limnichidworldlist

Up date: 5 January 2025

CHECKLIST OF THE WORLD LIMNICHIDAE 

By HIROYUKI YOSHITOMI 

CHECKLIST 

Family Limnichidae Erichson, 1846 

Limnichidae Erichson, 1846: 465. --- Brown, 1991; Hinton, 1939b, 1939c; Paulus, 1970b; Sato, 1996; Wooldridge, 1975, 1986.

Subfamily Hyphalinae Britton, 1971 

Hyphalinae Britton, 1971: 88. --- Britton, 1971, 1977.

Genus Hyphalus Britton, 1971 

Hyphalus Britton, 1971: 88.

1. Hyphalus crowsoni Hernando & Ribera, 2000b: 240. (Seychelles [Aldabra]) 

2. Hyphalus insularis Britton, 1971: 90. (Australia) 

3. Hyphalus kuscheli Britton, 1977: 82. (New Zealand) 

4. Hyphalus prolixus Britton, 1977: 85. (New Zealand) 

5. Hyphalus taekoae M. Satô, 1997: 110. (Japan [Ryukyu Isls.], Taiwan) 

6. Hyphalus ultimus Britton, 1977: 85. (New Zealand) 

7. Hyphalus wisei Britton, 1973: 121. (New Zealand) 

Subfamily Cephalobyrrhinae Champion, 1925 

Cephalobyrrhinae Champion, 1925: 174. --- Wooldridge, 1981b; 1984.

Genus Cephalobyrrhus Pic, 1923 

Cephalobyrrhus Pic, 1923b: 4.

1. Cephalobyrrhus bertiae Pütz, 1998b: 344. (China [Yunnan]) 

2. Cephalobyrrhus brevipalpus Pütz, 1998b: 346. (China [Yunnan]) 

3. Cephalobyrrhus emeishanensis Pütz, 1998b: 347. (China [Sichuan]) 

4. Cephalobyrrhus gibbicollis Champion, 1925a: 175. (India [Kumaon]) 

5. Cephalobyrrhus jaechi Pütz, 1998b, 351. (China [Sichuan]) 

6. Cephalobyrrhus japonicus Champion, 1925a: 176. (Japan) 

7. Cephalobyrrhus jiangxiensis Pütz, 1998b: 354. (China [Jiangxi]) 

8. Cephalobyrrhus jinggangshanensis Pütz, 1998b: 355. (China [Jiangxi]) 

9. Cephalobyrrhus latus Pic, 1923b: 4. (Taiwan) 

10. Cephalobyrrhus longipalpis Pütz, 1998: 359. (China [Jiangxi]) 

11. Cephalobyrrhus nepalensis Pütz, 1998b: 360. (Nepal [Myagdi]) 

12. Cephalobyrrhus schillhammeri Pütz, 1998b: 362. (China [Sichuan]) 

13. Cephalobyrrhus schuelkei Pütz, 1998b: 363. (China [Sichuan]) 

14. Cephalobyrrhus sichuanensis Pütz, 1998b: 365. (China) 

15. Cephalobyrrhus sublongatus Pic, 1923b: 4. (India [Sikkim]) 

16. Cephalobyrrhus amami Yoshitomi, 2019 (Japan)

Genus Parathroscinus Wooldridge, 1984 

Parathroscinus Wooldridge, 1984c: 121.

1. Parathroscinus corrinae Wooldridge, 1990: 254. (Singapore) 

2. Parathroscinus magnus Wooldridge, 1990: 254. (Singapore) 

3. Parathroscinus murphyi Wooldridge, 1990: 251. (Singapore) 

4. Parathroscinus oculatus (Motschulsky), 1858: 52. (Burma, India) 

5. Parathroscinus orientalis Wooldridge, 1984c: 122. (Philippines, Vietnam) 

Genus Throscinus LeConte, 1874 

Throscinus LeConte, 1874: 51.

1. Throscinus aethiops Darlington, 1936: 51. (Cayman Isl., Cuba, Dominican Republic, Haiti, Jamaica, Puerto Rico) 

2. Throscinus crotchi LeConte, 1874: 52. (Mexico, USA [CA]) 

3. Throscinus politus Casey, 1889: 162. (USA [TX]) 

4. Throscinus punctatus Wooldridge, 1981 a: 220. (Colombia) 

5. Throscinus schwarzi Schaeffer, 1904: 204. (Jamaica, USA [TX]) 

6. Throscinus simplex Wooldridge, 1981 a: 220. (Colombia) 

7. Throscinus spangleri Woodridge, 1981 a: 218. (Ecuador) 

Genus Erichia Reitter, 1895

Erichia Reitter, 1895: 79. (type species: Erichia longicornis Reitter, 1895, by monotypy.)

1. Erichia longicornis Reitter, 1895: 79. () 

= Jaechobyrrhinus amanosius Pütz, 1991

2. † Erichia cretacea Yu et al., 2018: 844. (Burmese Amber)  

Genus Ercus García & Jiménez-Ramos, 2019

Ercus García y Jiménez-Ramos, 2019: 223. (type species: Ercus bitipus García & Jiménez-Ramos, 2019, by monotypy.)

1. Ercus bitipus García & Jiménez-Ramos, 2019: 225. (Venezuela)

Genus Anomocephalobus Li, Jäch & Cai, 2022

† Anomocephalobus Li, Jäch & Cai, 2022: 2. (type species: )

1. † Anomocephalobus liuhaoi Li, Jäch & Cai, 2022: 3. (Burmese Amber) 

Subfamily Thaumastodinae Champion, 1924 

Thaumastodinae Champion, 1924a: 25. --- Spilman, 1959, 1972; Wooldridge, 1988.

Genus Acontosceles Champion, 1924 

Acontosceles Champion, 1924a: 27.

1. Acontosceles borneensis Yoshitomi, 2019 (Malaysia [Borneo])

2. Acontosceles hydroporoides Champion, 1924a: 29. (Taiwan, India) 

3. Acontosceles tagalog Spilman, 1959: 116. (Philippines) 

4. Acontosceles yorioi M. Satô, 1966: 60. (Japan: Ryukyu) 

5. Acontosceles chujoi Yoshitomi & Satô, 2005: @ (Laos) 

6. Acontosceles jaechi Pütz, 2008: 308 (Philippines: Bohol) 

7. Acontosceles negrosensis Pütz, 2008: 310 (Philippines: Negrosl) 

8. Acontosceles quatuordecimmaculosus Pütz, 2008: 312 (Myanmar) 

9. Acontosceles siwalikensis Pütz, 2008: 314 (Nepal) 

10. Acontosceles yunnanensis Pütz, 2008: 315 (China: Yunnan) 

11. Acontosceles zetteli Pütz, 2008: 317 (Thailand) 

12. Acontosceles javanicus Yoshitomi & Putra, 2011 (Indonesia, Java) 

Genus Martinius Spilman, 1959 

Martinius Spilman, 1959: 118.

1. Martinius ripisaltator Spilman, 1966: 124. (Cuba) 

2. Martinius tellepontis Spilman, 1959: 119. (Panama) 

3. Martinius temporalis Wooldridge, 1988: 314. (Ecuador) 

Genus Mexico Spilman, 1972 

Mexico Spilman, 1972: 113.

Babalimnichus M. Satô, 1994: 174. --- Yoshitomi & Satô, 2001: @@@

1. Mexico litoralis Spilman, 1972: 114. (Mexico)

2. Mexico morrisoni Skelley, 2005: 120. (Bahamas)

3. Mexico masamii (M. Satô, 1994): 175. (Japan: Honshu, Kyushu, Ryukyu) 

4. Mexico taiwanus (M. Satô, 1994): 174. (Taiwan) 

5. Mexico splendens (Hernando & Ribera, 2003): 270. (Tonga Isls.) 

6. Mexico baliensis Yoshitomi, 2019 (Indonesia [Bali Island])

7. Mexico borneensis Yoshitomi, 2019 (Malaysia [Borneo])

8. Mexico ogasawaraensis Yoshitomi, 2019 (Japan [Ogasawara Islands])

9. Mexico palauensis Yoshitomi, 2019 (Palau)

10. Mexico papuanus Yoshitomi, 2019 (Papua New Guinea)

Genus Pseudeucinetus Heller, 1921 

Pseudeucinetus Heller, 1921: 155.

Thaumastodus Champion, 1924a: 25.

Pseudoeuchinetus (misspelling): Satô, 1994: 173.

1. Pseudeucinetus novabritannica Delève, 1973: 18. (Bismark [New Britain]) 

2. Pseudeucinetus spilmani Spangler, 1995: 399. (Indonesia) 

3. Pseudeucinetus uenoi Spangler, 1995: 396. (Malaysia) 

4. Pseudeucinetus zygops Heller, 1921: 156. (India, Malaysia, New Ireland, Philippines, Solomon Isls.) (syn., Taumastodus fusiformis Champion, 1924a: 27) 

5. Pseudeucinetus javanicus Yoshitomi & Putra, 2010 

6. Pseudeucinetus solomonicus Yoshitomi & Putra, 2010 

7. Pseudeucinetus papuanus Yoshitomi, 2019 (Papua New Guinea)

Subfamily Limnichinae Erichson, 1846 

Limnichinae Erichson, 1846: 465 (including Bothriophorinae). --- Lawrence, 1987a; Wooldridge, 1977a; 1977b, 1980; 1981a; 1982; 1987.

Bothriophorini

Genus Bothriophorus Mulsant & Rey, 1852 

Bothriophorus Mulsant & Rey, 1852: 19.

1. Bothriophorus atomus Mulsant & Rey, 1852: 20. (Europe) (syn., ventetus Ferrari, 1864: 108) 

2. 
   

Wooldridgeini

Genus Platypelochares Champion, 1923 

Platypelochares Champion, 1923: 271.

Wooldridgeus Spangler, 1999: 181.

1. Platypelochares latimargo Champion, 1923: 272. (India [Kumaon]) 

2. Platypelochares perforatus (Spangler), 1999: 182. (Malaysia) 

3. Platypelochares periculosissimus Ribera & Hernando, 1999: 107. (Laos, Thailand, Vietnam) 

4. Platypelochares petrus Ribera & Hernando, 1999: 108. (Malaysia) 

5. Platypelochares trifidus Champion, 1923: 272. (Burma) 

6. Platypelochares sinensis Yoshitomi, 2011: 385 (China) 

Limnichini (Limnichus genus-group: sensu Hernando & Ribera, 2004)

Genus Chibidoronus M. Satô, 1966 

Chibidoronus M. Satô, 1966: 59.

1. Chibidoronus aureus M. Satô, 1966: 60. (Japan [Ryukyu]) 

Genus Geolimnichus Hernando & Ribera, 2003 

Geolimnichus Hernando & Ribera, 2003: 386. Type species: Geolimnichus endroedyi Hernando & Ribera, 2003: 387.

1. Geolimnichus endroedyi Hernando & Ribera, 2003: 387. (Republic of South Africa) 

2. Geolimnichus coprophilus Hernando & Ribera, 2003: 388. (Republic of South Africa) 

Genus Limnichoderus Casey, 1889 

Limnichoderus Casey, 1889: 147.

1. Limnichoderus amoenus Wooldridge, 1981b: 179. (Ecuador) 

2. Limnichoderus angustus Wooldridge, 1981b: 180. (Colombia, Ecuador, Panama, Venezuela) 

3. Limnichoderus barberi Wooldridge, 1981b: 174. (Haiti, Puerto Rico, Trinidad-Tobago) 

4. Limnichoderus conspectus Wooldridge, 1981b: 181. (Ecuador, Peru) 

5. Limnichoderus cupidus Wooldridge, 1981b: 178. (Brazil) 

6. Limnichoderus curtulus Sharp, 1902a: 678. (Costa Rica, Panama) (syn., minutus Pic, 1922: 4) 

7. Limnichoderus durus Wooldridge, 1981b: 182. (Brazil) 

8. Limnichoderus excelsus Wooldridge, 1993a: 35. 

9. Limnichoderus fragosus Wooldridge, 1981b: 178. (Ecuador) 

10. Limnichoderus imprioris Blackwelder, 1944: 274. (Argentina) (syn., oblongus Bréthes, 1922: 297; preoccupied) 

11. Limnichoderus insularis Wooldridge, 1981b: 188. (Dominican Republic, Haiti, Puerto Rico) 

12. Limnichoderus lutrochinus (LeConte), 1879: 515. (Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Panama, USA) (syn., indiscretus Sharp, 1902a: 678) (syn., lanosus Casey, 1912: 55) 

13. Limnichoderus magnus Wooldridge, 1981b: 179. (Costa Rica, Guatemala, Mexico) 

14. Limnichoderus modicus Wooldridge, 1981b: 176. (Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Panama) 

15. Limnichoderus maratus Wooldridge, 1981b: 185. (Colombia, Costa Rica, Guatemala, Mexico, Panama) 

16. Limnichoderus naviculatus (Casey), 1889: 156. (Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Puerto Rico, USA) 

17. Limnichoderus ovatus (LeConte), 1854: 117. (Mexico, USA) (syn., pulvereus Casey, 1912: 54) 

18. Limnichoderus placidus Wooldridge, 1987a: 330. (Costa Rica) 

19. Limnichoderus plenus Wooldridge, 1981b: 185. (Mexico) 

20. Limnichoderus punctiventris (Casey), 1889: 159. (USA) 

21. Limnichoderus seclusus Wooldridge, 1987a: 330. (Ecuador) 

22. Limnichoderus seriatus (Casey), 1889: 158. (USA) (syn., floridanus Casey, 1912: 55) 

23. Limnichoderus similis Wooldridge, 1981b: 182. (Peru) 

24. Limnichoderus vicinus Pic, 1922: 4. (Brazil, Colombia, Ecuador, Panama, Venezuela) 

Genus Limnichomorphus Pic, 1922 

Limnichomorphus Pic, 1922: 4.

1. Limnichomorphus curtulus Pic, 1922: 4. (Vietnam [Tonkin]) 

2. Limnichomorphus ohbayashii M. Satô, 1966: 58. (Japan [Ryukyu Isls.]) 

3. Limnichomorphus puetzi Hernando & Ribera, 2004: 45. (Nepal)

4. Limnichomorphus ciampori Hernando & Ribera, 2004: 45. (Malaysia [Borneo, Sabah])

Genus Limnichus Latreille, 1829 

Limnichus Latreille, 1829: 510.

1. Limnichus aeneipennis Pic, 1935a: 1. (Vietnam) 

2. Limnichus africanus Pic, Pic, 1923a: 3. (Ghana) 

3. Limnichus angustulus Weise, 1877: 300. (Sardinia) 

4. Limnichus astruci Pic, 1922: 3. (India) 

5. Limnichus ater Lea, 1920: 279. (Australia) 

6. Limnichus aurosericus Jacquelin de Val, 1857 (1859): 268. (Algeria, France, Italy, Spain) (syn., leprieuri Perris, 1864: 282) 

7. Limnichus australis Erichson, 1842: 153. (Australia, Tasmania) 

8. Limnichus brevior Fairmaire, 1898: 224. (Madagascar) 

9. Limnichus coomani Pic, 1935a: 1. (Vietnm) 

10. Limnichus cuneatus Champion, 1923: 275. (India [Kumaon]) 

11. Limnichus curtus Pic, 1922: 3. (Java) 

12. Limnichus decornus Broun, 1880: 247. (New Zealand) 

13. Limnichus demoulini Delève, 1968: 217. (Congo) 

14. Limnichus ellipticus Lea, 1920: 280. (Australia) 

15. Limnichus falsus Delève, 1971: 371. (India) 

16. Limnichus fragilicornis Wollaston, 1868: 80. (Cape Verde Islands) 

17. Limnichus fulvopubens Pic, 1922: 3. (China) 

18. Limnichus globosus Champion, 1923: 276. (India [Kumaon]) 

19. Limnichus incanus Liesenwetter, 1851: 584. (Mediterranean area, southern Europe, Caucasus, Turkestan) 

20. Limnichus inornatus Weise, 1887: 301. (Cyperus) (syn., punctipennis Baudi, 1870: 64) 

21. Limnichus javanicus Pic, 1922: 3. (Java) 

22. Limnichus latiusculus Reitter, 1901: 72. (Turkestan) 

23. Limnichus lederi Weise, 1877: 302. (Causasus) 

24. Limnichus lewisi Nakane, 1963: 43. (China, Japan, Russia) (syn., paululus Lewis, 1879: 12) 

25. Limnichus longitarsis Champion, 1923: 275. (India [Sarda]) 

26. Limnichus marmoratus Fairmaire, 1898: 224. (Madagascar) 

27. Limnichus micraspis Champion, 1923: 276. (India [Sarda]) 

28. Limnichus monticola Delève, 1968: 220. (Kenya) 

29. Limnichus murinus Baudi, 1870: 63. (Cyperus) 

30. Limnichus nigripes Broun, 1893: 1449. (New Zealand) 

31. Limnichus obliteratus Champion, 1923: 275. (India [Kumaon]) 

32. Limnichus orientalis Motschulsky, 1861: 112. (Sri Lanka) 

33. Limnichus palawanus Delève, 1973: 18. (Philippines [Palawan]) 

34. Limnichus picinus Broun, 1881: 671. (New Zealand) 

35. Limnichus pruinosus Fleischer, 1909: 244. (Samarkand) 

36. Limnichus pumilio Obenberger, 1916: 29. (Corfu) 

37. Limnichus punctatus Broun, 1880: 247. (New Zealand) 

38. Limnichus punctipennis Kraatz, 1858: 148. (Greece, Italy) 

39. Limnichus pygmaeus Sturm, 1807: 114. (Turkestan, middle Europe, Causasus) (syn., sericeus Stephens, 1832: 411) (syn., auricomus Reitter, 1877 (1878): 180) (syn., minutus Rey, 1889: 66) (syn., variegatus Guillebeau, 1886: 2) (syn., pygmaeus var. Reitter, 1887: 104) 

40. Limnichus ruficolor Pic, 1935a: 2. (N. Borneo) 

41. Limnichus rufipes Pic, 1923b: 6. (Ivory Coast) 

42. Limnichus rufulopubens Fairmaire, 1897: 368. (Madagascar) 

43. Limnichus rufus Pic, 1923b: 7. (Philippines [Luzon]) 

44. Limnichus saegeri Delève, 1968: 218. (Congo) 

45. Limnichus sericeus Dufschmid, 1825: 24. (Europe, Asia) 

46. Limnichus subchalybeus Reitter, 1881: 85. (Caucasus) 

47. Limnichus subelongatus Pic, 1922: 3. (Sumbawa) 

48. Limnichus subnotatus Champion, 1923: 273. (India [Kumaon]) 

49. Limnichus sulcatulus Pic, 1922: 3. (Guadeloupe) 

50. Limnichus tonkineus Fairmaire, 1888: 337. (Tonkin, Vietnam) 

51. Limnichus trachyformis Pic, 1922: 3. (India [Kumaon]) 

52. Limnichus turkestanicus Pic, 1905: 97. (Turkestan) 

53. Limnichus unistriatus Champion, 1923: 274. (India [Kumaon]) 

54. Limnichus vageguttatus Fairmaire, 1898: 223. (Madagascar) 

55. Limnichus waelbroecki Delève, 1968: 215. (Kinshasha) 

Mandersia genus-group of Limnichinae (sensu Hernando & Ribera, 2005)

Genus Euthryptus Sharp, 1902 

Euthryptus Sharp, 1902a: 682.

1. Euthryptus orbicularis Sharp, 1902a: 682. (Guatemala, Mexico, Panama) 

2. Euthryptus trigonalis Sharp, 1902a: 682. (Guatemala) 

Genus Resachus Delève, 1968 

Resachus Delève, 1968: 263.

1. Resachus lineatifrons Delève, 1968: 266. (Congo) 

2. Resachus striatellus Delève, 1968: 263. (Congo) 

3. Resachus gillianae Matsumoto, 2020 (Malawi)

4. Resachus schuhi Hernando & Ribera, 2006 (Madagascar)

5. Resachus bilardoi Hernando, 2024: 169. (Gabon)

Genus Simplocarina Pic, 1922 

Simplocarina Pic, 1922: 4.

= Cyrtolimnichus Delève, 1968: 267.

1. Simplocarina curticollis Pic, 1922: 4. (Congo) (syn., punctulatus Delève, 1968: 267) 

Genus Mandersia Sharp, 1902 

Mandersia Sharp, 1902b: 62.

1. Mandersia scymnoides Sharp, 1902b: 62. (Burma, China, India) 

Genus Pseudothryptus Hernando & Ribera, 2005 

Pseudothryptus Hernando & Ribera, 2005: 131. Type species: Caccothryptus multiseriatus Champion, 1923.

1. Pseudothryptus multiseriatus (Champion), 1923: 224. (India [Bengal]) 

Genus Caccothryptus Sharp, 1902 

Caccothryptus Sharp, 1902b: 63. --- Hernando & Ribera, 2005: 131 [redescription].

Macrobyrrhinus Pic, 1922: 4. (Type species: M. rouyeri Pic by monotypy; synonymized by Champion, 1923: 222.)

the compactus group

1. Caccothryptus abboti Matsumoto, 2021: 181. (India)

2. Caccothryptus compactus Sharp, 1902b: 63. (Borneo, Sumatra, Thailand) 

3. Caccothryptus larryi Matsumoto, 2021: 176. (India)

4. Caccothryptus maculosus Pic, 1923a: 3. (Cambodia, China [Hainan], India [Andaman], Indonesia [Bali, Java, Nias], Laos, Myanmar, Thailand, Vietnam) 

= Caccothryptus laosensis Pic, 1928: 2. (Laos) 

5. Caccothryptus schillhammeri Hernando et Ribera, 2017 (Myanmar)

6. Caccothryptus schuhi Hernando et Ribera, 2014 (Indonesia [Java])

7. Caccothryptus sulawesianus Hernando et Ribera, 2014 (Indonesia [Sulawesi])

8. Caccothryptus thai Hernando et Ribera, 2017 (Thailand)

the rouyeri group

9. Caccothryptus rouyeri (Pic, 1922) (Sumatra) 

the testudo group

10. Caccothryptus arakawae Matsumoto, 2021: 171. (Nepal)

11. Caccothryptus auratus Hernando et Ribera, 2014 (Thailand)

12. Caccothryptus brendelli Matsumoto, 2021: 169. (Nepal)

13. Caccothryptus championi Matsumoto, 2021: 174. (India)

14. Caccothryptus chayuensis Yoshitomi, 2015 (Tibet)

15. Caccothryptus fujianensis Hernando et Ribera, 2014 (China: Fujian)

16. Caccothryptus jendeki Hernando et Ribera, 2014 (India)

17. Caccothryptus malickyi Hernando et Ribera, 2014 (Vietnam)

18. Caccothryptus nepalensis Hernando et Ribera, 2014 (Nepal)

19. Caccothryptus occidentalis Hernando & Ribera, 2017 (India)

20. Caccothryptus orion Yoshitomi, 2015 (Japan [Okinawa])

21. Caccothryptus punctatus (Pic, 1923) (Vietnam)

22. Caccothryptus yunnanensis Yoshitomi, 2018 (China [Yunnan])

23. Caccothryptus ripicola Champion, 1923: 223. (India [W. Almora], Nepal) 

24. Caccothryptus sinensis Hernando et Ribera, 2014 (China [Fujian])

25. Caccothryptus taiwanus Yoshitomi, 2015 (Taiwan)

26. Caccothryptus tardarsauceae Matsumoto, 2021: 178. (India)

27. Caccothryptus testudo Champion, 1923: 222. (India [W. Almora], Nepal, Thailand) 

28. Caccothryptus tibetanus Yoshitomi, 2015 (Tibet)

the jaechi group

29. Caccothryptus jaechi Hernando et Ribera, 2014 (Indonesia: Sulawesi)

30. Caccothryptus nanus Hernando et Ribera, 2014 (Philippines [Luzon])

31. Caccothryptus ticaoensis Hernando et Ribera, 2014 (Philippines [Ticao])

32. Caccothryptus wooldridgei Hernando et Ribera, 2014 (Indonesia [Sulawesi])

the zetteli group

33. Caccothryptus luzonensis Hernando et Ribera, 2014 (Philippines [Luzon])

34. Caccothryptus zetteli Hernando et Ribera, 2014 (Philippines [Luzon])

Limnichinae incertae sedis

Genus Burmochares Kirejtshuk et Prokin, 2022

† Burmochares Kirejtshuk et Prokin, 2022: 3. (type species: Burmochares groehni Kirejtshuk et Prokin, 2022, by the original designation)

1. †  Burmochares groehni Kirejtshuk et Prokin, 2022: 7. (Burmese amber) 

Genus Burmochares Kirejtshuk et Prokin, 2022

†  Genus Hernandochares Kirejtshuk et Prokin, 2022: 9. (type species: Platypelochares electricus Hernando. Azawaryn et Ribero, 2018)

1. †  Hernandochares electricus Hernando. Azawaryn et Ribero, 2018 (Burmese amber) 

Genus Byrrhinus Motschulsky, 1858 

Byrrhinus Motschulsky, 1858: 50.

Byrrhininus Pic, 1922: 6.

Cyphonichus Sharp, 1902a: 679.

Notiocyphon Blackburn, 1896: 272.

Pelocherops Pic, 1923a: 3.

1. Byrrhinus amoenus Wooldridge, 1987a: 310. (Belize, British Guiana, Colombia, Ecuador) 

2. Byrrhinus andamanus (Pic), 1935a: 2. (Andaman Isls.) 

3. Byrrhinus angustatus Motschulsky, 1858: 51. (India, Thailand) 

4. Byrrhinus antipodium Fauvel, 1903: 348. (New Caledonia) 

5. Byrrhinus apicalis Pic, 1922: 5. (Katanga) 

6. Byrrhinus atricornis (Pic), 1935a: 3. (Congo, Sierra Leone) 

7. Byrrhinus atrolucidus (Pic), 1923a: 3. (New Guinea) 

8. Byrrhinus binhanus (Pic), 1923b: 5. (Vietnam) 

9. Byrrhinus biroi Pic, 1956: 75. (New Guinea) 

10. Byrrhinus bomansi (Delève), 1968: 245. (Tanganyika) 

11. Byrrhinus borneensis Pic, 1923b: 7. (Borneo) 

12. Byrrhinus boucardi Pic, 1923b: 7. (Sumatra) 

13. Byrrhinus brevior (Pic), 1927a: 65. (Sumatra) 

14. Byrrhinus convexus (Pic), 1923b: 5. (Philippines) 

15. Byrrhinus cribrosus Champion, 1923: 253. (India, Sri Lanka) 

16. Byrrhinus curtus Pic, 1935b, 471. (Sierra Leone) 

17. Byrrhinus densepunctatus (Pic), 1923b: 6. (Ivory Coast) 

18. Byrrhinus diversus (Pic), 1923b: 5. (Ivory Coast) 

19. Byrrhinus ellypticus Motschulsky, 1858: 51. (India) 

20. Byrrhinus euryaspis Champion, 1923: 250. (India [Nilgiri Hills]) 

21. Byrrhinus ferax Wooldridge, 1993b: 359. (Philippines, Borneo) 

22. Byrrhinus flavicornis Champion, 1923: 251. (India) 

23. Byrrhinus formosanus Pic, 1935a: 2. (Taiwan) 

24. Byrrhinus fuscus Motschulsky, 1858: 52. (India) 

25. Byrrhinus goldbachi Wooldridge, 1987b: 310. (Ecuador, Paraguay) 

26. Byrrhinus gracilicornis Pic, 1922: 5. (Brazil, Paraguay, Venezuela) 

27. Byrrhinus grandis (Pic), 1956: 74. (New Guinea) 

28. Byrrhinus grossepunctatus (Delève), 1968: 249. (Congo) 

29. Byrrhinus hargreavesi (Pic), 1935b: 470. (Sierra Leone) 

30. Byrrhinus hirsutulus Champion, 1923: 250. (India [Kumaon]) 

31. Byrrhinus hirsutus Pic, 1922: 5. (French Guiana, Trinidad) 

32. Byrrhinus hulstaerti (Pic), 1952: 9. (Kinshasa [Congo]) 

33. Byrrhinus improcerus Wooldridge, 1987b: 310. (Ecuador) 

34. Byrrhinus inlineatus Pic, 1923c: 15. (India) 

35. Byrrhinus irregularis Pic, 1922: 5. (Philippines) 

36. Byrrhinus jacobsoni Pic, 1927a: 65. (Sumatra) 

37. Byrrhinus latus Motschulsky, 1858: 50. (Bangladesh, Myanmar, India, Sri Lanka) 

38. Byrrhinus leleupi (Delève), 1968: 257. (Kivu [Zaire]) 

39. Byrrhinus leonensis (Pic), 1923b: 6. (Congo, Sierra Leone) 

40. Byrrhinus maculatus Wooldridge, 1987b: 307. (Ecuador) 

41. Byrrhinus magnus Wooldridge, 1987b: 307. (Ecuador, Perú) 

42. Byrrhinus marginatus Champion, 1923: 249. (India [Kumaon], Pakistan) 

43. Byrrhinus maroniensis Pic, 1923b: 7. (Brazil, Colombia, French Guiana, Trinidad-Tobago, Venezuela) 

44. Byrrhinus mimicus Wooldridge, 1987b: 308. (Colombia, Panama) 

45. Byrrhinus misellus (Sharp), 1902a: 681. (Colombia, Costa Rica, Mexico, Nicaragua, Panama) 

46. Byrrhinus monilicornis Champion, 1923: 251. (India [Kumaon]) 

47. Byrrhinus neoguineensis (Pic), 1956: 75. (New Guinea) 

48. Byrrhinus niger Pic, 1922: 5. (Vietnam) 

49. Byrrhinus nitidicollis Pic, 1956: 75. (New Guinea) 

50. Byrrhinus nitidulus Motschulsky, 1858: 51. (India, Thailand) 

51. Byrrhinus noctivagus Lea, 1920: 277. (Australia) 

52. Byrrhinus oculatus Motschulsky, 1858: 52. (India) 

53. Byrrhinus olibroides (Sharp), 1902a: 679. (Belize, El Salvador, Guatemala, Mexico) (syn., selatus Sharp, 1902a: 680) 

54. Byrrhinus orientalis (Sharp), 1902b: 63. (Taiwan, Malaysia [Borneo]) 

55. Byrrhinus padangus Pic, 1927b: 49. (Sumatra) 

56. Byrrhinus panamensis (Sharp), 1902a: 679. (Belize, Colombia, Costa Rica, Guatemala, Panama) (syn., mollis Sharp, 1902a: 681)(syn., rufipennis Pic, 1923b: 7) 

57. Byrrhinus peguensis Pic, 1923c: 15. (India) 

58. Byrrhinus plenus Wooldridge, 1987b: 305. (Boliva, Ecuador) 

59. Byrrhinus pubiventris Lea, 1920: 277. (Australia) 

60. Byrrhinus punctaticeps Pic, 1927b: 49. (Sumatra) 

61. Byrrhinus punctatus (Pic), 1922: 5. (Philippines) 

62. Byrrhinus punctulatus (Delève), 1968: 243. (Upemba [Zaire]) 

63. Byrrhinus reitteri (Pic), 1923b: 6. (Ghana) 

64. Byrrhinus rhomboideus Champion, 1923: 253. (India) 

65. Byrrhinus rufoapicalis (Pic), 1955: 132. (Ruanda, Kiva [Zaire]) 

66. Byrrhinus rufus Pic, 1935a: 2. (Thailand) 

67. Byrrhinus schoutedeni (Pic), 1952: 9. (Congo) 

68. Byrrhinus semirufus Pic, 1923b: 8. (Trinidad-Tobago, Venezuela) 

69. Byrrhinus seydeli (Delève), 1968: 252. (Elisabethville [Zaire]) 

70. Byrrhinus sikkimensis Champion, 1923: 249. (India [Sikkim]) 

71. Byrrhinus subglobosus subglobosus (Delève), 1968: 254. (Elisabethville [Zaire]) 

72. Byrrhinus subglobosus upembanus (Delève), 1968: 256. (Upemba [Zaire]) 

73. Byrrhinus subregularis Pic, 1923c: 15. (Vietnam) 

74. Byrrhinus subtestaceous Pic, 1923b: 8. (Philippines) 

75. Byrrhinus tarawakanus Delève, 1973: 23. (Borneo [Sabah], Philippines [Tawi Tawi]) 

76. Byrrhinus testaceicornis (Pic), 1923b: 5. (Philippines) 

77. Byrrhinus testaceipes Pic, 1922: 5. @@@@ 

78. Byrrhinus subregularis Pic, 1923b: 7. (Lapsus Calami, Sumatra) @@@@ 

79. Byrrhinus tschoffeni (Delève), 1968: 235. (Boma [Congo]) 

80. Byrrhinus vatovai (Pic), 1953: 113. (Ethiopia) 

81. Byrrhinus vestitus (Sharp), 1902a: 681. (Guatemala, Mexico) 

82. Byrrhinus vicinus (Delève), 1968: 248. (Congo) 

83. Byrrhinus wittei (D elève), 1968: 253. (Upemba [Zaire]) @@@@ wittmeri?? 

84. Byrrhinus helicophallus Hernando et Ribera, 2014 (Yemen)

85. Byrrhinus socotrensis Hernando et Ribera, 2014 (Socotra)

Genus Limnichites Casey, 1889 

Limnichites Casey, 1889: 147.

1. Limnichites browni Wooldridge, 1977a: 187. (El Salvador, Mexico) 

2. Limnichites confertus (Sharp), 1902a: 676. (Costa Rica, Guatemala, Honduras, Jamaica, Mexico, Nicaragua) 

3. Limnichites foraminosus Casey, 1912: 43. (Mexico, USA) 

4. Limnichites huronicus Casey, 1912: 43. (Canada, USA) 

5. Limnichites imparatus Wooldridge, 1977a: 181. (Mexico) 

6. Limnichites nebulosus (LeConte), 1879: 515. (Mexico, USA) (syn., densissimus Casey, 1912: 41) 

7. Limnichites olivaceus (LeConte), 1854: 116. (USA) (syn., virginicus Casey, 1912: 42) 

8. Limnichites perforatus (Casey), 1889: 155. (Mexico, USA) 

9. Limnichites porrectus Wooldridge, 1977a: 182. (Mexico) 

10. Limnichites punctatus (LeConte), 1854: 116. (Canada, USA) (syn., austinianus Casey, 1912: 42) 

11. Limnichites rudis Wooldridge, 1977a: 183. (Cuba, Jamaica) 

12. Limnichites simplex Wooldridge, 1977a: 188. (Costa Rica, Mexico) 

Genus Paralimnichus Delève, 1973 

Paralimnichus Delève, 1973: 24. --- Kasap & Crowson, 1975; Wooldridge, 1983.

1. Paralimnichus castaneus (Lea), 1920: 279. (Australia) 

2. Paralimnichus panamensis Wooldridge, 1983: 232. (Panama) 

3. Paralimnichus punctatus (Pic), 1923b: 5. (Borneo, Guadalcanal, Philippines [Banguey]) (syn., punctatus var. obscurior Pic, 1923b: 5) (syn., rufescens Pic, 1923b: 5) 

Genus Pelochares Mulsant & Rey, 1869 

Pelochares Mulsant & Rey, 1869: 161.

1. Pelochares acuminatus Champion, 1923: 271. (India [Sikkim]) 

2. Pelochares acutipennis Pic, 1935a: 2. (Malaysia) 

3. Pelochares alluaudi Delève, 1968: 226. (Africa Prientale) 

4. Pelochares basilewski (Pic), 1955: 131. (Ruanda) 

5. Pelochares biroi (Pic), 1956: 75. (New Guinea) 

6. Pelochares congoensis Delève, 1968: 221. (Congo) 

7. Pelochares densatus Pic, 1922: 3. (Java) 

8. Pelochares diversus (Pic), 1922: 3. (Philippines [Los Banos] 

9. Pelochares emarginatus Mulsant & Rey, 1869: 161. (Europe) 

10. Pelochares gibbipennis Champion, 1923: 269. (Europe, Burma, India [Bengal], Sri Lanka) 

11. Pelochares minutus Delève, 1968: 224. (Kivu [Zaire]) 

12. Pelochares murinus Baudi, 1870: 63. (Cyperus, Caucasus) 

13. Pelochares oblongulus Champion, 1925b: 263. (India [Kumaon]) 

14. Pelochares orientalis (Motschulsky), 1861: 112. (Sri Lanka) (syn., horni Delève, 1971: 368) 

15. Pelochares philippinensis Delève, 1973: 21. (Philippines [Tawi Tawi]) 

16. Pelochares rugiventrus Champion, 1923: 270. (India [Kumaon], Sri Lanka) (syn., seylanicus Delève, 1971: 370) 

17. Pelochares ryukyuensis M. Satô, 1966: 57. (Taiwan, Japan [Ryukyu Isls.]) 

18. Pelochares sulciger Champion, 1923: 270. (Sri Lanka) 

19. Pelochares trifidus Champion, 1923: 272. (Burma) 

20. Pelochares versicolor Waltl, 1838: 273. (Europe, New Caledonia) var., opacus Reitter, 1881: 84. (syn., emarginatus Mulsant & Rey, 1869: 161) 

21. Pelochares vitalisi Pic, 1935a: 1. (Cambodia) 

Genus Physemus LeConte, 1854 

Physemus LeConte, 1854: 117.

Ditaphrus Casey, 1886: 250.

Genus Afrolimnichus Delève, 1968 

Afrolimnichus Delève, 1968: 259

1. Afrolimnichus brevicornis Delève, 1968: 260. (Congo) 

2. Afrolimnichus oblongus Delève, 1974: 40. (South Africa) 

Genus Cephalobyrrhinus Pic, 1923 

Cephalobyrrhinus Pic, 1923b: 6.

Cephalobyrrhinus brevis Wooldridge, 1986a: 512. (Ecuador)

1. Cephalobyrrhinus curticornis Pic, 1923b: 6. (Costa Rica, Mexico, Nicaragua, Panama, Venezuela) 

2. Cephalobyrrhinus impressopunctatus Wooldridge, 1977b: 32. (Bolivia) 

3. Cephalobyrrhinus lineatus Wooldridge, 1986a: 513. (Costa Rica, Panama) 

4. Cephalobyrrhinus robustus Wooldridge, 1986a: 512. (Ecuador, Peru) 

5. Cephalobyrrhinus sedatus Wooldridge, 1986a: 513. (Ecuador) 

Genus Corrinea Woodridge, 1980 

Corrinea Woodridge, 1980: 69.

1. Corrinea admirandus (Sharp), 1902a: 677. (Mexico) 

2. Corrinea audax Wooldridge, 1980: 75. (Mexico) 

3. Corrinea aurata Wooldridge, 1980: 71. (Brazil, Ecuador) 

4. Corrinea delicata Wooldridge, 1980: 75. (Colombia, Costa Rica, Mexico, Panama) 

5. Corrinea edita Wooldridge, 1993a: 36. (Venezuela) 

6. Corrinea incondita Wooldridge, 1980: 73. (Bolivia, Colombia, Ecuador) 

7. Corrinea lecta Wooldridge, 1980: 74. (Bolivia) 

8. Corrinea parva Wooldridge, 1980: 74. (Bolivia, Ecuador, Peru) 

9. Corrinea placida Wooldridge, 1980: 76. (Trinidad) 

10. Corrinea salebrosa Wooldridge, 1980: 75. (Guatemala) 

11. Corrinea spicula Wooldridge, 1980: 73. (Ecuador) 

12. Corrinea splendida Wooldridge, 1980: 71. (Ecuador, Peru 

13. Corrinea vianai (Pic), 1939: 249. (Argentina) 

Genus Cyclolimnichus Delève, 1968 

Cyclolimnichus Delève, 1968: 270.

1. Cyclolimnichus dentoni Hernando & Ribera, 2000a: 216. (Cameroon) 

2. Cyclolimnichus jaechi Hernando & Ribera, 2000a: 215. (Kenya) 

3. Cyclolimnichus ovalis Hernando & Ribera, 2000a: 215. (Cameroon) 

4. Cyclolimnichus presignis Delève, 1968: 272. (Congo) 

Genus Ersachus Erichson, 1847 

Ersachus Erichson, 1847: 497.

1. Ersachus erichsonianus Sharp, 1902a: 638, pl. 20. (Guatemala, Mexico, Panama) 

2. Ersachus lugubris Sharp, 1902a: 684. (Guatemala) 

3. Ersachus mexicanus Sharp, 1902a: 684. (Mexico) 

4. Ersachus variegatus Erichson, 1847: 497. (Colombia) 

Genus Eulimnichus Casey, 1889 

Eulimnichus Casey, 1889: 146.

1. Eulimnichus acutus Wooldridge, 1979: 7. (Costa Rica, Colombia, El Salvador, Guatemala, Honduras, Mexico, Panama) 

2. Eulimnichus analis (LeConte), 1879: 515. (Mexico, Panama, USA) (syn., analis rugiceps Casey, 1912: 47) (syn., analis coloradensis Casey, 1912: 47) 

3. Eulimnichus ater (LeConte), 1854: 117. (Argentina, Belize, Bolivia, Brazil, British Guiana, Colombia, Costa Rica, Cuba, Ecuador, Guatemala, Honduras, Jamaica, Mexico, Panama, Peru, USA, Venezuela) (syn., guatemalicus Sharp, 1902a: 676) (syn., sculpticeps Casey, 1912: 50) (syn., Limnichoderus oblongus Pic, 1922: 4) 

4. Eulimnichus californicus (LeConte), 1879: 515. (USA) 

5. Eulimnichus coheni Wooldridge, 1979: 4. (Bolivia, Ecuador) 

6. Eulimnichus confertus Sharp, 1902a: 676. (Mexico, Guatemala) 

7. Eulimnichus corrineae Wooldridge, 1979: 3. (Argentina, Brazil, Venezuela) 

8. Eulimnichus epistemus Sharp, 1902a: 673. (Costa Rica, Guatemala, Nicaragua, Mexico) 

9. Eulimnichus evanescens Casey, 1912: 48. (USA) 

10. Eulimnichus expeditus Wooldridge, 1984b: 57. (Guatemala, Honduras) 

11. Eulimnichus impostus Wooldridge, 1978: 172. (Brazil) (syn., elongatus Pic, 1923b: 7; preoccupied) 

12. Eulimnichus improcerus Wooldridge, 1979: 4. (USA) 

13. Eulimnichus incultus Wooldridge, 1979: 6. (Bolivia, Ecuador) 

14. Eulimnichus langleyae Wooldridge, 1979: 3. (Ecuador, Peru) 

15. Eulimnichus montanus (LeConte), 1879: 514. (Mexico, USA) (syn., laeticulus Sharp, 1902a: 674) 

16. Eulimnichus nitidulus (LeConte), 1854: 117. (Argentina, Brazil, Costa Rica, Ecuador, Mexico, Nicaragua, Panama, USA) 

17. Eulimnichus obscurus (LeConte), 1854: 116. (USA) 

18. Eulimnichus optatus Sharp, 1902a: 675. (Guatemala, Mexico, USA) 

19. Eulimnichus pellucidus Wooldridge, 1979: 5. (Venezuela) 

20. Eulimnichus perpolitus Casey, 1889: 151. (USA) 

21. Eulimnichus plebius Sharp, 1902a: 675. (Argentina, Brazil, Colombia, Dominican Republic, Ecuador, El Salvador, Guatemala, Haiti, Honduras, Mexico, Puerto Rico, Peru, Trinidad, Venezuela) (syn., Limnichoderus convexus Pic, 1922: 4) (syn., Limnichoderus breyeri Pic, 1930: 43) (syn., Limnichoderus peruvianus Pic, 1954: 178) 

22. Eulimnichus rugolosus Wooldridge, 1979: 6. (USA) 

23. Eulimnichus rusticus Wooldridge, 1979: 6. (Mexico) 

24. Eulimnichus sharpi Wooldridge, 1979: 5. (Mexico) 

25. Eulimnichus sordidus Sharp, 1902a: 676. (Guatemala) 

26. Eulimnichus spangleri Wooldridge, 1979: 8. (El Salvador, Guatemala, Honduras, Mexico) 

27. Eulimnichus subtitus Wooldridge, 1979: 8. (USA) 

28. Eulimnichus sublaevis Sharp, 1902a: 674. (Guatemala, Panama) 

29. Eulimnichus visendus Wooldridge, 1979: 1. (Mexico) 

Genus Jaechobyrrhinus Pütz, 1991 

Jaechobyrrhinus Pütz, 1991a: 132.

1. Jaechobyrrhinus amanosius Pütz, 1991a: 132. (Turkey) 

Genus Lichminus Casey, 1889 

Lichminus Casey, 1889: 147.

1. Lichminus tenuicornis Casey, 1889: 155. (USA) 

Genus Phalacrichus Sharp, 1902 

Phalacrichus Sharp, 1902a: 678.

Limnichalia Casey, 1912: 52.

1. Phalacrichus atomarinus Sharp, 1902a: 679. (Beize, Bolivia, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Surinum, Trinidad, Venezuela) (syn., colonica Casey, 1912: 52) 

2. Phalacrichus diligens Wooldridge, 1982: 389. (Bolivia, Brazil, Colombia, Venezuela) 

3. Phalacrichus durus Wooldridge, 1982: 387. (Bolivia, Brazil, Ecuador, Paraguay, Peru, Venezuela) 

4. Phalacrichus golbachi Wooldridge, 1982: 388. (Paraguay) 

5. Phalacrichus latus Wooldridge, 1982: 386. (Brazil, Colombia, Ecuador, Peru, Venezuela) 

6. Phalacrichus punctatus Wooldridge, 1982: 389. (Bolivia, Brazil, Colombia, Panama, Venezuela) 

7. Phalacrichus rudis Wooldridge, 1982: 385. (Surinam, Venezuela) 

8. Phalacrichus simplex Wooldridge, 1982: 387. (Bolivia, Brazil, Venezuela) 

9. Phalacrichus spangleri Wooldridge, 1982: 386. (Ecuador, Venezuela) 

10. Phalacrichus sublimus Wooldridge, 1993a: 36. (Venezuela) 

11. Phalacrichus vianai (Pic), 1938: 77. (Argentina, Brazil, British Guiana, Trinidad) 

Genus Physemus LeConte, 1854 

Physemus LeConte, 1854: 117.

Ditaphrus Casey, 1886: 250.

1. Physemus excavatus Wooldridge, 1976: 182. (El Salvador, Guatemala, Honduras) 

2. Physemus latifrons Wooldridge, 1976: 180. (Panama) 

3. Physemus levis Wooldridge, 1984a: 538. (Brazil, Trinidad, Venezuela) 

4. Physemus minutus LeConte, 1854a: 117. (Mexico, USA) (syn., Ditaphrus scymnoides Casey, 1886: 250) 

5. Physemus mirus Wooldridge, 1984a: 538. (Ecuador) 

6. Physemus punctatus Wooldridge, 1976: 180. (Colombia) 

Genus Tricholimnichus Hernando & Ribera, 2001 

Tricholimnichus Hernando & Ribera, 2001: 153.

1. Tricholimnichus maior Hernando & Ribera, 2001: 159. (Malaysia) 

2. Tricholimnichus minor Hernando & Ribera, 2001: 160. (Malaysia) 

3. Tricholimnichus sabahensis Hernando & Ribera, 2001: 160. (Malaysia)