Around 1150 the Austrian margraves, who were also dukes of Bavaria at the time, transferred their residence to Vienna. They founded Vienna's oldest monastery, Sankt Maria zu den Schotten (St. Mary of the Scots). This (now) direct contact with the sovereigns contributed strongly to Vienna's unfolding into a veritable town. It is interesting to underline that Vienna was not "founded" by a lord, but that, on the other hand, the role of the sovereign was a significant one. The shining light emulated was most of all the old Bavarian capital of Ratisbon, which Vienna outdid, however, within a few decades.

High Middle Ages
The outward sign of any medieval city was its surrounding wall. Vienna's city wall was built around 1200. The community was granted its first privilege in 1221. Vienna was termed one of the principal cities of the empire after Cologne. At that time, in the early 13th century, Vienna was already part and parcel of an extensive network of trade relations. This network made use of the Danube route and benefited from intensive contacts with Venice. A major upgrading came with the granting of the so-called "Stapelrecht". Under this privilege non-resident tradesmen had to display their goods for sale in the city. This afforded the Viennese the very profitable opportunity to act as middlemen. The city's expansion and development is particularly reflected in the increasing splendour and number of churches, although the foundations thereto had been laid by the Babenbergs still. There was growing construction at the time even in the suburban area, outside the walls, and development crystallised along the main arteries leading into and out of the city.

The late medieval Period
The late Middle Ages deserve to be called a heyday of burgher autonomy although conflicts with the sovereigns - to wit the Habsburgs from 1276/83 - recurred again and again. It is interesting to note in this context that the Habsburgs lost the imperial crown upon the death of Frederick the Fair (1330) for more than a century. They did not regain it until 1438, when Albert II came to power. In the 14th century, the cityscape was refashioned in Gothic style. This appearance was to prevail until the early 17th century. The most obvious example is doubtless the redesign of the parish church of St. Stephen's, which did not become a bishopric until 1469. The intellectual appeal of the city, which was largely dominated by the Habsburg court, was dramatically enriched with the establishment of the University of Vienna in the 14th century still.

In the late Middle Ages political and social tensions among the burghers of Vienna often mirrored dynastic quarrelling within the House of Habsburg. Nevertheless the city was spared the kinds of constitutional conflicts known from many other European cities. On the contrary, 1396 saw the number represented within the city government of various groups of burghers, especially the so-called "Erbbürger", or patricians, as well as tradesmen and craftsmen set down in a well-regulated way.

Trade
The economic mainstay of the city was, on the one hand, trade, and, on the other hand, wine-growing. Wine was the principal export, and this considerable wine production was protected by import bans on foreign, particularly Hungarian, wines. With this in mind, it is hardly surprising that the political unrest of the 15th century, beginning with the Hussite Wars and including the strife between Emperor Frederick III and his brother, Archduke Albert VI, and the Hungarian occupation of Vienna under King Matthias I Corvinus (1485 to 1490), affected these very pillars of Vienna's economy drastically. Especially the "Stapelrecht" was becoming more and more ineffective. The pressure exerted by foreign tradesmen who had been doing business in the city for a long time as well as the growing dependence of the Habsburgs on the financial prowess of powerful trade dynasties such as the Fuggers eventually prompted Emperor Maximilian to repeal the privilege in the early 16th century.

MGS Locality G25 Set

Avar:MGS001,0.125205,0.113739,0.013953,-0.017119,0.01908,-0.008367,-0.002115,0.001846,0,0.007107,0.004872,0.001049,-0.001933,-0.003165,-0.002172,0.005436,0.001565,0.003801,-0.001383,-0.001876,-0.001872,-0.003462,-0.002465,0,0.000718

Avar:MGS002,0.12862,0.132019,0.028284,0.009044,0.028313,-0.001673,0.007285,0.004846,-0.001636,0.00656,0.00341,0.009741,-0.018731,-0.006193,-0.005565,0.003978,0.012908,-0.000507,0.00176,-0.006003,-0.002496,-0.001978,0.001602,0.00012,-0.00946

Avar:MGS003,0.136588,0.138112,0.033564,0.007429,0.034468,0.002231,-0.00846,0.002769,0.005727,0.013303,0.000974,0.008842,0.000595,0.008808,-0.009908,-0.011535,-0.011604,-0.000253,0.001885,-0.004502,-0.003743,-0.002968,0.002342,0.007832,-0.001557

Avar:MGS005,0.122929,0.1046,0.037712,0.000646,0.02462,0.006972,-0.009635,0.006231,0.005318,0.012939,0.001461,-0.001049,0.001189,0.001789,-0.008686,-0.002519,-0.00326,0.001774,0.004022,-0.009379,0.006863,0.008532,0.012202,-0.010242,-0.002994

Avar:MGS005.A0201,0.118376,0.105615,0.030547,0.017442,0.010156,0.011713,0.00564,0.011999,0.009204,0.014579,0.005521,0.013788,0.008028,-0.002615,-0.008822,-0.005569,-0.010431,-0.001267,-0.008673,0.000875,0.003743,-0.005564,0.001725,-0.007712,0.007664

Avar:MGS006,0.132035,0.12491,0.041483,0.005168,0.036622,0.002789,0.00564,0.007615,0,-0.010205,0.003085,-0.002248,0.003271,0.011973,-0.014522,0.005171,0.012778,-0.007348,0.008673,0.001751,-0.007237,0.00643,0.010846,0.004097,-0.002036

Avar:MGS007,0.118376,0.121864,0.015462,-0.015827,0.018465,-0.008646,0.00141,0.001846,0.003886,0.022962,0.003735,0.005845,-0.011447,-0.006881,-0.003393,-0.002387,0.009648,-0.005068,0.012067,-0.012381,-0.006863,0.003215,-0.008258,0.008676,-0.00012

Avar:MGS008,0.135449,0.136081,0.039598,0.025194,0.038161,0.015618,0.00235,0.008769,0.000614,-0.00656,-0.007145,-0.010041,0.011447,0.023396,-0.018051,-0.002784,0.008214,0.006968,0.004777,-0.005753,-0.009109,-0.002473,-0.00037,-0.011929,0.001916

Avar:MGS009,0.122929,0.14319,0.048271,0.024548,0.036622,0.016176,0.000235,-0.001385,-0.001227,0.007472,-0.001137,-0.007493,0.009514,0.011423,-0.014794,-0.010872,0.001565,-0.003167,0.001131,-0.006003,-0.011854,-0.005317,0.002958,-0.005543,-0.002994

Avar:MGS010,0.119514,0.1046,0.037712,-0.00323,0.018465,-0.002231,0.011986,0.000923,-0.006136,0.004738,0.000974,0.000899,-0.011447,0.00289,-0.005429,-0.015911,-0.013038,0.003167,0.009176,-0.005127,0,-0.004451,-0.001602,0.006025,-0.002994

Avar:MGS011,0.133173,0.127957,0.060339,0.034561,0.03416,0.005857,0.00282,0.004615,0.011658,-0.012574,-0.007632,-0.004046,0.010258,0.020368,-0.021308,-0.008618,0.011213,-0.004941,-0.00264,0.002876,-0.006738,0.013849,0.005546,-0.011206,0.005149

Avar:MGS012,0.114961,0.085304,0.018102,-0.029393,0.010463,-0.008088,0.005875,0.003231,-0.003477,0.021139,0.004547,0.005995,-0.012042,-0.002752,-0.013843,-0.005967,0.008866,0.00076,0.002011,-0.004627,-0.005116,-0.002226,-0.007641,0.005904,-0.000838

Avar:MGS013,0.124067,0.148267,0.028284,0,0.025235,0.000279,0.00423,0.005077,0.011044,0.013668,-0.002111,0.000899,0.003419,0.001376,-0.008822,0.009679,0.024773,-0.004307,0.004274,-0.005378,-0.005865,-0.004822,0.000739,-0.003374,0.002275

Avar:MGS014,0.130897,0.132019,0.03017,0.010336,0.032314,0.008367,0.006345,0.006461,0.002659,-0.004191,0.00747,0.003147,0.010555,0.003028,-0.012622,0.012066,0.022426,0.002407,0.002263,0.001626,-0.000998,-0.007296,-0.001232,-0.006386,-0.000718

Avar:MGS015,0.132035,0.111708,0.039975,0.01292,0.022466,0.000279,0.003055,0.002077,0.003272,0.013668,-0.001949,0.001948,0.002379,0.004404,-0.006515,0.003845,0.004563,0.00076,0.002388,0.002751,-0.008984,0.000742,-0.003081,-0.00253,0.002395

Avar:MGS016,0.103579,0.028435,0.046386,0.011305,0.008617,0.005299,0.001175,0.011076,-0.007567,-0.004191,-0.009743,-0.006894,0.004311,0.004404,0.0019,0.002917,0.019427,0.00228,0.004022,0.009379,-0.014474,0.004575,0.001849,0.003253,-0.008622

Avar:MGS017,0.129758,0.129988,0.07467,0.05168,0.044624,0.012829,0.000235,0.008077,0.007158,-0.015126,-0.009094,-0.01154,0.00996,0.016652,0.000271,-0.003845,0.004824,-0.00266,0.000377,0.002501,-0.004866,-0.002844,0.002835,0.002169,0.001437

Avar:MGS018,0.122929,0.128972,0.063356,0.038437,0.032929,0.011435,-0.004935,0.012692,-0.002045,-0.00164,0.000325,0.00015,-0.001933,0.020506,-0.012486,-0.009149,0.001956,0.000887,0.00088,0.001251,-0.010856,0.001855,0.005176,0.00253,0.00491

Avar:MGS019,0.129758,0.146236,0.026021,-0.011305,0.026466,0.006136,0.005875,-0.006461,0.001636,0.005285,0.002273,0.005695,-0.007284,0.005643,-0.013029,-0.002121,0.007171,0.00228,0.004022,0.003252,-0.001497,-0.000866,-0.006409,-0.000723,-0.007065

Avar:MGS020,0.134311,0.141159,0.050911,0.036176,0.037853,0.003347,-0.0047,0.007615,0.000614,-0.005832,-0.005196,0.000899,0.000297,-0.009634,-0.000814,0.000133,-0.004433,-0.006081,0.005154,0.013381,-0.004118,0.00507,0.005423,-0.000482,-0.012095

Avar:MGS021,0.125205,0.101553,0.043746,0.013566,0.01508,0.004183,0.004465,0.006,-0.003068,0.002916,-0.007795,-0.001948,0.008028,0.010459,-0.008007,0,0.005737,-0.00266,0.004274,0.002376,-0.000749,0.000989,-0.002095,0.00253,-0.008502

Avar:MGS022,0.137726,0.141159,0.049026,0.040698,0.044008,0.022032,0.002585,0.00923,0.007158,0.000182,0.002598,0.004196,-0.002379,0.020506,0.000271,-0.013524,-0.008736,-0.00038,0.006913,-0.002001,-0.006613,0.007172,0.003697,-0.003856,-0.007185

Avar:MGS023,0.12862,0.13405,0.030924,0.011628,0.031083,0.006136,0.00094,0.009692,0.005318,0.018588,-0.004872,-0.004646,-0.001784,0.009771,-0.008279,0.00305,0.006128,0.003801,0.006411,-0.006253,0.002246,0.005935,-0.003081,0.004097,0.002395

Avar:MGS024,0.106994,0.052808,0.035449,-0.011951,0.008925,-0.010877,0.003995,0.003692,0.004704,0.017312,-0.005359,-0.002098,0.000892,0.000963,-0.009908,-0.003315,0.015646,-0.00152,-0.000754,-0.00075,-0.013102,0.002349,0.001109,0.006868,0.001078

Avar:MGS025,0.104717,0.051792,0.038089,-0.00969,0.009848,-0.007251,0.00517,-0.000923,0,0.017677,-0.012504,0.003297,-0.003419,-0.001239,-0.006379,0.005304,0.01356,0.00038,0.003268,-5e-04,-0.005241,-0.000124,-0.000493,0.007471,-0.008861

Avar:MGS026,0.119514,0.111708,0.023004,-0.001938,0.018157,-0.009203,0.01034,0.006923,-0.003272,0.008018,-0.001461,-0.002698,-0.001189,-0.003165,-0.013843,0.007027,0.023078,-0.002534,0.002388,0.001876,-0.007487,-0.006801,0.007395,-0.002892,0.00946

Avar:MGS027,0.121791,0.147252,0.038843,-0.004522,0.035391,-0.004741,0.001175,0.007846,-0.004704,0.01713,-0.000325,0.007343,-0.011298,0.003578,-0.013029,-0.002917,0.004824,-0.000507,0.009804,5e-04,-0.006613,0.002349,0.000863,0.004338,-0.00012

Avar:MGS028,0.122929,0.122879,0.031678,0.002261,0.019696,0.001673,0.003525,0.004615,-0.001636,0.006925,0.014128,0.004196,-0.003122,-0.004404,-0.000679,0.013126,0.023078,-0.010515,-0.00817,-0.001126,-0.000998,0.001237,-0.00037,-0.002651,-0.004311

Avar:MGS029,0.130897,0.126941,0.025267,-0.000969,0.025235,0.003347,0.006345,0.011769,-0.002045,0.009112,0.002761,0.001349,0.001487,-0.003716,-0.000543,0.008618,0.016559,-0.00228,-0.010433,-0.002126,-0.003119,0.003091,0.003944,-0.005302,-0.000479

Avar:MGS030,0.132035,0.121864,0.045632,0.01615,0.032621,0.004183,0.011516,0.006692,-0.001227,0.000911,0.002436,-0.001649,0.011596,-0.00578,-0.002714,0.012198,0.023991,-0.005321,-0.001383,0.009129,-0.010232,-0.000247,0.002835,0.001325,-0.00479

Avar:MGS031,0.129758,0.136081,0.030924,-0.000646,0.020311,0.006972,0.00705,0.009923,-0.001841,0.001822,0.004222,-0.007194,-0.002527,0.006744,-0.010043,0.000398,0.010822,-0.006334,-0.000377,-0.003252,-0.01722,-0.007666,-0.001602,0,0.000838

Avar:MGS032,0.133173,0.141159,0.040729,0.014212,0.028313,-0.006693,0.011516,0.004615,0.004704,0.004374,-0.000974,-0.001499,-0.000595,0.00234,-0.011129,-0.015513,-0.01708,-0.002534,0.003268,-0.00988,-0.006988,0.005564,-0.003451,-0.008435,0.000599

Avar:MGS033,0.125205,0.127957,0.035826,0.017119,0.029236,0.003904,0.006345,0.008769,0.0045,0.001276,-0.001137,-0.003447,0.001338,0.007156,-0.011808,-0.001458,0.016428,0.004181,0.004902,-0.001126,-0.012603,0.003957,0.001725,-0.006627,0.005508

Avar:MGS034,0.127482,0.147252,0.046386,0.019057,0.029236,0.006136,0.0047,0.002077,0.007158,0.001458,0.001949,0.001349,-0.005946,0.004954,-0.005565,0.000928,0.006128,0.001647,0.011816,-0.001876,-0.003868,0.005564,0.000616,0.002771,-0.007305

Avar:MGS035,0.129758,0.14319,0.029415,-0.000969,0.02739,-0.000558,0.001645,0.004384,-0.002045,-0.003462,0.004709,-0.001798,0.003122,0.001376,-0.013165,0.013126,0.021774,-0.009248,-0.005908,-0.004752,-0.010107,0.002844,-0.00419,0.00241,-0.001676

Avar:MGS036,0.130897,0.145221,0.04186,0.010013,0.027082,-0.000279,0.00611,-0.001154,0.002454,0.006925,-0.002111,-0.002847,0.003717,0.017753,-0.007193,-0.010475,-0.006389,-0.00038,0.003268,-0.002376,-0.010606,0.01286,-0.002465,-0.010242,-0.003113

Avar:MGS037,0.12862,0.15436,0.035072,-0.002907,0.041854,0.000837,0.011516,0.008077,0.004295,0.014761,-0.00406,0.003147,-0.0055,0.020919,-0.014386,-0.007027,-0.011474,-0.001267,0.005531,-0.009505,-0.006364,-0.000124,-0.006039,-0.001446,-0.00958

Avar:MGS038,0.120652,0.117801,0.024513,-0.008398,0.025235,-0.003904,0.003055,0.003461,0.004909,0.018953,0,-0.00045,-0.007284,-0.006193,-0.001764,0.007027,0.022817,-0.003041,0.001131,0.008504,-0.005989,-0.00136,-0.004314,-0.010001,0.001317

Avar:MGS039,0.118376,0.146236,0.02489,0.006783,0.027697,-0.002231,-0.002115,0.010846,0.002863,0.003645,0.001461,0.001049,-0.00446,0.003578,-0.005836,-0.007558,0.009648,-0.005448,0.003017,0.001251,0,-0.005193,-0.002835,0.001446,0.002515

Avar:MGS040,0.114961,0.082258,0.039598,0.012597,0.020311,-0.005578,0.00188,0.002308,0.004295,0.006743,-0.001786,-0.001049,-0.004608,0.009771,-0.0019,0.004773,0.0103,-0.003927,0.006285,0.008754,-0.009483,0.002349,-0.000986,0.002169,-0.008861

Avar:MGS044,0.130897,0.142174,0.035449,0.002907,0.028928,0.000837,0.004935,0.001846,0.000205,0.016219,-0.005521,0.002698,-0.00669,0.002477,0.002172,0.004773,-0.000391,-0.0019,0.003771,-0.007128,-0.000873,0.006677,-0.000739,0.001205,-0.003712

Avar:MGS045,0.120652,0.147252,0.022627,0.002261,0.016926,0.000837,0.00658,0.001846,-0.007976,0.009112,-0.005359,0.009891,-0.019475,0.000826,-0.00665,0.004243,-0.004824,0.012542,0.016466,-0.005878,0.018717,0.001113,0.003697,-0.003494,-0.016166

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