NOTE: For many of these references, I have pdfs. This list includes all known papers on the fossil record of thylacocephalans. A number of fossils originally interpreted as phyllocarids have subsequently been reinterpreted as thylacocephalans. If you think I am missing something, please let me know!
Thylacocephalan references (117)
1. Affer, D., and G. Teruzzi. 1999. Thylacocephalan crustaceans from the Besano Formation, Middle Triassic, N. Italy. Rivista del Museo Civico di Scienze Naturali “Enrico Caffi” 20: 5-8.
2. Alessandrello, A., P. Arduini, G. Pinna, and G. Teruzzi. 1991. New observations on the Thylacocephala (Arthropoda, Crustacea). In The Early Evolution of Metazoa and the Significance of Problematic Taxa, A. M. Simonetta, and S. Conway Morris, eds. Cambridge University Press, Cambridge, p. 245-251.
3. Arduini, P., G. Pinna, and G. Teruzzi. 1982. Il giacimento sinemuriano di Osteno in Lombardia e i suoi fossili [The Sinemurian deposits of Osteno in Lombardy and their fossils]. In Palaeontology, Essential of Historical Geology [Paleontologia come Scienza Geostorica], E. Montanaro Gallitelli, ed. STEM Mucchi Modena Press, Modena, p. 495-522.
4. Arduini, P., and A. Brasca. 1984. Atropicaris: Nuovo genere della classe Thylacocephala. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 125: 87-93.
5. Arduini, P., G. Pinna, and G. Teruzzi. 1984. Ostenocaris nom. nov. pro Ostenia Arduini, Pinna & Teruzzi, 1980. Atti Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 125: 48.
6. Arduini, P. 1988. Microcaris and Atropicaris, two genera of the class Thylacocephala. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 129: 159-163.
7. Arduini, P., and G. Pinna. 1989. I Tilacocefali: una nuova classe di crostacei fossili. Natura 80: 1-34.
8. Arduini, P. 1990. Studies on Permo-Trias of Madagascar-n. 1. Thylacocephala from Lower Trias of Madagascar. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 131: 197-204.
9. Arduini, P. 1991. Clausocaris pinnai n. sp., (Order Clausocarida nov.), thylacocephalan crustacean from the Norian of the Preone Valley (Udine, N. Italy) and morphological considerations on Thylacocephala. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 132: 265-272.
10. Boucot, A. J. 1990. Evolutionary Paleobiology of Behavior and Coevolution. 1-725.
11. Braig, F., J. T. Haug, M. Schädel, and C. Haug. 2019. A new thylacocephalan crustacean from the Upper Jurassic lithographic limestones of southern Germany and the diversity of Thylacocephala. Palaeodiversity 12: 69.
12. Brayard, A., L. J. Krumenacker, J. P. Botting, J. F. Jenks, K. G. Bylund, E. Fara, E. Vennin, N. Olivier, N. Goudemand, T. Saucède, S. Charbonnier, C. Romano, L. Doguzhaeva, B. Thuy, M. Hautmann, D. A. Stephen, C. Thomazo, and G. Escarguel. 2017. Unexpected Early Triassic marine ecosystem and the rise of the Modern evolutionary fauna. Science Advances 3: e1602159.
13. Briggs, D. E. G., and W. D. I. Rolfe. 1983. New Concavicarida (new order: ?Crustacea) from the Upper Devonian of Gogo, Western Australia, and the palaeoecology and affinities of the group. Special Papers in Palaeontology 30: 249-276.
14. Briggs, D. E. G., H. P. Liu, R. M. McKay, and B. J. Witzke. 2015. Bivalved arthropods from the Middle Ordovician Winneshiek Lagerstätte, Iowa, USA. Journal of Paleontology 89: 991-1006.
15. Broda, K., T. A. Hegna, and M. Zatoń. 2015. Fossils Explained 65: Thylacocephala. Geology Today 31: 116-120.
16. Broda, K., M. Wolny, and M. Zatoń. 2015. Palaeobiological significance of damaged and fragmented thylacocephalan carapaces from the Upper Devonian of Poland. Proceedings of the Geologists’ Association 126: 589-598.
17. Broda, K., and M. Zatoń. 2017. A set of possible sensory system preserved in cuticle of Late Devonian thylacocephalan arthropods from Poland. Historical Biology 29: 1045-1055.
18. Broda, K., L. Marynowski, M. Rakociński, and M. Zatoń. 2019. Coincidence of photic zone euxinia and impoverishment of arthropods in the aftermath of the Frasnian-Famennian biotic crisis. Scientific Reports 9: 16996.
19. Broda, K., Š. Rak, and T. A. Hegna. 2020. Do the clothes make the thylacocephalan? A detailed study of Concavicarididae and Protozoeidae (?Crustacea, Thylacocephala) carapace micro-ornamentation. Journal of Systematic Palaeontology 18: 911-930.
20. Bürgin, T., U. Eichenberger, H. Furrer, and K. Tschanz. 1991. Die Prosanto-Formation―eine fischreiche Fossil-Lagerstätte in der Mitteltrias der Silvretta-Decke (Konton Graubünden, Schweiz) [The Prosanto Formation―a fish-rich fossil deposit in the Middle Triassic of the Silvretta Nappe (Konton Graubünden, Switzerland)]. Eclogae Geologicae Helvetiae 84: 921-990.
21. Calzada, S., and R. Mañé. 1993. Primera cita de un Tilacocéfalo (Crustacea) (Ferrecaris n. gen. magransi n. sp.) en el Ladiniense español. Trabajos del Museo Geológico del Seminario C. de Barcelona 224: 12-16.
22. Charbonnier, S. 2009. Le Lagerstätte de la Voulte: un Environment Bathyal au Jurassique. Publications Scientifiques du Muséum, Paris.
23. Charbonnier, S., J. Vannier, P. Hantzpergue, and C. Gaillard. 2010. Ecological significance of the arthropod fauna from the Jurassic (Callovian) La Voulte Lagerstätte. Acta Palaeontologica Polonica 55: 111-132.
24. Charbonnier, S., G. Teruzzi, D. Audo, M. Lasseron, C. Haug, and J. T. Haug. 2017. New thylacocephalans from the Cretaceous Lagerstätten of Lebanon. Bulletin de la Société Géologique de France 188: 19.
25. Charbonnier, S., A. Brayard, K. G. Bylund, G. Escarguel, E. Fara, N. Goudemand, P. Gueriau, J. F. Jenks, L. J. Krumenacker, N. Olivier, D. A. Stephen, C. Thomazo, M. Thoury, and E. Vennin. 2019. New thylacocephalans from the Early Triassic Paris Biota (Bear Lake County, Idaho, USA). Geobios 54: 37-43.
26. Cooper, C. L. 1932. A crustacean fauna from the Woodford Formation of Oklahoma. Journal of Paleontology 6: 346-352.
27. Dalla Vecchia, F. M., and G. Muscio. 1990. Occurrence of Thylacocephala (Arthropoda, Crustacea) from the Upper Triassic of Carnic Prealps (N.E. Italy). Bolletino della Societa Paleontologica Italiana 29: 39-42.
28. Dalla Vecchia, F. M. 1992. Segnalazione di crostacei nell’unità Fonte Santa (Triassico sup.) presso Filettino (Lazio, Italia) [Record of crustaceans from Fonte Santa Unit (Upper Triassic) near Filettino (Lazio, Italy)]. Gortania 14: 59-69.
29. Dames, W. 1886. Ueber einige crustaceen aus den kreideablagerungen des libanon. Zeitschrift der Deutsche Geologische Gesellschaft 38: 551-575.
30. Ehiro, M., O. Sasaki, H. Kano, J. Nemoto, and H. Kato. 2015. Thylacocephala (Arthropoda) from the Lower Triassic of the South Kitakami Belt, Northeast Japan. Paleontological Research 19: 269-282.
31. Ehiro, M., and H. Kano. 2024. A new species of Parisicaris (Microcarididae, Thylacocephala) from the upper Olenekian (Lower Triassic) Osawa Formation in the South Kitakami Belt, Northeast Japan. Bulletin of the Tohoku University Museum 23: 23-34.
32. Feldmann, R. M., C. E. Schweitzer, S. Hu, J. Huang, C. Zhou, Q. Zhang, W. Wen, T. Xie, and E. Maguire. 2015. Spatial distribution of Crustacea and associated organisms in the Luoping Biota (Anisian, Middle Triassic), Yunnan Province, China: evidence of periodic mass kills. Journal of Paleontology 89: 1022-1037.
33. Fischer, J.-C. 2003. Invertébrés remarquables du Callovien inférieur de la Voulte-sur-Rhône (Ardèche, France) [Remarkable invertebrates from the Lower Callovian of la Voulte-sur-Rhone (Ardeche, France)]. Annales de Paléontologie 89: 223-252.
34. Forchielli, A., and P. Pervesler. 2013. Phosphatic cuticle in thylacocephalans: a taphonomic case study of Austriocaris (Arthropoda, Thylacocephala) from the fossil-lagerstätte Polzberg (Reingraben Shales, Carnian, Upper Triassic, Lower Austria). Austrian Journal of Earth Science 106: 46-61.
35. Frey, L., A. Pohle, M. Rücklin, and C. Klug. 2020. Fossil‐Lagerstätten, palaeoecology and preservation of invertebrates and vertebrates from the Devonian in the eastern Anti‐Atlas, Morocco. Lethaia 53: 242-266.
36. Fröhlich, F., A. Mayrat, B. Riou, and S. Secretan. 1992. Structures rétiniennes phosphatisées dans l’œil géant de Dollocaris un crustacé fossile. Annales de Paléontologie 78: 193-204.
37. Garwood, R. 2007. Tomographic reconstructions of the enigmatic thylacocephalan arthropod Dollocaris ingens. Department of Earth Science & Engineering MSci: 36.
38. Gass, K. C., and S. J. Braddy. 2023. The Waukesha Biota: a wonderful window into early Silurian life. Geology Today 39: 169-176.
39. Haig, D. W., S. K. Martin, A. J. Mory, S. McLoughlin, J. Backhouse, R. W. Berrell, B. P. Kear, R. Hall, C. B. Foster, G. R. Shi, and J. C. Bevan. 2015. Early Triassic (early Olenekian) life in the interior of East Gondwana: mixed marine–terrestrial biota from the Kockatea Shale, Western Australia. Palaeogeography, Palaeoclimatology, Palaeoecology 417: 511-533.
40. Haug, C., and J. T. Haug. 2014. Neues zu den Thylacocephala, einer gruppe (immer noch) seltsamer krebse [News about the Thylacocephala, a group of (still) strange crabs]. Archaeopteryx 32: 8-14.
41. Haug, C., D. E. G. Briggs, D. G. Mikulic, J. Kluessendorf, and J. T. Haug. 2014. The implications of a Silurian and other thylacocephalan crustaceans for the functional morphology and systematic affinities of the group. BMC Evolutionary Biology 14: 1-15.
42. Haug, J. T., C. Haug, and M. Ehrlich. 2008. Erster fossiler nachweis einer fangschreckenkrebslarve (Arthropoda: Crustacea) und eine neue methode zur dokumentation von Solnhofen-Fossilien (Oberjura, Süddeutchland) [First fossil evidence of a stomatopod larva (Arthropoda: Crustacea) and a new method for documentation of Solnhofen fossils (Upper Jurassic, South Germany)]. Archaeopteryx 26: 19-26.
43. Haug, J. T., C. Haug, and M. Ehrlich. 2008. First fossil stomatopod larva (Arthropoda: Crustacea) and a new way of documenting Solnhofen fossils (Upper Jurassic, Southern German). Palaeodiversity 1: 103-109.
44. Hegna, T. A., F. J. Vega, and K. A. González-Rodríguez. 2014. First Mesozoic thylacocephalans (Arthropoda, ?Crustacea; Cretaceous) in the western hemisphere: new discoveries from the Muhi Quarry Lagerstätte. Journal of Paleontology 88: 606-616.
45. Herter, S., G. Eberle, M. Freif, and C. Klug. 2025. Devonische knorpelfische aus dem marokkanischen Anti-Atlas [Devonian cartilaginous fishes from the Moroccan Anti-Atlas]. Fossilien 2025: 4-13.
46. Hu, S.-x., Q.-y. Zhang, W. Wen, J.-y. Huang, C.-y. Zhou, T. Xie, T. Lü, T.-l. Liu, and M. J. Benton. 2014. The Luoping Biota: A Taphonomic Window on Triassic Biotic Recovery and Radiation. Yunnan Publishing Group Co. & Yunnan Science and Technology Press, Kunming, CHina.
47. Jauvion, C., S. Bernard, P. Gueriau, C. Mocuta, S. Pont, K. Benzerara, and S. Charbonnier. 2020. Exceptional preservation requires fast biodegradation: thylacocephalan specimens from La Voulte‐sur‐Rhône (Callovian, Jurassic, France). Palaeontology 63: 395-413.
48. Ji, C., A. Tintori, D. Jiang, and R. Motani. 2017. New species of Thylacocephala (Arthropoda) from the Spathian (Lower Triassic) of Chaohu, Anhui Province of China. Paläontologische Zeitschrift 91: 171-184.
49. Ji, C., A. Tintori, D. Jiang, R. Motani, and F. Confortini. 2021. New Thylacocephala (Crustacea) assemblage from the Spathian (Lower Triassic) of Majiashan (Chaohu, Anhui Province, South China). Journal of Paleontology 95: 305-319.
50. Ji, C., and A. Tintori. 2024. New material of Thylacocephala from the Early Ladinian (Middle Triassic) of Northern Grigna (Lecco, Lombardy, Northern Italy). Diversity 16: 677.
51. Jobbins, M., C. Haug, and C. Klug. 2020. First African thylacocephalans from the Famennian of Morocco and their role in Late Devonian food webs. Scientific Reports 10: 5129.
52. Kobayashi, T. 1937. An occurrence of a new Permian phyllocarid in South Chosen. Journal of the Geological Society of Japan 44: 337-340.
53. Križnar, M., and T. Hitij. 2010. Nevretenčarji (Invertebrates) Strelovške formacije. Scopolia Suppl. 5: 91-107.
54. Kühl, G., and J. Rust. 2012. Captopodus poschmanni gen. et sp. nov. a new stem-group arthropod from the Lower Devonian Hünsruck Slate (Germany). Arthropod Structure & Development 41: 609-622.
55. Lange, S., C. H. J. Hof, F. R. Schram, and F. A. Steeman. 2001. New genus and species from the Cretaceous of Lebanon links the Thylacocephala to the Crustacea. Palaeontology 44: 905-912.
56. Lange, S., and F. R. Schram. 2002. Possible lattice organs in Cretaceous Thylacocephala. Contributions to Zoology 71: 159-169.
57. Laville, T., C. Haug, J. T. Haug, M.-B. Forel, and S. Charbonnier. 2021. Morphology and anatomy of the Late Jurassic Mayrocaris bucculata (Eucrustacea?, Thylacocephala) with comments on the tagmosis of Thylacocephala. Journal of Systematic Palaeontology 19: 289-320.
58. Laville, T., J. T. Haug, and C. Haug. 2021. New species of Thylacocephala, Eodollocaris keithflinti n. gen., n. sp., from the Mazon Creek Laterstätte, Illinois, United States (c. 307 Ma) and redescription of other Mazon Creek thylacocephalans. Geodiversitas 43: 295-310.
59. Laville, T., C. P. A. Smith, M.-B. Forel, A. Brayard, and S. Charbonnier. 2021. Review of early Triassic Thylacocephala. Rivista Italiana di Paleontologia e Stratigrafia 127: 73-101.
60. Laville, T., M.-B. Forel, and S. Charbonnier. 2022. Resolving the tagmatisation and the phylogenetic affinities of thylacocephala: Input from 3D data. 8th Symposium on Fossil Decapod Crustaceans. Palaeontological Publications 1: 68-69.
61. Laville, T., T. Hitij, R. Gašparič, J. Žalohar, M.-B. Forel, and S. Charbonnier. 2022. The Triassic thylacocephalans from Slovenia: Implications for their evolution and diversification. 8th Symposium on Fossil Decapod Crustaceans. Palaeontological Publications 1: 70-71.
62. Laville, T., T. A. Hegna, M.-B. Forel, S. Darroch, and S. Charbonnier. 2023. New look at Concavicaris woodfordi (Euarthropoda: Pancrustacea?) using micro-computed tomography. Palaeontologia Electronica 26: a1.
63. Laville, T., M.-B. Forel, and S. Charbonnier. 2023. Re-appraisal of thylacocephalans (Euarthropoda, Thylacocephala) from the Jurassic La Voulte-sur-Rhône Lagerstätte. European Journal of Taxonomy 898: 1-61.
64. Laville, T., T. Hitij, M.-B. Forel, R. Gašparič, J. Žalohar, and S. Charbonnier. 2024. Middle and Late Triassic thylacocephalans from Slovenia: Implications for their diversification. Bulletin of Geosciences 99: 219-253.
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66. Lukeneder, A., and P. Lukeneder. 2021. The Upper Triassic Polzberg palaeobiota from a marine Konservat-Lagerstätte deposited during the Carnian Pluvial Episode in Austria. Scientific Reports 11: 16644.
67. Lukeneder, A., and P. Lukeneder. 2023. New data on the marine Upper Triassic palaeobiota from the Polzberg Konservat-Lagerstätte in Austria. Swiss Journal of Palaeontology 142: 1-18.
68. Mergl, M. 2020. The cuticles of (?) thylacocephalan arthropod from the basal Choteč Event (Choteč Formation, Eifelian; Barrandian Area, Czech Republic). Folia Musei rerum naturalium Bohemiae occidentalis. Geologica et Paleobiologica 54: 1-12.
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