Family Leptoceridae Leach
Author: Nobuyuki KATSUMA
E-mail: setodesujiensisakagi1960[AT]gmail.com
Last update: 5 May 2026
Subfamily Leptocerinae Leach
Genus Adicella McLachlan
Adicella longiramosa Yang and Morse 2000
Distribution. Japan (Honshu), China.
Adicella makaria Malicky and Chantaramongkol 2002
Distribution. Ishigaki-jima, Iriomote-jima, Yonaguni-jima, Taiwan.
Adicella odamiyamensis Katsuma and Yamamoto 2015
Distribution. Japan (Shikoku, Yaku-shima).
Adicella paludicola Ito and Kuhara 2013
Distribution. Japan (Hokkaido, Honshu).
Adicella strigillata Katsuma and Ito 2013
Distribution. Japan (Honshu).
Adicella trichotoma Ito and Kuhara 2013
Distribution. Japan (Hokkaido, Honshu, Shikoku, Amami-oshima, Okinawa-jima).
Genus Athripsodes Billberg
Athripsodes tsudai (Akagi 1960)
Leptocerus sp. LA: Akagi 1953
Leptocerus tsudai Akagi 1960
Ceraclea tsudai: Tanida 1985
Leptocerus seikunis Kobayashi 1987
Distribution. Japan (Honshu).
Genus Ceraclea Stephens
Two unnamed species accompanied with an alphabetic code were described for the larvae, which have not yet associated with adults. Tanida (1997, 2003) recorded one undescribed species from Ryukyu Islands. Yamamoto and Nozaki (2025) recorded an unidentified species similar to a Chinese species C. parakamonis Yang and Morse 2000.
Ceraclea albimacula (Rambur 1842)
Mystacida albimacula Rambur 1842
Leptocerus alboguttatus Hagen 1860
Leptocerus spinosus Tsuda 1942b
Leptocerus biwaensis: Tsuda and Kuwayama 1950
Athripsodes albimacula: Fischer 1965
Athripsodes alboguttatus: Fischer 1965
Ceraclea biwaensis: Morse 1975
Ceraclea alboguttata: Morse 1975
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), Europe, China, North and South Korea.
Ceraclea complicata (Kobayashi 1984)
Leptocerus complicatus Kobayashi 1984a
Leptocerus funashiensis Kobayashi 1985
Ceraclea miyakonis (non Tsuda 1942): Nozaki 1987
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki-no-shima, Tsushima, Yaku-shima), Russia (Sakhalin).
Ceraclea coreana Kumanski 1991
Distribution. Japan (Kyushu), Russia (Primorskii krai, Khabarovskii krai), North and South Korea.
Ceraclea hastata (Botosaneanu 1970)
Athripsodes hastatus Botosaneanu 1970
Distribution. Japan (Honshu), Russia (Far East), North Korea, Mongolia.
Leptocerus kamonis Tsuda 1942b
Athripsodes kamonis: Fischer 1965
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu).
Ceraclea lobulata (Martynov 1935)
Leptocerus lobulatus Martynov 1935
Leptocerus miyakonis Tsuda 1942b
Athripsodes lobulatus: Fischer 1965
Athripsodes miyakonis: Fischer 1965
Ceraclea miyakonis: Morse 1975
Distribution. Japan (Honshu, Shikoku), Russia (Amur Region), China, South Korea.
Leptocerus mitis Tsuda 1942b
Athripsodes mitis: Fischer 1965
Leptocerus takatsunis Kobayashi 1987
Distribution. Japan (Honshu, Shikoku), South Korea.
Ceraclea nigronervosa (Retzius 1783)
Phryganea nigronervosa Retzius 1783
Leptocerus nigronervosus: Tsuda 1942b
Athripsodes nigronervosus: Fischer 1965
Distribution. Japan (Honshu), Europe, Russia.
Leptocerus superbus Tsuda 1942b
Athripsodes superbus: Fischer 1965
Distribution. Japan (Honshu), Russia (Prymorskii krai, Khabarovskii krai) China.
Ceraclea valentinae Arefina 1997
Distribution. Japan (Hokkaido), Russia (south Kuril Islands).
Ceraclea variabilis (Martynov 1935)
Leptocerus variabilis Martynov 1935
Athripsodes variabilis: Fisher 1965
Distribution. Japan (Kyushu), Russia (Amur Region).
Leptocerus sp. LB: Akagi 1957
Leptocerus sp. LC: Akagi 1959
Athripsodes sp. LC: Tsuda and Akagi 1974
Ceraclea species indeterminate
Genus Erotesis McLachlan
Only one species, E. japonica Iwata 1930 was described from Honshu based only on the larval stage. Uenishi et al. (1994) recorded an unidentified species based on adult material from Kyushu.
Distribution. Japan (Honshu).
Genus Leptocerus Leach
Setodes biwae Tsuda 1942b
Distribution. Japan (Hokkaido, Honshu, Kyushu), Russia (Primorye), China.
Leptocerus fluminalis Ito and Kuhara 2009
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu).
Leptocerus moselyi (Martynov 1935)
Setodes moselyi Martynov 1935
Distribution. Japan (Hokkaido, Honshu, Kyushu), Russia (Amur region).
Leptocerus valvatus (Martynov 1935)
Setodes valvata Martynov 1935
Leptocerus utonaiensis Kobayashi 1977
Distribution. Japan (Hokkaido, Honshu, Kyushu), Russia (Amur Region).
Genus Mystacides Berthold
Two unnamed species accompanied with an alphabetic code were described for the larvae, which were not associated with adults. Tsuda (1942b) recorded Mystacides longicornis (Linnaeus) from Japan, but Uenishi (1993) suggested that it was a misidentification of an undescribed species.
Mystacides azureus (Linnaeus 1761)
Phryganea azurea: Linnaeus 1761
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Rebun-to, Sado-ga-shima, Oki-no-shima, Tsushima, Fukue-jima, Koshiki-jima, Yaku-shima), Russia, Europe.
Mystacides bifidus Martynov 1924
Distribution. Japan (Honshu), Russia (Amur Region).
Mystacides moritai Kuhara, Nozaki, Zhang and Zhou 2023
Distribution. Japan (Honshu).
Distribution. Japan (Hokkaido), Russia (Sakhalin).
Mystacides rivularis Kuhara, Nozaki, Zhang and Zhou 2023
Distribution. Japan (Hokkaido, Honshu).
Mystacides sibiricus Martynov 1935
Distribution. Japan (Hokkaido), Russia (Amur Region), China.
Mystacides sp. MA: Tsuda and Akagi 1962
Mystacides sp. MB: Tsuda and Akagi 1962
Genus Oecetis McLachlan
Kuwayama (1930) reported a European species, Oecetis testcea (Curtis 1984), based on two female specimens collected from Hokkaido. After that, Kumanski (1991) described a subspecies, O. testacea orientalis (now treated as O. kumanskii), based on specimens collected from North Korea. Furthermore, Arefina and Armitage (2005) described other subspecies, O. testacea sakhalinica (junior synonym of O. tsudai), from South Sakhalin. Both O. kumanskii and O. tsudai are distributed in Hokkaido, therefore, Kuwayama's female specimens may belong to either of those species. On the other hand, although Tanida (2016) listed O. testacea testacea in his checklist of Japanese Trichoptera, no reliable record of this nominotypical subspecies is found from East Asia.
Tanida (1997) recorded one undescribed species from Ryukyu Islands. O. dilata Yang and Morse 2000 was recorded from Hokkaido without collecting data by Malicky (2005b).
Oecetis antennata (Martynov 1935)
Oecetodella antennata Martynov 1935
Distribution. Japan (Honshu), Russia (Ussuri), China.
Oecetis brachyura Yang and Morse 1997
Distribution. Japan (Hokkaido, Honshu, Kyushu, Yaku-shima), Russia (Far East), China.
Oecetis caucula Yang and Morse 2000
Distribution. Japan (Honshu, Shikoku, Kyushu), South Korea, China.
Mystacida furva Rambur 1842, 512, France.
Distribution. Japan (Honshu), Russia (South Khabarovsk, Prymorskii krai), Europe.
Oecetis hamochiensis Kobayashi 1984
Oecetis dilata non Yang and Morse 2000: Nozaki and Nakamura 2002
Oecetis ochracea non Curtis 1825: Kawase and Morita 2010
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Sado-ga-shima, Oki-no-shima).
Oecetis kumanskii Yang and Morse 2000
Oecetis testacea orientalis Kumanski 1991
Oecetis testacea kumanskii Yang and Morse, 2000
Distribution. Japan (Hokkaido), North Korea.
Distribution. Japan (Hokkaido, Honshu, Kyushu, Rebun-to), Russia (Prymorskii krai), China.
Oecetis nigropunctata Ulmer 1908
Oecetis pallidipunctata Martynov 1935, 207, 249-253, male, Russia (Amur Region). Synonymized by Kumanski (1991).
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Koshiki-jima, Ishigaki-jima), Russia (Amur Region, Sakhalin), North and South Korea, China, Taiwan, Vietnam, Laos.
Distribution. Japan (Honshu).
Distribution. Ryukyu Island (Yonaguni-jima), Taiwan, China, Jawa.
Oecetis tripunctata (Fabricius 1793)
Phryganea tripunctata Fabricius 1793, 81, Germany.
Distribution. Japan (Hokkaido, Honshu), North Korea, Russia, Europe.
Oecetis orientalis Tsuda 1942b
Oecetis testacea sakhalinica Arefina 2005
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Oki-no-shima, Yaku-shima).
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), Russia (Far East), North and South Korea.
Genus Parasetodes McLachlan
Parasetodes respersellus (Rambur 1842)
Setodes respersella Rambur 1842
Distribution. Japan, Russia (Amur Region), Egypt, Europe.
Genus Setodes Rambur
Two unnamed species accompanied with an alphabetic code were described for the larvae, which were not associated with adults. Aoyagi (2025) recorded an unidentified species from Ishigaki-jima.
Setodes argentatus Matsumura 1907
Setodes iris: Matsumura 1927
Setodes appendiculata Martynov 1933
Oecetis turbata Navas 1933
Setodes uenoi Tsuda 1942a
Setodes turbatus: Schmid 1950
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), Russia (Far East), North Korea, China.
Setodes chirotheca Katsuma 2018
Distribution. Japan (Hokkaido, Honshu).
Setodes hinumaensis Katsuma 2009
Distribution. Japan (Honshu).
Setodes sp. SA: Akagi 1957
Setodes curviseta Kobayashi 1959
Distribution. Japan (Honshu, Shikoku, Kyushu).
Distribution. Japan (Honshu, Shikoku).
Setodes pseudofurcatulus Katsuma 2018
Distribution. Japan (Honshu).
Setodes shirasensis Kobayashi 1984
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu, Sado-ga-shima).
Trichosetodes ujiensis Akagi 1960
Distribution. Japan (Honshu), North Korea.
Setodes sp. SB: Tsuda and Akagi 1962
Genus Triaenodes McLachlan
Three species belonging to this genus have been recorded from Japan. Furthermore, one unnamed species accompanied with an alphabetic code were described for the larvae, which were not associated with adults. Tanida (1997, 2003) recorded one unidentified species from Yaeyama Islands.
Triaenodes pellectus Ulmer 1908
Triaenodella gracillima Martynov 1935
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), South Kuriles, Russia (Amur Region), China, Borneo, Laos, Malaysia, Sumatra, Thailand, Vietnam.
Triaenodes qinglingensis Yang and Morse 2000
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), China, Laos, Thailand.
Triaenodes unanimis McLachlan 1877
Triaenodes niwai Iwata 1927
Triaenodes yamamotoi Tsuda 1942b
Distribution. Japan (Hokkaido, Honshu, Shikoku, Kyushu), South Kuriles, Russia (Amur Region), Korea, China, Europe.
Triaenodes sp. TA: Tsuda and Akagi 1962
Genus Trichosetodes Ulmer
Trichosetodes japonicus Tsuda 1942
Trichosetodes polonorum Botosaneanu 1970
Distribution. Japan (Honshu, Shikoku, Kyushu), Russia (Far East), North Korea.
Subfamily Triplectidinae Ulmer
Genus Triplectides Kolenati
Triplectides misakianus (Matsumura 1931)
Tobikera misakiana Matsumura 1931
Triplectides magna non Walker 1842: Tsuda, 1942b
Notanatolica magna (non Walker 1842): Tsuda, 1959
Distribution. Japan (Honshu, Kyushu).