Up-to-date citations can be found here.



Brandl, H.B.*, Farine, D.R.*, Funghi, C., Schuett, W., Griffith, S.C. (2019) Early-life social environment predicts social network position in wild zebra finchesProceedings of the Royal Society B 286: 20182579. (link)   *authors contributed equally

Hillemann, F., Cole, E.F., Keen, S.C., Sheldon, B.C., Farine, D.R. (accepted) Diurnal variation in the production of vocal information about food supports a model of social adjustment in wild songbirds. Proceedings of the Royal Society B 286: 20182740. (link)

Carter, G.G., Schino, G., Farine, D.R. (in press) Challenges in assessing the roles of nepotism and reciprocity in cooperation networksAnimal Behaviour (link)

Lang, S.D.J., Mann, R.P., Farine, D.R. (in press) Temporal activity patterns of predators and prey across broad geographic scalesBehavioural Ecology (link)

He, P., Maldonado-Chaparro, A.A., Farine, D.R. (2019) The role of habitat configuration in shaping social structure: a gap in studies of animal social complexityBehavioural Ecology & Sociobiology 73: 9. (link

Wilkinson, G.S., Carter, G.G., Bohn, K.M., Baspers, B., Chaverri, G., Farine, D.R., Günther, L., Kerth, G., Knörnschild, M., Mayer, F., Nagy, M., Ortega, J., Patriquin, K. (2019) Kinship, association and social complexity in batsBehavioural Ecology & Sociobiology 73: 7. (link)


Somveille, M., Firth, J.A., Aplin, L.M., Farine, D.R., Sheldon, B.C., Thompson, R.N. (2018) Movement and conformity interact to establish local behavioural traditions in animal populations. PLoS Computational Biology 14(12): e1006647. (link)

Maldonado-Chaparro, A.A., Alarcón-Nieto, G., Klarevas-Irby, J.A., Farine, D.R. (2018) Experimental disturbances reveal group-level costs of social instability. Proceedings of the Royal Society B 285: 20181577. (link)

Smaldino, P.E., Aplin, L.M., Farine, D.R. (2018) Sigmoidal Acquisition Curves Are Good Indicators of Conformist Transmission. Scientific Reports 8: 14015. (link)

Boogert, N.J., Lachlan, R.F., Spencer, K.A., Templeton, C.N., Farine, D.R. (2018) Stress hormones, social associations and song learning in zebra finches. Philosophical Transactions of the Royal Society B 373: 20170290. (link)

Marthy, W., Farine, D.R. (2018) The potential impacts of the songbird trade on mixed-species flocking. Biological Conservation 222: 222-231. (link)

Cantor, M., Farine, D.R. (2018) Simple foraging rules in competitive environments can generate socially-structured populations. Ecology and Evolution 8(10): 4978-4991. (link)

Davis, G.H., Crofoot, M.C., Farine, D.R. (2018) Estimating the robustness and uncertainty of animal social networks using different observational methods. Animal Behaviour 141: 29-44. (link)

Alarcón-Nieto, G., Graving, J.M., Klarevas-Irby, J.A., Maldonado-Chaparro, A.A., Mueller, I., Farine, D.R. (2018) An automated barcode tracking system for behavioural studies in birds. Methods in Ecology and Evolution  9(6): 1536-1547. (link)

Moyers, S.C., Adelman, J.S., Farine, D.R., Moore, I.T., Hawley, D.M. (2018) Exploratory behavior is linked to stress physiology and social network centrality in free-living house finches (Haemorhous mexicanus). Hormones and Behaviour 102: 105-113. (link)


Maldonado-Chaparro, A.A., Montiglio, P.-O., Forstmeier, W., Kempenaers, B., Farine, D.R. (2018) Linking the fine-scale social environment to mating decisions: a future direction for the study of extra-pair paternity. Biological Reviews 93(3): 1558-1577. (link)

Moyers, S.C., Adelman, J.S., Farine, D.R., Thomason, C.A., Hawley, D.M. (2018) Feeder density enhances house finch disease transmission in experimental epidemicsPhilosphical Transactions of the Royal Society B 373: 20170090. (link)

Montiglio, P.O., McGlothlin, J.W., Farine, D.R. (2018) Social structure modulates the evolutionary consequences of social plasticity: taking a social network perspective of interacting phenotypesEcology and Evolution  8(3): 1451–1464. (link

Sánchez-Tójar, A. Schroeder, J., Farine, D.R. (2018) A practical guide for inferring reliable dominance hierarchies and estimating their uncertaintyJournal of Animal Ecology 87(3): 594-608. (link)

Strandburg-Peshkin, A., Papageorgiou, D., Crofoot, M.C., Farine, D.R. (2018) Inferring influence and leadership in moving animal groupsPhilosophical Transactions of the Royal Society B 373: 20170006. (link)

Hoppitt, W., Farine, D.R. (2018) Association indices for quantifying social relationships: how to deal with missing observations of individuals or groupsAnimal Behaviour 136: 227-238. (link)

Farine, D.R. (2018) When to choose dynamic vs. static social network analysisJournal of Animal Ecology 87(1): 128-138. (link)



McDonald, G.C., Farine, D.R., Foster, K.R., Biernaskie, J.M. (2017) Assortment and the analysis of natural selection on social traits. Evolution 71(11): 2693-2702. (link)

Lang, S.D.J., Farine, D.R. (2017) A multidimensional framework for studying social predation strategies. Nature Ecology & Evolution 1: 1230-1239. (link)

Radersma, R., Garroway, C.J., Santure, A., de Cauwer, I., Farine, D.R., Slate, J., Sheldon, B.C. (2017) Social and spatial effects on genetic variation between foraging flocks in a wild bird population. Molecular Ecology 26(20): 5807–5819. (link)

Carter, G.G., Farine, D.R., Wilkinson, G.S. (2017) Social bet-hedging in vampire batsBiology Letters 13: 20170112. (link)

Isbell, L.A., Bidner, L.R., Crofoot, M.C., Matsumoto-Oda, A., Farine, D.R. (2017) GPS-identified, low-level nocturnal activity of vervets (Chlorocebus pygerythrus) and olive baboons (Papio anubis) in Laikipia, KenyaAmerican Journal of Physical Anthropology 164(1): 203-211. (link)
 [44] Farine, D.R.*, Strandburg-Peshkin, A.*, Couzin, I.D., Berger-Wolf, T.Y., Crofoot, M.C. (2017) Individual variation in local interaction rules can explain emergent patterns of spatial organisation in wild baboonsProceedings of the Royal Society B 284: 20162243. (link)


Farine, D.R. (2017) A guide to null models for animal social network analysis. Methods in Ecology and Evolution 8(10): 1309-1320. (link)

Johnson, K.V.-A, Aplin, L.M., Cole, E.F., Farine, D.R., Firth, J.A., Patrick, S.C., Sheldon, B.C. (2017) Male great tits assort by personality during the breeding seasonAnimal Behaviour 128: 21-32. (link)

Farine, D.R. (2017) The dynamics of transmission and the dynamics of networks. Journal of Animal Ecology 86(3): 415-418. (link)

Strandburg-Peshkin, A., Farine, D.R., Crofoot, M. C. & Couzin, I.D. (2017) Habitat and social factors shape individual decisions and emergent group structure during baboon collective movementeLife 6:e19505. (link)
 [39] Francesiaz, C., Farine, D.R., Laforge, C., Béchet, A., Sadoul, N., Besnard, A. (2017) Familiarity drives social philopatry in an obligate colonial breeder with weak interannual breeding-site fidelityAnimal Behaviour 124: 83-94. (link)

 [38] Miller, E.T., Farine, D.R., Trisos, C.H. (2017) Phylogenetic community structure metrics and null models: a review with new methods and softwareEcography 40(4): 461–477. (link)


Farine, D.R., Strandburg-Peshkin, A., Berger-Wolf, T., Ziebart, B., Brugere, I., Li, J., Crofoot, M.C. (2016) Both Nearest Neighbours and Long-term Affiliates Predict Individual Locations During Collective Movement in Wild Baboons. Scientific Reports 6: 27704. (link)

Firth, J.A., Sheldon, B.C., Farine, D.R. (2016) Pathways of information transmission amongst wild songbirds follow experimentally imposed changes in social foraging structure. Biology Letters 12: 20160144. (link)

Crates, R.A., Firth, J.A., Farine, D.R., Garroway, C.J., Kidd, L.R., Aplin, L.M., Radersma, R., Milligan, N.D., Voelkl, B., Culina, A., Verhelst, B.L., Hinde, C.A., Sheldon, B.C. (2016) Individual variation in winter supplementary food consumption and its consequences for reproduction in wild birds. Journal of Avian Biology 47(5): 678-689. (link)

Shizuka, D., Farine, D.R. (2016) Measuring the robustness of network community structure using assortativityAnimal Behaviour 112: 237-246. (link)  

Leu, S.T., Farine, D.R., Wey, T.W., Sih, A., Bull, M. (2016) Environment modulates population social structure: experimental evidence from replicated social networks of wild lizardsAnimal Behaviour 111:23-31. (link)


Aplin, L.M., Farine, D.R., Morand-Ferron, J., Cockburn, A., Thornton, A., Sheldon, B.C. (2015) Counting conformity: evaluating the units of measurement in frequency-dependent social learning. Animal Behaviour 110: e5-e8. (link)

Firth, J.A., Voelkl, B., Farine, D.R., Sheldon, B.C. (2015) Experimental evidence that social relationships determine individual foraging behaviour. Current Biology 25: 1-6. (link)
News highlights include: Science, IFLScience, Discovery, BBC.

Farine, D.R., Strandburg-Peshkin, A. (2015) Estimating uncertainty and reliability of social network data using Bayesian inference. Royal Society Open Science 2: 150367. (link) * all authors contributed equally
This paper was written as a 24 hour science challenge live online, representing the first truly open scientific publication (read more about it here).

Farine, D.R., Montiglio, P.O., Spiegel, O. (2015) From individuals to groups and back: the evolutionary implications of group phenotypic composition. Trends in Ecology and Evolution 30(10): 609-621. (linkall authors contributed equally

Strandburg-Peshkin, A.*, Farine, D.R.*, Couzin, I.D., Crofoot, M.C. (2015) The wisdom of baboon decisionsScience 349(6251): 935-936. (link)

Adelman, J.S., Moyers, S.C., Farine D.R., Hawley, D.M. (2015) Feeder use predicts both acquisition and transmission of a contagious pathogen in a North American songbird. Proceedings of the Royal Society B 282: 20151429. (link)

Farine, D.R.*, Spencer, K.A., Boogert, N.J.* (2015) Early-Life Stress Triggers Juvenile Zebra Finches to Switch Social Learning Strategies. Current Biology 25(16): 2184-2188. (link* equal contributors 
News highlights include IFLScience and Audubon Magazine.

Strandburg-Peshkin, A.*, Farine, D.R.*, Couzin, I.D., Crofoot, M.C. (2015) Shared decision-making drives collective movement in wild baboons. Science 348(6241): 1358-1361. (link* joint first authors
Media highlights altmetric score including a nice article in National Geographic and a video by Untamed Science.

Farine, D.R., Whitehead, H. (2015) Constructing, conducting, and interpreting animal social network analysis. Journal of Animal Ecology 84(5): 1144-1163. (link)

Aplin, L.M., Firth, J.A., Farine, D.R., Voelkl, B., Crates, R.A., Culina, A., Garroway, C.J., Hinde, C.A., Kidd, L.R., Psorakis, I., Milligan, N.D., Radersma, R., Verhelst, B., Sheldon, B.C. (2015) Consistent individual differences in the social phenotypes of wild great tits (Parus major). Animal Behaviour 108: 117-127. (link)

Farine, D.R., Firth, J.A., Aplin, L.M., Crates, R.A., Culina, A., Garroway, C.J., Hinde, C.A., Kidd, L.R., Milligan, N.D., Psorakis, I., Radersma, R., Verhelst, B., Voelkl, B., Sheldon, B.C. (2015) The role of social and ecological processes in structuring animal populations: a case study from automated tracking of wild birds. Royal Society Open Science 2: 150057. (link)

Psorakis, I., Voelkl, B., Garroway, C.J., Radersma, R., Aplin, L.M., Crates, R.A., Culina, A., Farine, D.R., Firth, J.A., Hinde, C.A., Kidd, L.R., Milligan, N.D., Roberts, S.J., Verhelst, B., Sheldon, B.C. (2015) Inferring Social Structure from Temporal Data. Behavioural Ecology and Sociobiology 69(5): 857-866. (link)

Farine, D.R., Sheldon, B.C. (2015) Selection for territory acquisition is modulated by social network structure in a wild songbirdJournal of Evolutionary Biology 28(3): 547-556. (link)
Farine, D.R., Aplin, L.M., Sheldon, B.C., Hoppitt, W. (2015) Interspecific social networks promote information transmission in wild songbirdsProceedings of the Royal Society B 282: 20142804(link) 


Aplin, L.M., Farine, D.R., Morand-Ferron, J., Cockburn, A., Thornton, A., Sheldon, B.C. (2015) Experimentally induced innovations lead to persistent culture via conformity in wild birdsNature 518: 538-541. (link)
Media highlights: altmetric score.

Farine, D.R. (2015) Proximity as a proxy for interactions: issues of scale in social network analysisAnimal Behaviour 104: e1-e5. (link) 



Boogert, N.J.*, Farine, D.R.*, Spencer, K.A. (2014) Developmental stress predicts social network positionBiology Letters 10(10): 20140561. (link) * joint first authors


Godfrey, S.S., Anasri, T.H., Gardner, M.G., Farine, D.R., Bull, C.M. (2014) A contact based social network of lizards is defined by low genetic relatedness among strongly-connected individuals. Animal Behaviour 97: 35-43. (link)  


Farine, D.R., Aplin, L.M., Garroway, C.J., Mann, R.P., Sheldon, B.C. (2014) Collective decision-making and social interaction rules in mixed-species flocks of songbirdsAnimal Behaviour 95: 173-182. (link)
2nd most read article in Animal Behaviour September - December 2014  (see my short discussion on null models from this paper here)

Aplin, L.M.*, Farine, D.R.*, Mann, R.P., Sheldon, B.C. (2014) Individual-level personality influences social foraging and collective behaviour in wild birdsProceedings of the Royals Society B 218 (1789): 20141016. (link) * joint first authors
3rd most read article in Proceedings B in July 2014

Farine, D.R., Downing, C.P., Downing, P.A. (2014) Mixed-species associations can arise without heterospecific attractionBehavioural Ecology 25(3): 574-581(link) 


Farine, D.R. (2014) Measuring phenotypic assortment in animal social networks: weighted associations are more robust than binary edgesAnimal Behaviour 89: 141-153. (link) 

Farine, D.R. (2013) Animal Social Network Inference and Permutations for Ecologists in R using asnipeMethods in Ecology and Evolution 4(12): 1187–1194. (link) 

Farine, D.R., Lang, S.D. (2013) The early bird gets the worm: foraging strategies of wild songbirds lead to the early discovery of food sourcesBiology Letters 9(6): 20130578. (link)
Media links: (see AltMetric) Scientific American BBC News CBS News The Times Daily Mail Science Daily Orange (FR) & featured on BBC Winterwatch, and BBC, Irish, German & Canadian radio stations.

Aplin, L.M., Farine, D.R., Morand-Ferron, J., Cole, E.F., Cockburn, A., Sheldon, B.C. (2013) Individual personalities predict social behaviour in wild networks of great tits (Parus major)Ecology Letters 16(11): 1365–1372. (link)
See highlights of media coverage on AltMetric and recommendation by F1000 (featured in Top 10 & Hidden Jewels).

Farine, D.R. & Milburn, P.J. (2013) Social organisation of thornbill-dominated mixed-species flocks using social network analysisBehavioural Ecology and Sociobiology 67(2): 321-330. (link) 


Farine, D.R., Garroway, C.J., Sheldon, B.C. (2012) Social Network Analysis of mixed-species flocks: exploring the structure and evolution of interspecific social behaviourAnimal Behaviour 84: 1271-1277. (link) 


Aplin, L.M., Farine, D.R., Morand-Ferron, J., Sheldon, B.C. (2012) Social networks predict patch discovery in a wild population of songbirdsProceedings of the Royal Society B 279(1745): 4199-4205. (link)
Media links: Australian Geographic

Herr, A., O'Connell, D., Farine, D., Dunlop, M., Crimp, S., Poole, M. (2012) Watching grass grow in Australia: is there sufficient production potential for a biofuel industry? Biofuels, Bioproducts and Biorefining 6(3): 257-268. 


Farine, D.R., O'Connell, D.A., Raison, R.J., plus 12 co-authors (2011) An assessment of biomass for bioelectricity and biofuel, and for greenhouse gas emission reduction in AustraliaGlobal Change Biology Bioenergy 4(2): 148-175.
Top 15 most downloaded GCBB paper for 2012.

Rodriguez, L.C., May, B., Herr, A., Farine, D., O'Connell, D. (2011) Biofuel excision and the viability of ethanol production in the Green Triangle, Australia.  Energy Policy 39(4): 1951-1957. 


Farine, D.R., O'Connell, D.A., Grant, T., Poole, M.L. (2010) Opportunities for energy efficiency and biofuel production in Australian wheat farming systems.  Biofuels 1(4): 547-561. 

Book chapters

O'Connell, D., Farine, D.R., O'Connor, M.H., Dunlop, M. (2013) Integration and Implementation Research: Would CSIRO contribute to, and benefit from, a more formalised I2S approach? In Bammer, G. Disciplining Interdisciplinarity: Integration and Implementation Sciences for Researching Complex Real-World Problems. Australian National University Press, Canberra, Australia. (link)

Selected Talks

University of Zurich (Anthropology Departmental Seminar)

ASAB Easter Meeting (Plenary speaker)
CNRS Strasbourg (Seminar)
University of Bern (Departmental Seminar)
University of Zurich (Departmental Seminar)
University of Oulu (Departmental Seminar)

Gottinger Freilandtag (Plenary speaker)
University of Konstanz (Departmental Seminar)
University of Zurich (Departmental Seminar)
University of Oxford (EGI Seminar)

ECBB 2016, Vienna, Austria (Plenary: The collective side of animal personalities symposium)
ISBE 2016, Exeter, UK (contributed talk + From parents to peers: the consequences of information use for evolution, development and social behaviour symposium)
EcoSummit 2016, Montpellier, France (Mixed-species flocking symposium)

ABS 2015, Anchorage, Alaska (President's Symposium)
ASP 2015, Bend, Oregon (invited speaker in networks symposium)
IEC 2015, Cairns, Australia

University of Cambridge (Departmental Seminar)
University of California Davis (Departmental Seminar)
NIMBioS Investigative Workshop on Animal Social Networks (details here)
AmoveE, Raleigh NC (video)
ABS 2014, Princeton University
ISBE 2014, Hunter College, NYC
Simon Fraser University, Vancouver (Departmental Seminar)
Virginia Tech (Departmental Seminar)

University of California Davis (ABGG seminar)
Behaviour 2013, Newcastle
Animal Swarms Workshop (Israel)
University of Neuchatel (invited seminar)

University of York Centre for Complex Systems Analysis (Departmental seminar)


Journal of Animal Ecology
Animal Behaviour


Current Biology, PNAS, Trends in Ecology and Evolution, American Naturalist, Proceedings of the Royal Society B,  Ecology Letters, Biology Letters, Methods in Ecology and Evolution, Ecology, Scientific Reports, Journal of the Royal Society Interface, PLOS Computational Biology, Journal of Animal Ecology, Animal Behaviour, Behavioural Ecology, Journal of Ornithology, Ibis, Conservation Biology, Canadian Journal of Zoology, New Generation Computing, Population Ecology, PLOS One, Behavioural Ecology and Sociobiology, Current Zoology, The Auk, plus others (I'm losing track). Also NERC, NSF, and other granting agencies.