Inflorescences

What is an inflorescence?

  • An inflorescence is a cluster of flowers arranged along a stem.

  • The branching of the flowering stalk creates most of the inflorescence types

  • Inflorescences contain pedicels (stem that directly holds a flower) and peduncles (stems that support other stems and pedicels)

  • There are several types of inflorescences:

Spike

  • Unbranched inflorescence with sessile flowers (no pedicels)

https://upload.wikimedia.org/wikipedia/commons/0/04/Woolly_mullein_flower-spike.jpg

Above: The spike of flowers in mullein (Verbascum thapsus)

Raceme

  • Unbranched inflorescence with flowers on pedicels

https://upload.wikimedia.org/wikipedia/commons/9/9d/Muscari.jpg

Above: The raceme of Grape hyacinth (Muscari)

Panicle

  • A branched or compound raceme

  • Main rachis with branches bearing flowers on pedicels

https://upload.wikimedia.org/wikipedia/commons/d/d4/Buddleja_davidii_%27Fascination%27_panicle.jpg

Above: A panicle of Butterfly Bush (Buddleja)

Catkin

  • A spike-like inflorescence of unisexual flowers

  • Catkins are usually wind-pollinated and lack petals

  • Typical inflorescence of willow (Salix), cottonwood (Populus), oak (Quercus), alder (Alnus), and birch (Betula)

  • This feature has evolved several times, in different groups of angiosperms

https://upload.wikimedia.org/wikipedia/commons/8/84/Black_cottonwood_male_catkin_and_leaf_buds.JPG

Above: A male catkin of Cottonwood (Populus)

Cyme

  • A group of flowers in which the oldest flower terminates the main axis, and the inflorescence continues branching through axillary buds

  • Scorpioid cymes have 1-sides branching, forming a coiled inflorescence

  • e.g. Boraginaceae (forget-me-not family) and Hydrophyllaceae (waterleaf family)

https://upload.wikimedia.org/wikipedia/commons/8/86/Scorpioid_cyme_%286940408930%29.jpg

Above: Scorpioid cyme of the Boraginaceae

Umbel

  • All flower stalks (pedicels) originate from the same point with equal lengths, creating a flat-topped appearance

  • Simple umbel: inflorescence contains a peduncle and several equal-length pedicels

  • Compound umbel: inflorescence contains a peduncle, which gives rise to several "rays", which give rise to equal-length pedicels

  • e.g. Apiaceae (carrot family), Asclepidaceae (milkweed family)

Above: simple umbel of milkweed (Asclepias)

Above: Compound umbel of Queen Anne's Lace (Daucus carota)

Corymb

  • Similar in branching to a panicle, but similar in shape to an umbel

  • The proximal (lower) flowers have longer pedicels

  • The pedicels become shorter with more distal flowers

  • The result is an inflorescence with a flat top appearance

  • e.g. Yarrow (Achillea), many species in the Maloideae, such as hawthorns and Sorbus

https://upload.wikimedia.org/wikipedia/commons/7/77/Achillea_millefolium_bloem.jpg

Above: The corymb of Yarrow (Achillea millefolium)

Head or Composite

  • Inflorescence in which many flowers are attached to a dome-like receptacle, and appears to be 1 flower

  • e.g. Daisy family (Asteraceae or Compositae)

Ray flowers

  • Bilateral flowers, with fused petals, that make the entire ray flower look like 1 petal

Disk flowers

  • Radial flowers, fused at the base, that form the "eye" of a composite inflorescence

Composite species may have a combination of disk flowers and/or ray flowers

  • Sunflower (Helianthus) is an example with both types of flowers

  • Dandelion (Taraxacum) is an example with just ray flowers

  • Pineapple weed (Matricaria discoidea) is an example with disk flowers only

https://upload.wikimedia.org/wikipedia/commons/5/57/Ursinia_speciosa_%28Compositae%29_flower.JPG

Above: Head of a composite inflorescence (Ursinia) with both disk and ray flowers

Above: Dandelion (Taraxacum officinale) is a composite with ray flowers only

Above: Pineapple weed (Matricaria discoidea) is a composite with disk flowers only

Spadix and Spathe

  • A thick, fleshy spike of unisexual flowers lacking petals (spadix), often surrounded by a vase-shaped modified leaf (spathe)

  • e.g. Arum family (Araceae)

https://upload.wikimedia.org/wikipedia/commons/f/f6/Arum_maculatum.jpeg

Above: The spadix and spathe of wild arum (Arum maculatum)

What is the largest inflorescence?

  • The talipot palm (Corypha umbraculifera) has the largest inflorescence in the world

    • The inflorescences of many palms (Arecaceae) are panicles

  • The inflorescence grows up to 25 m (80 ft) with stems up to 1.3 m (4.25 ft) in diameter

  • This palm grows in eastern and southern India and Sri Lanka

https://upload.wikimedia.org/wikipedia/commons/c/cc/Flowering_Talipot_Palm_06.jpg

Above: The inflorescence of the talipot palm (Corypha umbraculifera); the largest in the world

What about the Titan Arum?

  • The largest unbranched inflorescence is the titan arum, Amorphophallus titanum

    • The titan arum is in the Araceae, which means that it has a spathe and spadix inflorescence

  • The inflorescence grows over 3m (10 ft )

  • It is native to western Sumatra but is commonly grown and displayed at botanical gardens

Above: the spathe and spadix inflorescence of the Titan Arum (Amorphophallus titanum)