Family Philopotamidae Stephens
Author: Naotoshi KUHARA
E-mail: naotoshi.kuhara[AT]nifty.com
Last updated: 4 October 2026
Subfamily Chimarrinae Rambur
Genus Chimarra Stephens
The larva tentalively described by Furuya (1974) as Chimarra sp. CB belongs to the genus Wormaldia (Kuhara 2017).
Chimarra (Chimarra) tsudai Ross 1956
Chimarra concolor non Ulmer 1905: Tsuda 1939b, 1941, 1942a, 1943
Chimarra sp. CA: Kumode 1952
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki-shoto, Tsushima, Koshikijima-retto, Ryukyu (Yakushima)), Korea.
Chimarra species indeterminate
Subfamily Philopotaminae Stephens
Genus Dolophilodes Ulmer
The larva tentalively described by Tanida (1985, 2005) as Dolophilodes sp. DC belongs to the genus Wormaldia (Kuhara 2017).
Dolophilodes angustata Kuhara 2005
Dolophilodes sp. DA: Tanida 1985
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu).
Dolophilodes auriculata Martynov 1933
Sortosa auriculata: Ross 1956
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima).
Dolophilodes babai (Kobayashi 1980)
Sortosa babai Kobayashi 1980
Distribution. Japan (Honshu (central)).
Dolophilodes commata (Kobayashi 1980)
Sortosa commata Kobayashi 1980
Sortosa kaishoensis Kobayashi 1985a
Distribution. Japan (Honshu, Shikoku, Kyushu, Tsushima, Oki-shoto, Ryukyu (Yakushima, Tanegashima)).
Dolophilodes dilatata Kuhara 2005
Distribution. Japan (Hokkaido, Honshu).
Dolophilodes iroensis (Kobayashi 1980)
Sortosa iroensis Kobayashi 1980
Dolophilodes sp. DD: Tanida 1985
Dolophilodes kunashirensis Ivanov 1996 (in Arefina et al. 1996)
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Ryukyu (Yakushima)), Russia (southern and northern Kurils).
Dolophilodes japonica (Banks 1906)
Philopotamus japonica Banks 1906
Dolophilodes exscisa Martynov 1933
Sortosa japonica: Ross 1956
Dolophilodes sp. DB: Furuya 1974
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Sado-ga-shima), Russia (Sakhalin, southern Kurils).
Dolophilodes nomugiensis (Kobayashi 1980)
Sortosa nomugiensis: Kobayashi 1980
Wormaldia triangulata Kobayashi 1984a
Wormaldia sp. WA: Tanida 1985
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Rishiri-to, Rebun-to, Oki-shoto), Russia (Kamchatka, Sakhalin, Kurils).
Dolophilodes shinboensis (Kobayashi 1980)
Sortosa shinboensis Kobayashi 1980
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Rishiri-to).
Dolophilodes species indeterminate
Genus Kisaura Ross
Kisaura ashizuriensis Kuhara 2019
Distribution. Japan (Shikoku).
Kisaura borealis (Kuhara 1999)
Dolophilodes borealis Kuhara 1999
Distribution. Japan (Hokkaido, Honshu (north to central)), Russia (Sakhalin, southern Kurils).
Distribution. Japan (Honshu, Shikoku).
Kisaura curvispina Kuhara 2019
Distribution. Japan (Shikoku).
Kisaura dichotoma Kuhara and Arefina 2004
Dolophilodes sp.: Kuhara 1997a
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Rebun-to), Russia (southern Kurils).
Kisaura hattorii (Kuhara 1999)
Dolophilodes hattorii Kuhara 1999
Distribution. Japan (Hokkaido, Honshu (north to central)), Russia (southern Kurils).
Kisaura kisoensis (Tsuda 1939)
Dolophilodes kisoensis Tsuda 1939b
Sortosa kisoensis: Ross 1956
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima).
Kisaura minakawai Arefina 2005 (in Arefina and Armitage 2005)
Kisaura imparis Hur and Morse 2006
Distribution. Japan (Hokkaido, Honshu, Kyushu, Ryukyu (Yakushima, Tanegashima)), Russia (Sakhalin).
Kisaura niitakaensis (Kobayashi 1973)
Sortosa niitakaensis Kobayashi 1973
Distribution. Japan (Hokkaido, Honshu).
Dolophilodes nozakii Kuhara 1999
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu).
Kisaura tsudai (Botosaneanu 1970)
Dolophilodes tsudai Botosaneanu 1970
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu, Ryukyu (Yakushima)), Russia (southern Kurils).
Kisaura nozakii species complex : Suzuki et al. 2023
Genus Wormaldia McLachlan
Larval records of Wormaldia subnigra in Japan by Iwata (1927a, 1927b) were considered to be misidentification of other species (Tsuda 1942b, 1958, Kuhara 2005b).
Wormaldia amamiensis Kuhara 2016
Distribution. Japan (Amami-oshima).
Wormaldia apophysis Kuhara 2016
Distribution. Japan (Amami-oshima, Tokuno-shima).
Wormaldia carinata (Schmid 1991)
Doloclanes carinata Schmid 1991
Distribution. Japan (Amami-oshima, Okinawa-jima).
Wormaldia fujinoensis Kobayashi 1980
Distribution. Japan (Honshu (central, west)).
Wormaldia ishigakiensis Kuhara 2016
Distribution. Japan (Ishigaki-jima).
Distribution. Japan (Ryukyu (Yakushima)).
Wormaldia kadowakii Kobayashi 1980
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu).
Wormaldia kisoensis (Tsuda 1942)
Naganagapetus kisoensis Tsuda 1942a
Distribution. Japan (Honshu (central), Shikoku).
Wormaldia nabewarina Kobayashi 1969
Distribution. Japan (Honshu (central), Shikoku, Kyushu, Koshikijima-retto).
Wormaldia niiensis Kobayashi 1985
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Tsushima, Koshikijima-retto, Ryukyu (Yakushima)), Russia (South Primorye), Korea, China (Liaoning), Taiwan.
Wormaldia okinawaensis Kuhara 2016
Distribution. Japan (Okinoerabu-jima, Okinawa-jima).
Wormaldia rara (Kobayashi 1959)
Dolophilodes rara Kobayashi 1959
Wormaldia kurokawana Kobayashi 1968
Sortosa rara: Fischer 1971
Wormaldia saekiensis Kobayashi 1980
Wormaldia yunotakiensis Kobayashi 1980
Distribution. Japan (Honshu (central to west), Shikoku, Kyushu).
Wormaldia sumuharana Kobayashi 1980
Distribution. Japan (Honshu (central)).
Distribution. Japan (Okinawa-jima).
Wormaldia uonumana Kobayashi 1980
Distribution. Japan (Hokkaido, Honshu, Shikoku).
Wormaldia yakuensis Kobayashi 1980
Distribution. Japan (Kyushu, Fukue-jima, Ryukyu (Yakushima)).
Wormaldia species indeterminate