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Soil Evolution
  • Home
    • Start
      • Soil & Civilisation
      • Seeing Soil
      • Soil Science
      • New Science
      • Short story
    • What is Soil?
      • Clay
      • Soil Structure
      • Biome
      • Glomalisation
        • Testing
      • Soil Functions
        • Energy
          • Entropy
      • Decomposition
        • Mineralisation
        • De-lignification
        • Humification
      • Types
        • Europe
    • Challenge
      • Terrestrialisation
      • Theories so far
      • Tools
    • Darwin's version
    • Timeline
      • Copy of 100mya - 0 mya
      • Copy of 200-100 mya
      • Copy of 300-200 mya
      • Copy of 400-300 mya
      • Copy of 500-400 mya
  • 500-400 mya
    • No Soil
    • 4.500 - 1000 mya
    • 1000 - 500 mya
    • Periods
      • Cambrian
      • Ordovician
      • Silurian
    • Biology
      • Plants
      • Animals
      • Bacteria
  • 400-300 mya
    • 400-360 mya Late Devonian
      • Green cover
      • Vascular Plants
      • Mycorrhiza (AMF)
      • Animals
        • Springtails
        • Arachnids
    • 360-300mya Carboniferous
      • Plants
        • Vascular
      • Early Soils
        • Micro-aggregation
      • Animals - Early Carb
        • Oribatids - Lower
        • Origin of Insects
      • Animals - Late Carb
      • Worms
  • 300-200 mya
  • 200-100 mya
    • 200-145 mya Jurassic
    • 145-66 mya Cretaceous
  • 100mya - 0 mya
    • 66 - 0 mya Cenozoic
  • Now
    • Present State of Soil
      • Desertification
      • Concretisation
      • Globalisation
    • Practices affecting Soil
      • Chemical
        • Fertilisers
        • Carbon
        • Pesticides
      • Problem
      • Biological
    • Soil & Global Warming
      • Soil Surfaces & Global Warming
      • Soil Carbon
      • Soil & Water
      • Soil Temperature
      • Soil Biota
      • Climate Change
    • Save our Soil!
      • Soil Health
      • Regenerate
      • Ecology
      • Economics
Soil Evolution
  • Home
    • Start
      • Soil & Civilisation
      • Seeing Soil
      • Soil Science
      • New Science
      • Short story
    • What is Soil?
      • Clay
      • Soil Structure
      • Biome
      • Glomalisation
        • Testing
      • Soil Functions
        • Energy
          • Entropy
      • Decomposition
        • Mineralisation
        • De-lignification
        • Humification
      • Types
        • Europe
    • Challenge
      • Terrestrialisation
      • Theories so far
      • Tools
    • Darwin's version
    • Timeline
      • Copy of 100mya - 0 mya
      • Copy of 200-100 mya
      • Copy of 300-200 mya
      • Copy of 400-300 mya
      • Copy of 500-400 mya
  • 500-400 mya
    • No Soil
    • 4.500 - 1000 mya
    • 1000 - 500 mya
    • Periods
      • Cambrian
      • Ordovician
      • Silurian
    • Biology
      • Plants
      • Animals
      • Bacteria
  • 400-300 mya
    • 400-360 mya Late Devonian
      • Green cover
      • Vascular Plants
      • Mycorrhiza (AMF)
      • Animals
        • Springtails
        • Arachnids
    • 360-300mya Carboniferous
      • Plants
        • Vascular
      • Early Soils
        • Micro-aggregation
      • Animals - Early Carb
        • Oribatids - Lower
        • Origin of Insects
      • Animals - Late Carb
      • Worms
  • 300-200 mya
  • 200-100 mya
    • 200-145 mya Jurassic
    • 145-66 mya Cretaceous
  • 100mya - 0 mya
    • 66 - 0 mya Cenozoic
  • Now
    • Present State of Soil
      • Desertification
      • Concretisation
      • Globalisation
    • Practices affecting Soil
      • Chemical
        • Fertilisers
        • Carbon
        • Pesticides
      • Problem
      • Biological
    • Soil & Global Warming
      • Soil Surfaces & Global Warming
      • Soil Carbon
      • Soil & Water
      • Soil Temperature
      • Soil Biota
      • Climate Change
    • Save our Soil!
      • Soil Health
      • Regenerate
      • Ecology
      • Economics
  • More
    • Home
      • Start
        • Soil & Civilisation
        • Seeing Soil
        • Soil Science
        • New Science
        • Short story
      • What is Soil?
        • Clay
        • Soil Structure
        • Biome
        • Glomalisation
          • Testing
        • Soil Functions
          • Energy
            • Entropy
        • Decomposition
          • Mineralisation
          • De-lignification
          • Humification
        • Types
          • Europe
      • Challenge
        • Terrestrialisation
        • Theories so far
        • Tools
      • Darwin's version
      • Timeline
        • Copy of 100mya - 0 mya
        • Copy of 200-100 mya
        • Copy of 300-200 mya
        • Copy of 400-300 mya
        • Copy of 500-400 mya
    • 500-400 mya
      • No Soil
      • 4.500 - 1000 mya
      • 1000 - 500 mya
      • Periods
        • Cambrian
        • Ordovician
        • Silurian
      • Biology
        • Plants
        • Animals
        • Bacteria
    • 400-300 mya
      • 400-360 mya Late Devonian
        • Green cover
        • Vascular Plants
        • Mycorrhiza (AMF)
        • Animals
          • Springtails
          • Arachnids
      • 360-300mya Carboniferous
        • Plants
          • Vascular
        • Early Soils
          • Micro-aggregation
        • Animals - Early Carb
          • Oribatids - Lower
          • Origin of Insects
        • Animals - Late Carb
        • Worms
    • 300-200 mya
    • 200-100 mya
      • 200-145 mya Jurassic
      • 145-66 mya Cretaceous
    • 100mya - 0 mya
      • 66 - 0 mya Cenozoic
    • Now
      • Present State of Soil
        • Desertification
        • Concretisation
        • Globalisation
      • Practices affecting Soil
        • Chemical
          • Fertilisers
          • Carbon
          • Pesticides
        • Problem
        • Biological
      • Soil & Global Warming
        • Soil Surfaces & Global Warming
        • Soil Carbon
        • Soil & Water
        • Soil Temperature
        • Soil Biota
        • Climate Change
      • Save our Soil!
        • Soil Health
        • Regenerate
        • Ecology
        • Economics

 500 - 400 mya

No Soil  4500-1000mya 1000-500mya 

Cambrian   Ordovician  Silurian         Microbes  Plants  Animals 


500 - 400mya

There was no soil half a billion (500mya)  years ago, although we have seen in the previous 500 million years, metazoan creatures. However, the first signs of soil may have been emerging during these next 100 million years developed via biofilm.

Biofilms
Liverworts
Moss - Takakia 
Rhniophytes
Cambrian           Ordovician       Silurian
Retallack 2022
Silurian                                                           Devonian
Royal Society graphic 
Tardigrades

500mya 

Entomobryoid Springtails
Nematodes
Millipedes

Wet

We tend to describe the creation of life on land as 'conquering the dry conditions'. This is true. But It may well be better to consider the 'extension of wet conditions' too. This would be via thin water films - biofilms. There would be wierd water tensions, and those that can cope with that can go further inland. It is these biofilms that holds the rest together. Bacteria would be in them as well as tiny nematode worms. The biofilm thus becomes an environment of its own, somewhere other organisms could live. The more water the better it does that. After the average composition of living organisms is 65-90% water!

Healthy soil today is about 1/4 air and 1/4 water…but we have a long way to get there.

Dry

The big development over these 100 million years were the number of plants and creatures that could survive both wet and dry; this combination is essential for characters to live in soil. There are often long dry periods between the wet ones, so particular mechanisms have to be in place. 

In particular, mosses which grow in moist conditions can withstand long periods of drought too. They increasingly covered much of the Earth, and housed the key characters, thereby creating the conditions for soil formation. We'll explore the next great soil development in the next phase.

Protozoans are the first visible organisms to emerge from their cysts, the protective cases they produced when the moss dried out. Twenty minutes later, the single cells have been joined by some tiny animals that have come out of dormancy. We can also find there rotifers, the smallest members of the animal kingdom. The little beasts appear to have two sets of rotating wheels—an illusion produced by tufts of cilia that beat in a wave - which gives these animals their name, Rotifers.

Biofilms  for billion years

A biofilm is a community of microorganisms that attach to each other and to a surface, and are embedded in a slimy matrix Biofilms will carry bacteria and algae to live on land., but not widespread. Grow into lycopods. Biofilms full of bacteria were spreading across the plains, where lichens and fungi were starting to grow but not widespread

There were no lumps of ‘earth’. There are still no soils to be found. This means that for the first four billion years of this planet’s life there was no soil. Not a ped anywhere. There were not even the characters or conditions needed to make soil, hat give this planet’s name – Earth.  However, there were early signs of some of the soil's processes and key components. which continued to emerge over the next hundred millions of years..Half a billion years ago there was nothing that looked like soil does today. But over theset 100 my we can find mosses, fungi, and bacteria that started creating debris. Key characters were arriving to make use of that.  

Key Function in Soil evolution

We have seen the presence of various microbes, and the arrival of several plants and animals, some of which are going to make an impact on soil formation. Several key soil functions appeared and grew during the 100 million years from 500- 400mya. Perhaps the most significant was their ability to live in moist conditions but capable of withstanding dry periods. 

Wet

We tend to describe the creation of life on land as 'conquering the dry conditions'. This is true. But It may well be better to consider the 'extension of wet conditions' too. This would be via thin water films - biofilms. There would be wierd water tensions, and those that can cope with that can go further inland. It is these biofilms that holds the rest together. Bacteria would be in them as well as tiny nematode worms. The biofilm thus becomes an environment of its own, somewhere other organisms could live. The more water the better it does that. After the average composition of living organisms is 65-90% water!

Healthy soil today is about 1/4 air and 1/4 water…but we have a long way to get there.

The extent and survival of mosses  by around  400mya play a crucial part in soil evolution. They die leaving behind organic matter, which provides food for microbes and create the conditions for a host of small animals to live in stable conditions. Some of the inhabitants (like tardigrades and rotifers) may have existed much the same in the same environment since that time. Others, like nematodes and small creatures provide a mixture of undigested organic matter and metabolic waste products., that are recycled as nutrients. Small  soil animal poo plays a vital role in soil formation. But there was no real soil structure. 

Many creatures ran around then. The springtails could have sprung along the biofilms to find their food - bacteria, by following the raindrops and the smell of the earth, produced by those bacteria, that we know today. They developed mechanisms to jump along the films, but also be able to jump back to safety. This was a massive evolutionary advantage and a great leap for life. 

 Start the soil story at 500mya

This site is set up by Dr Charlie Clutterbuck
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