A flower is reproductive structure in angiosperms that has up to four (4) whorls of modified leaves on a determinate stem (receptacle / pedicel). Flowers can have some or all of the following parts:
Sepals (Calyx)
Petals (Corolla)
Stamens (Androecium)
Carpels / Pistil (Gynoecium)
Above: flower bud of Lily in transverse section
Modified leaves used for protecting the delicate structures on the flower while in bud stage
Some sepals appear quite petal-like, and if it is difficult to distinguish, the term tepals is used
Sepals are sometimes leaf-like structures
Collectively sepals are known as calyx
Above: the green leaf-like sepals of a Geranium flower
Modified leaves that are showy structures to attract pollinators
Absent in wind- and water-pollinated flowers (see catkins)
Collectively known as corolla
Perianth: The term "perianth" refers to both the corolla and calyx
Tepal: If the sepals and petals look identical, the term "tepal" is used.
Nectaries: organ with sugar-rich liquid called nectar, usually found at the base of the perianth for animal pollinators
Above: the petals of Serviceberry (Amelanchier)
Also known as the androecium
Pollen-producing or male portion of the flower
Most stamens have two parts: anther and filament
Pollen-producing structures
Pollen is the dispersed microspores of a seed plant that contain the male gametophyte of the plant
Site of meiosis to create pollen (or microspores)
Most angiosperm anthers are four (4) fused microsporangia
Tapetum: a layer of cells inside of microsporangia that provides nutrition to developing pollen
Stalk which elevates anther to dust pollinators
Over evolutionary time, anther-bearing leaves reduced into the filament of derived angiosperms
Some ancient angiosperms (e.g. Austrobaileya, Degeneria) still have leaf-like or petal-like stamens
Above: the anthers of an Amaryllis flower
Above: petalloid stamens of a daylily cultivar
Above: Flower of Austrobaileya with petalloid stamens
The female portion of the flower; also called the gynoecium
The pistil usually has three parts: stigma, style, and ovary (see below)
The term "carpel" refers to a single-seeded structure; it is the ancestral state.
Some ancestral angiosperms exhibit carpels, not pistils (e.g. Magnolia - see image below)
A pistil is the evolutionary result of the fusion of several carpels
Developmentally, pistils/carpels are modified leaves that completely envelope the seeds
Over evolutionary time, ovule-bearing leaves folded lengthwise to form the carpel
This is thought to be an adaptation to protect the ovules
Some ancient angiosperms (e.g. Akebia, Degeneria) have carpels that do not completely close during development, called conduplicate carpels (see image below)
A landing pad for pollen; the site of pollination
A stalk that elevates the stigma to the height of the anthers
The ovary protects ovules and develops into the fruit
Ovules are unfertilized seeds
Ovules are called "seeds" after fertilization
Above: the ripening pistil of a flower: an immature fruit
Above: multiple carpels of Magnolia
Above: Flower of Akebia with conduplicate carpels
The corpse flower (Rafflesia arnoldii) is the largest known flower
The flower grows up to one meter (3 ft) in diameter and weighs up to 11 kilograms (24 lbs)
Rafflesia grows in old-growth rain forests of Borneo and Sumatra
This plant is a parasitic plant with haustoria that tap into the stems of tropical grapevines (Tetrastigma rafflesiae)
Watch a time-lapse video of Rafflesia opening
N.B. Some people will report that the largest flower is Amorphophallus titanum, but this is not a single flower. Instead, it has a cluster of flowers, called an inflorescence, on a long stalk called a spadix.
Above: The corpse flower (Rafflesia arnoldii); the largest flower in the world
Droughts can reduce the caloric value of flower nectar by up to 95%, study finds (Phys.org 24Jun2025)
└Frigero et al. (2025) Extreme events induced by climate change alter nectar offer to pollinators in cross pollination-dependent crops
Plants Sweeten Nectar When They Hear (Yes, Hear) Bees Buzzing Nearby (Science Blog 21May2025)
Ancient pollen reveals stories about Earth's history, from dinosaurs to the Mayan collapse (Phys.org 20May2025)
New species of Rafflesia discovered (Galindon et al., 2016)
Parasitic flower pirates genes from its host (Scientific American, 2012)
Aquatic eel-grass is pollinated by zooplankton (Hakai Magazine, 2016)