Family Phryganeidae Leach
Author: Takao NOZAKI
E-mail: takao.nozaki[AT]nifty.com
Last update: 11 October 2026
Subfamily Phryganeinae Burmeister
Genus Agrypnia Curtis
Larvae recorded as a European species, Phryganea varia, by Iwata (1927a, 1927b) undoubtedly belong to this genus, but cannot be identified as a species (Wiggins 1998). A. colorata Hagen recorded by Kuwayama (1973) as Dasystegia principalis (Martynov) is a misidentification of A. acristata or A. sordida (Wiggins 1998).
Agrypnia acristata Wiggins 1998
Dasystegia ulmeri (non Martynov 1909): Kuwayama 1973
Distribution. Japan (Hokkaido, Honshu, Kyushu), Sakhalin, Kuriles.
Agrypnia incurvata Wiggins 1998
Distribution. Japan (Honshu, Shikoku, Kyushu, Sado), Korea.
Distribution. Japan (Hokkaido, Honshu), Sakhalin, Kuriles, Russia, Mongolia, Tibet, China, Korea, Europe.
Agrypnia sordida (McLachlan 1871)
Phryganea sordida McLachlan 1871
Dasystegia (Prophryganea) umbrina Martynov 1924
Prophryganea sordida: Matsumura 1931
Agrypnia umbrina: Martynov 1935
Phryganea (Prophryganea) sordida: Tsuda 1942a
Dasystegia (Prophryganea) sordida: Fischer 1964
Dasystegia sordida: Kuwayama 1973
Dasystegia umbrina: Nozaki and Itou 1988
Distribution. Japan (Hokkaido, Honshu, Shikoku, Sado), Sakhalin, Kuriles, Russia (continental part of Far East), Korea.
Agrypnia ulmeri (Martynov 1909)
Phryganea ulmeri Martynov 1909
Phryganea ulmerina Navás, 1920 ("1919")
Dasystegia (Prophryganea) ulmeri: Fischer 1964
Distribution. Japan (Hokkaido, Honshu), Russia (Ussuri, Siberia).
Genus Eubasilissa Martynov
Eubasilissa imperialis (Nakahara 1915)
Phryganea imperialis Nakahara 1915
Phryganea regina ab. imperialis: Kuwayama 1922
Distribution. Japan (Honshu, Oki).
Eubasilissa regina (McLachlan 1871)
Holostomis maclachlani var. regina (an sp. distincta?) McLachlan 1871
Neuronia (Holostomis) regina: McLachlan 1894
Holostomis regina: Matsumura 1904
Neuronia regina: Ulmer 1907a
Neuronia reginella Nakahara 1913a
Phryganea regina ab. reginella: Kuwayama 1922
Regina regina: Martynov 1924
Phryganea regina: Esaki 1930
Neuronia (Eubasilissa) regina: Tsuda 1942a
Neuronia maxima Iwata 1927a
Eubasilissa reginella: Kuwayama 1970
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Sado, Oki, Tsushima), Sakhalin, Kuriles, China, Korea, Taiwan.
Genus Hagenella Martynov
Hagenella apicalis (Matsumura 1904)
Neuronia apicalis Matsumura 1904
Neuronia dentata Martynov 1909
Neuronia (Hagenella) apicalis: Kuwayama 1949
Distribution. Japan (Hokkaido, Yururi-to), Sakhalin, Kuriles.
Genus Oligostomis Kolenati
Oligostomis wigginsi Nishimoto and Kawase 2005
Distribution. Japan (Honshu).
Genus Oligotricha Rambur
Oligotricha fluvipes (Matsumura 1904)
Neuronia fluvipes Matsumura 1904
Neuronia (Oligostomis) fluvipes
Oligostomis fluvipes: Matsumura 1931
Oligostomis hirayamai Matsumura 1931
Distribution. Japan (Hokkaido?, Honshu, Shikoku).
Oligotricha hybridoides Wiggins and Kuwayama 1971
Oligotricha fluvipes (non Matsumura 1904): Kuwada 1965, 165
Distribution. Japan (Hokkaido), Sakhalin, Kuriles.
Oligotricha kawamurai (Iwata 1927)
Neuronia kawamurai Iwata 1927a
Distribution. Japan (Honshu).
Oligotricha spicata Wiggins and Kuwayama 1957
Distribution. Japan (Honshu).
Genus Phryganea Linnaeus
Phryganea japonica McLachlan 1866
Phryganea (Colpomera) japonica: Martynov 1924a, 213.
Colpomera japonica: Kuwayama 1973
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki, Tsushima), South Kuriles, Korea, China.
Genus Semblis Fabricius
Semblis melaleuca (McLachlan 1871)
Holostomis melaleuca McLachlan 1871
Neuronia (Holostomis) melaleuca: Tsuda 1942a
Neuronia melaleuca: Nakahara 1913b
Holostomis chishimana Kuwayama 1967
Semblis phalaenoides: Tanida 1982, 359, Honshu (Gumma). Misidentification mentioned by Tanida (2019).
Distribution. Japan (Hokkaido, Honshu), Sakhalin, Kuriles.
Semblis phalaenoides (Linnaeus 1758)
Phryganea phalaenoides Linnaeus 1758
Holostomis phalaenoides: Matsumura 1931
Neuronia (Holostomis) phalaenoides: Kuwayama
Neuronia phalaenoides: Navás 1920
Holostomis coreana Kuwayama 1967
Distribution. Japan (Hokkaido, Honshu), Russia (European, Siberia, Far East, Sakhalin), China (north), Korea, Mongolia, Europe (north).