The vilelands leucrocotta is the smallest (by a small margin), least numerous (despite a wide range), and most reclusive culture of any of the leucrocotta people. They are also the most genetically isolated into the modern day, and have had virtually no intermixing of genes with any other group in at least 120,000 years and possibly as long as 200,000 (Littorals, though isolated for longer historically, now intermix slightly with continentals, and there is again some gene flow between populations.) For this reason, they are considered their own distinct species in the Truxmonoceros genus, though it is recognized by the author that this is arbitrary (taxonomy is all arbitrary.)
As their name indicates, this race lives mainly throughout the vilelands, a vast, inhospitable arid region near the equator characterized by brackish waterways and saline lakes, rocky calcified soil (the remnants of coastal reefs), and forests of cactaiga. They are a descendant species of the littoral leucrocotta, diverging some 300,000 years ago, and still carry a resemblance, but have also been changed by their isolation. They are shorter, with stockier legs and more robust necks, which suit them to clamber over rocky, uneven ground and move between thickets of spiny vegetation. Body weight is also around 120 lbs, but height is shorter than littorals on average. Their jaws are short, very narrow and highly serrated, with relatively long fang teeth. The digits resemble those of littorals but the dewclaws are not quite as dexterous, a reversion toward a more primitive state due to a lack of use. The neck mane is much shorter, but a ridge of hair runs down the entire length of the back. The crests are long and narrow, set closer together than in most littorals and always arced backwards like the horns of a goat. Though some littorals may have even longer crests - especially males - in those the crests usually have an outward arc. The eyes of the vilelands leucrocotta are positioned slightly more forward than in any other form and provide improved binocular vision, a trait which is beneficial as these leucrocotta preferentially live on sloped terrain, as it lets them flee other predators with less stable footing. Though the base colors can vary, light greys and tans similar to littorals are the typical pelage; striping is scarce, darkest on the legs and throat. All vilelands leucrocotta, except for rare cases, have highly contrasted black and white facial patterns which resemble skulls, and this trait - which increases visibility in low light - lends them their latin name of "masked leucrocotta." This is the only leucrocotta which has dark eyebrow markings rather than white, and they are most often a bright shade of red. A very striking black tear mark serves to catch glare and keep sunlight out of the eye during the day.
Today, the vilelands leucrocotta is the only one which still lives among larger carnivore species which both threaten and compete with it; continental leucrocotta have exterminated their rivals early in their history, and littorals retreated to hostile outlying regions where few rivals could survive (only the stalkvulture still persists, though even it has become very rare in the densely settled regions of Nexus Peninsula.) Natural selection has favored brains over brawns in this species, which still utilizes deception and trickery to outwit rivals much as earlier ancestral leucrocotta and their even earlier kelpie precursors did. Skillful mimics and master manipulators, vilelands leucrocotta still skillfully imitate the other species around them to lure prey, frighten enemies, and turn rivals against each other. The small size of the species lets them hide in cactaiga forests and flee up steep rocky slopes where they can orchestrate plans to lure and trap other animals while being untouchable. They use the terrain to their full advantage, drawing animals with vocal mimicry to fall into pitfall traps set up in natural low-lying ravines and chasms. In this way, much of their diet remains large prey, much too big for them to kill with their jaws alone, even in numbers. Their success at bringing down larger rivals has protected them from decimation from roaming continental leucrocotta in both ancient and recent history. While littorals tend to flee, vilelands leucrocotta will stand their ground and fight, using their knowledge of their land to win against stronger foes.
Though they use minimal tools, their skill in using the environment they live in to its full advantage to exploit and take down both rivals and prey is unequaled, and their memory of the landscape and its many varying routes through dangers is very sharp, to an almost photographic degree. In acting upon the vilelands leucrocotta, living in a more complex habitat than most, evolution seems to have rewired the part of its brain which in other leucrocotta allows rapid speech and language acquisition, to instead memorize visual details instantly and in great depth. Though they can still emulate most any sound, their ability to acquire new languages is significantly worse than other leucrocotta, and this has the effect of isolating them much more quickly, both from other species and within their kind. Vilelands leucrocotta are extremely tribal and kill their own species more than any other leucrocotta, for language barriers lead to rapid cultural change and an ease of viewing rivals as others. Their clans are small and break apart easily, rarely reaching more than three generations, and cooperative interactions, such as trade, is very limited across clans even of a single dialect. Vilelands leucrocottas can be considered war-like, prone to frequent conflict and orchestrated attacks on rival groups. This has the effect as well of making them anxious and always on edge, for danger can come from anywhere, and traps can be set at any time. Vilelands leucrocotta have an extremely short average life expectancy of as little as 8 to years, and very rarely live past 20; they are thus independent and sooner than others, as early as age 3, though they would not be considered adult in many other cultures for at least two more years.
A broad extent of range (but a sparse distribution over it) means that vilelands leucrocotta could theoretically meet littoral, steppe-continental, and eastern leucrocotta, though whisperwing claim of the land east of the Nexus peninsula has driven this species out of this extreme edge of its range so that they now do not typically meet the littorals today - and when they did, they were fierce rivals. Conflict with nomadic steppe continentals is ongoing, for the land where the two meet is considered a lawless frontier, while the Kir view these leucrocotta as the most unevolved and savage in the many-tiered hierarchy of leucrocotta in which they place themselves at the top. They have been so villified that they are hardly more than a pest, and generally viewed as below personhood. In earlier - and less enlightned - periods, they were considered so fierce and unsuitable for civilized life that they were not even considered usable as slaves. Kir have vastly superior technology, and have firmly pushed back the vilelands leucrocotta's range far from the green grassy fields of crater bay - where they once could be found on the rocky slopes nearest the sea - and exiling them to the harshest desert regions where no settlements are worth building. It is believed that in earlier eras, perhaps as recently as 3,000 years ago, that some coastal whisperwings formed trade relationship with at least some vilelands leucrocotta and may have cooperated in finding food and shared the spoils. This relationship no longer exists, as the vast majority of coastal wings are now urbanized, and this leucrocotta has long since been chased away from the northernmost reaches of its historic range where they would meet.