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Most articles are posted as pdfs, for the convenience of people making personal copies for their own use.  If you wish copies of articles for commercial purposes or for distribution to a class, please write the publishers.  For a selection of the best-cited papers published prior to 2000, please see the
link at the bottom of the page. My CV contains a complete bibliography, and my Google Scholar Profile tallies how other people have used my work.
I may be able to provide you with copies of other articles I have written that are not posted here -- write me at bpattersonfieldmuseum.org. If some of these links ask you to sign in, sorry!  Try accessing them with Google Chrome.
In Press

Patterson, B.D. & C. Lopez W. In press.  Mammals/Mamíferos.  In: Perú: Cordillera Escalera (N. Pitman et al., eds.). Rapid Biological and Social Inventories No. 26,

Patterson, B.D., M.F. Smith & P. Teta. In press. Abrothrix, in: Mammals of South America, Vol. 2: Rodents (J.L. Patton, U.F.J. Pardiñas, and G. D’Elía, eds.). University of Chicago Press, Chicago.

Patterson, B.D.  In press.  Family Caenolestidae, shrew-opossums, in: Handbook of Mammals of the World, Vol. 5. Monotremes and Marsupials (D.E. Wilson, ed.). Lynx Editions, Barcelona.

Dittmar, K., S.F. Morse, C.W. Dick & B.D. Patterson. In press. Bat fly evolution from the Eocene to the Present (Hippoboscoidea, Streblidae and Nycteribiidae), in: Parasite Diversity and Diversification: Evolutionary Ecology Meets Phylogenetics (S. Morand, B. Krasnov and T. Littlewood, eds). Cambridge University Press, Cambridge.

Ramírez Chaves, H.E. & B.D. Patterson. In press. Mustela felipei (Carnivora, Mustelidae). Mammalian Species.


Artibeus lituratus

An Artibeus lituratus
peers down from its perch in lowland tropical forest, part of the fauna analyzed by Cisneros et al. (2014)

Velazco., P.M. & B.D. Patterson. 2014. Two new species of yellow-shouldered bats, Genus Sturnira Gray, 1842 (Chiroptera, Phyllostomidae) from Costa Rica, Panama and western Ecuador.  ZooKeys 402:43-66.
pdf (6.6 Mb)

Cisneros, L.M., K.R. Burgio, L.M. Dreiss, B.T. Klingbeil, B.D. Patterson, S.J. Presley & M.R. Willig. 2014. Multiple dimensions of bat biodiversity along an extensive tropical elevational gradient.  Journal of Animal Ecology doi: 10.1111/1365-2656.12201. pdf


Oct 2013 cover

The 5th known species of northern shrew-opossum, Caenolestes sangay Ojala-Barbour et al, 2013

Ojala-Barbour, R., C.M. Pinto, J. Brito M., L. Albuja V., T.E. Lee, Jr. & B.D. Patterson. 2013. A new species of shrew-opossum (Caenolestidae, Paucituberculata) with a phylogeny of extant caenolestids. Journal of Mammalogy 94:967-982 + cover. pdf  SOM1  SOM2

Upham, N.S., R. Ojala-Barbour, J. Brito, P.M. Velazco & B.D. Patterson›*››. 2013. Transitions between Andean and Amazonian centers of endemism in the radiation of some arboreal rodents. BMC Evolutionary Biology 13:191. *corresponding author. link to Open Access 

Velazco, P.M. & B.D. Patterson*. 2013. Diversification of the Yellow-shouldered bats, Genus Sturnira (Chiroptera: Phyllostomidae) in the New World tropics. Molecular Phylogenetics and Evolution 68:683-698.
*corresponding author: pdf (4.07 Mb)

Dubach, J.M., M.B. Briggs, P.A. White, B.A. Ament & B.D. Patterson*. 2013. Genetic perspectives on "Lion Conservation Units" in Eastern and Southern Africa. Conservation Genetics 
14:741-755. *corresponding author:
pdf  SOM

Patterson, B. D. 2013. What's in a name?
(Re)naming the lions of Africa. In the Field 84(2):9

Packer, C.
et al. (numerous co-authors including BDP). 2013. Conserving large carnivores: Dollars and fence. Ecology Letters 16:635-641 pdf (234 kb)  SOM figs   SOM tabs

Morse, S.F., S.E, Bush, B.D. Patterson, C.W. Dick, M.E. Gruwell & K. Dittmar. 2013
Evolution, multiple acquisition, and localization of endosymbionts in bat flies (Hippoboscoidea, Streblidae and Nycteribiidae). Applied and Environmental Microbiology
79:2952-2961 pdf (2.8 Mb)  SOM (66 kb)

Patterson, B.D. 2013. Squirrels of the World [book review]. Journal of Wildlife Management 77:643–644 pdf (59 kb)

Amori, G., F. Chiozza, B.D. Patterson, C. Rondinini, J. Schipper & L. Luiselli. 2013. Correlates of species richness and the distribution of South American rodents, with conservation implications. Mammalia 77:1-19. pdf (588 kb)


The University of Chicago Press volume that resulted from IMC-10 in Mendoza, Argentina

Patterson, B.D. & P.W. Webala.  2012.  Keys to the bats (Mammalia: Chiroptera) of East Africa. Fieldiana: Life and Earth Sciences, 6:1-63. pdf (6.7 Mb)

Morse, S.F., C.W. Dick, B.D. Patterson & K. Dittmar. 2012. Some like it hot -- Evolution and ecology of novel endosymbionts in bat flies of cave-roosting bats (Hippoboscoidea, Nycterophiliinae). Applied and Environmental Microbiology 78:8639-8649. pdf (2.7 Mb)

Morse, S.F., K.J. Olival, M. Kosoy, S. Billeter, B.D. Patterson, C.W. Dick & K. Dittmar. 2012. Global distribution and genetic diversity of Bartonella in bat flies (Hippoboscoidea, Streblidae, Nycteribiidae). Infection, Genetics and Evolution 12:1717-1723. pdf (588 kb)

Presley, S.J., L.M.Cisneros, B.D. Patterson & M.R. Willig. 2012. Vertebrate meta-community structure along an extensive elevational gradient in the tropics: a comparison of bats, rodents, and birds. Global Ecology and Biogeography 21:968-976. pdf (950 kb)

Pilosof, S., C.W. Dick, C. Korine, B.D. Patterson & B.R. Krasnov. 2012. Effects of anthropogenic disturbance and climate on patterns of bat fly parasitism. PLoS One 7(7): e41487. html

Jiménez, F.A. & B.D. Patterson. 2012. A new species of Pterygodermatites (Nematoda: Rictulariidae) from the Incan shrew opossum, Lestoros inca. Journal of Parasitology, 98(3):604-607. pdf (939 kb)

Patterson, B.D. & L.P. Costa (eds.). 2012.  Bones, Clones, and Biomes: The history and geography of Recent Neotropical mammals University of Chicago Press, Chicago. 419 pp. webpage --  J Mamm review
  • Patterson, B.D. & L.P. Costa. 2012.  Introduction. Pp. 1-5 in Bones, Clones, and Biomes: The history and geography of Recent Neotropical mammals (B.D. Patterson & L.P. Costa, eds.).  University of Chicago Press, Chicago.
  • Solari, S., P.M. Velazco & B.D. Patterson. 2012. Hierarchical organization of Neotropical mammal diversity and its historical basis. Pp. 145-156 in Bones, Clones, and Biomes: The history and geography of Recent Neotropical mammals (B.D. Patterson & L.P. Costa, eds.). University of Chicago Press, Chicago.
  • Patterson, B.D., S. Solari & P.M. Velazco. 2012. The role of the Andes in the diversification and biogeography of Neotropical mammals. Pp. 351-378 in Bones, Clones, and Biomes: The history and geography of Recent Neotropical mammals (B.D. Patterson & L.P. Costa, eds.).  University of Chicago Press, Chicago pdf (326 kb).

Patterson, B. D. 2012. The man-eating lions of Tsavo and the science behind the killings. Open Skies Apr 2012:84-93. pdf (2.4 Mb)

Upham, N.S. & B.D. Patterson. 2012. Diversification and biogeography of the Neotropical caviomorph lineage Octodontoidea (Rodentia: Hystricognathi). Molecular Phylogenetics and Evolution 63:417-429. pdf (1 Mb)

Patterson, B.D. 2012. Su-Lin, America's first panda. In the Field 83(1):12. pdf (300 kb)


Incidence and average local abundance for 29 Egyptian rodents, from Abu Baker & Patterson (2011)

Meyer, C.F.J. et al. (numerous co-authors including BDP). 2011. Accounting for detectability improves estimates of species richness in tropical bat surveys. Journal of Applied Ecology 48:777-787. pdf (590 kb)

Laurance, W.F. et al. (numerous co-authors including BDP). 2011. Global warming, elevational ranges and the vulnerability of tropical biota.  Biological Conservation 144:548-557. pdf (870 kb)

Abu Baker, M. & B.D. Patterson. 2011. Patterns in the local assembly of Egyptian rodent faunas: co-occurrence and nestedness. Journal of Arid Environments 75:14-19. pdf (504 kb)


Map showing extent of Patagonian glaciers and exposed continental shelf at the
Late Glacial Maximum in South America,
(Fig. 1 from Patterson 2010).

Abu Baker, M. & B.D. Patterson. 2010.  Patterns in the local assembly of Egyptian rodent faunas: areography and species combinations.  Mammalian Biology 75:510-522. pdf (1.1 Mb)

Patterson, B.D. 2010.  Climate change and faunal dynamics in the uttermost part of the earth.  Molecular Ecology 19:3019-3021. pdf (222 kb)

Velazco, P.M., A.L. Gardner & B.D. Patterson. 2010. Systematics of the Platyrrhinus helleri complex (Chiroptera: Phyllostomidae), with descriptions of two new species.  Zoological Journal of the Linnean Society 159:785-812. pdf (1.49 Mb)


Earthwatch lions Bahati and Kabochi on the cover of the 10 Nov 2009 issue of PNAS (Yeakel et al. 2009).
Yeakel, J.D., B.D. Patterson, K. Fox-Dobbs, M.M. Okumura, T.E. Cerling, J.W. Moore, P.L. Koch & N.J. Dominy. 2009. Cooperation and individuality among man-eating lions. Proceedings of the National Academy of Sciences 106:19040-19043 + cover. pdf (833 kb)

Williams, P.R., B.D. Patterson, M. Thayer & K. Angielczyk. 2009. Museum curators and the AAUP. Illinois Academe Fall 2009:2. pdf (584 kb)

Patterson, B.D., C.W. Dick & K. Dittmar. 2009. Nested distributions of bat flies (Diptera: Streblidae) on Neotropical bats: artifact and specificity in host-parasite studies. Ecography 32:481-487. pdf (159 kb)
Dittmar, K., C.W. Dick, B.D. Patterson, M.F. Whiting & M. Gruwell. 2009. Pupal deposition and ecology of bat flies (Diptera: Streblidae): Trichobius sp. (caecus group) in a Mexican cave habitat. Journal of Parasitology 95:308-314. pdf (1021 kb)

Three bat fly species commonly found together on the bat Carollia perspicillata (Fig.1 of Patterson et al. 2009 Ecography)

Velazco, P.M. & B.D. Patterson. 2008. Phylogenetics and biogeography of the broad-nosed bats, Genus Platyrrhinus (Chiroptera: Phyllostomidae). Molecular Phylogenetics and Evolution 49:749-759. pdf (1.3 Mb)
Dick, C.W. & B.D. Patterson. 2008. An excess of males: skewed sex ratios in bat flies (Diptera: Streblidae). Evolutionary Ecology 22:757-769. pdf (216 kb).

Fig. 1a from Schipper et al. (2008), showing the global distribution of mammalian species richness
Schipper, J. et al. (numerous co-authors including BDP). 2008. The status of the world's land and marine mammals: diversity, threat and knowledge. Science 322:225-230 + online suppl. pdf (655 kb) supplement (494 kb)
Patterson, B.D., C.W. Dick & K. Dittmar. 2008b. Parasitism by bat flies (Diptera: Streblidae) on Neotropical bats: effects of host body size, distribution and abundance. Parasitology Research 103:1091-1100. pdf (355 kb).
Patterson, B.D., C.W. Dick & K. Dittmar. 2008a. Sex biases in parasitism of Neotropical bats by bat flies (Diptera: Streblidae). Journal of Tropical Ecology 124:387-396. pdf (110 kb).
Fig. 1 from Patterson & Velazco (2008) showing locations of molecular samples for six species of
(Rodentia, Echimyidae) used in phylogenetic analyses
Patterson, B.D. & P.M. Velazco. 2008. Phylogeny of the rodent genus Isothrix (Hystricognathi, Echimyidae) and its diversification in Amazonia and the Eastern Andes. Journal of Mammalian Evolution 15:181-201. pdf (534 kb).

Patterson, B.D. & M.A. Rogers 2008. Microbiotheria, Pp. 117-119 in Mammals of South America, Volume 1: Marsupials, Xenarthrans, Shrews, and Bats (A.L. Gardner, ed.). University of Chicago Press, Chicago, 912 pp. pdf (720 kb).

Patterson, B.D. 2008b. Paucituberculata, Pp. 119-120 in Mammals of South America, Volume 1: Marsupials, Xenarthrans, Shrews, and Bats (A.L. Gardner, ed.). University of Chicago Press, Chicago, 912 pp.
pdf (662 kb).

Timm, R.M. & B.D. Patterson. 2008. Caenolestes, Pp. 120-124 in Mammals of South America, Volume 1: Marsupials, Xenarthrans, Shrews, and Bats (A.L. Gardner, ed.). University of Chicago Press, Chicago, 912 pp.
pdf (822 kb).

Patterson, B.D. 2008a. Rhyncholestes, Pp. 126-127 in Mammals of South America, Volume 1: Marsupials, Xenarthrans, Shrews, and Bats (A.L. Gardner, ed.). University of Chicago Press, Chicago, 912 pp.
pdf (652 kb)
Veron, G., B.D. Patterson & R. Reeves. 2008. Global diversity of mammals (Mammalia) in freshwater. In Freshwater Animal Diversity Assessment (E.V. Balian, C. Leveque, H. Segers, K. Martens, eds.). Hydrobiologia 595:607-617. pdf (295 kb).


A tale of two lions--Bahati (a) and Charlie (b), both six years old and from Tsavo, but raised
under different climatic regimens
(Fig. 1 in Patterson [2007] Evol. Biol. article)
Patterson, B.D. 2007. A new 'Age of Discovery' for mammals [Review of Mammal Species of the World, 3rd ed]. Journal of Mammalian Evolution 14:67-69. pdf (166 kb).

Patterson, B.D., C.W. Dick & K. Dittmar. 2007. Roosting habits of bats affect their parasitism by bat flies (Diptera: Streblidae). Journal of Tropical Ecology 23:177-189.
pdf (247 kb).

Patterson, B.D. 2007. The meaning of manes. Hunting Insight Africa 9:41-45.
pdf (1.08 Mb).

Dick, C.W. & B.D. Patterson. 2007. Against all odds: explaining host specificity in dispersal-prone ectoparasites. International Journal of Parasitology 37:871-876.
pdf (340 kb).

Patterson, B.D. 2007. On the nature and significance of variability in lions (Panthera leo). Evolutionary Biology 34:55-60.
pdf (258 kb).

Vargas Espinoza, A. & B.D. Patterson. 2007. Comunidades de murcielagos montanos en Bolivia. Pp 82-86, In L.F. Aguirre (Ed.). Historia Natural, Distribucion y Conservacion de los Murcielagos de Bolivia. Editorial, Centro de Ecologi­a y Difusion Simon I. Patinyo, Santa Cruz, Bolivia.
pdf (945 kb).

Patterson, B.D. 2007. The quintessential naturalist: honoring the life and legacy of Oliver P. Pearson [Review of Kelt et al. eds, University of California Publications in Zoology]. Mastozoologí­a Neotropical 14:299.
pdf (258 kb).

New species Isothrix barbarabrownae Patterson & Velazco 2006 on the cover of Mastozoología Neotropical.
Drawing by Velizar Simeonovski.
Patterson, B.D. & P.M. Velazco. 2006. A distinctive new cloud-forest rodent (Hystricognathi: Echimyidae) from the Manu Biosphere Reserve, Peru. Mastozoología Neotropical 13:175-191 + cover. pdf (799 kb).

Patterson, B.D., D.F. Stotz & S. Solari (eds.). 2006. Mammals and Birds of the Manu Biosphere Reserve, Peru. Fieldiana: Zoology, new series 110:1-49. pdf (93 kb).

Patterson, B.D., D.F. Stotz & S. Solari. 2006. Biological surveys and inventories in Manu. Pp. 3-12 in Mammals and Birds of the Manu Biosphere Reserve, Peru (B.D. Patterson, D.F. Stotz, and S. Solari, eds.). Fieldiana: Zoology, new series 110.
pdf (1.3 Mb).

Solari, S., V. Pacheco, L. Luna, P.M. Velazco & B.D. Patterson. 2006. Mammals of the Manu Biosphere Reserve. Pp. 13-22 in Mammals and Birds of the Manu Biosphere Reserve, Peru (B.D. Patterson, D.F. Stotz, and S. Solari, eds.). Fieldiana: Zoology, new series 110.
pdf (221 kb).

Dick, C. W. & B. D. Patterson. 2006. Bat flies - obligate ectoparasites of bats. Pp. 179-194 In Micromammals and macroparasites: from evolutionary ecology to management (S. Morand, B. Krasnov, and R. Poulin, eds.). Springer-Verlag, Tokyo. pdf (121 kb)


Earthwatch lion "Cassius" on the cover of Journal of Mammalogy in April 2006

Patterson, B.D., R.W. Kays, S.M. Kasiki & V.M. Sebestyen. 2006. Developmental effects of climate on the mane of the lion (Panthera leo). Journal of Mammalogy 87:193-200 + cover. pdf (457 kb).  
D'Elía, G., L. Luna, E.M. González & B.D. Patterson. 2006. On the Sigmodontinae radiation (Rodentia, Cricetidae): An appraisal of the phylogenetic position of Rhagomys. Molecular Phylogenetics and Evolution 38:558-564.
pdf (265 kb)

Patterson, B.D. 2006. [Review] Foundations of Biogeography: classic papers with commentaries, edited by Lomolino et al. Perspectives in Biology and Medicine 49:152-154.
pdf (65 kb).

Earthwatch lion "Bahati" on the cover of a popular article in April 2005

Patterson, B.D. 2005. Living with lions in Tsavo, or some notes on managing man-eaters. Travel News & Lifestyle (East Africa) 129 (Feb 2005): 28-31 + cover. pdf (889 kb).

Dubach, J, B.D. Patterson, M.B. Briggs, K. Venzke, J. Flammand, P. Stander, L. Scheepers & R. Kays. 2005. Molecular genetic variation across the southern and eastern geographic ranges of the African lion, Panthera leo. Conservation Genetics 7:15-24.
pdf (314 kb).

Patterson, B.D. 2005. [Review] Evolving Eden: an illustrated guide to the evolution of the African large-mammal fauna, by A. Turner and M. Anton. Journal of Mammalogy 86:1052-1053.
pdf (93 kb).

Patterson, B.D., N. Yamaguchi, J.M. Dubach & D. York. 2005. Molecular genetics and morphological variation of lions (Panthera leo). African Lion Newsletter 6(2):17-23.
pdf (296 kb).

At long last, the book was finally published in January 2004!

Patterson, B.D. 2004. The lions of Tsavo: exploring the legacy of Africa's notorious man-eaters. McGraw-Hill, New York, 231 pp. examine book here

Patterson, B.D. 2004. Maneless and misunderstood: the lions of Tsavo. Earthwatch Institute 23(2):12-15. pdf (920 kb)

Patterson, B.D., S.M. Kasiki, E. Selempo & R.W. Kays. 2004. Livestock predation by lions (Panthera leo) and other carnivores on ranches neighboring Tsavo National Park, Kenya. Biological Conservation 119:507-516.  pdf (430 kb)

Patterson, B.D. 2004. Fatal attraction: a predilection for livestock threatens Tsavo's lions. Earthwatch Institute - Australia April/May 2004:6-7. pdf (840 kb).

Solari, S., R.A. Van Den Bussche, S.R. Hoofer & B.D. Patterson. 2004. Geographic distribution, ecology, and phylogenetic affinities of Thyroptera lavali Pine 1993. Acta Chiropterologica 6:293-302. pdf (362 kb).

Distributions of 1700 species of mammals native to the Western Hemisphere

Patterson, B.D., E.J. Neiburger & S.M. Kasiki. 2003. Tooth breakage and dental disease as causes of carnivore-human conflicts. Journal of Mammalogy 84:190-196. pdf (340 kb).

Patterson, B.D. & S.M. Kasiki. 2003. Expedition briefing: Lions of Tsavo. Earthwatch Institute, Maynard, Massachusetts, 30 pp + appendices.
pdf (970 kb)..

Luna W., L. & B.D. Patterson. 2003. A remarkable new mouse from Peru (Muridae: Sigmodontinae), with comments on the affinities of Rhagomys rufescens (Thomas, 1886). Fieldiana: Zoology, new ser. 101:1-24.
pdf (1.2 Mb).

Patterson, B.D., M.R. Willig & R.D. Stevens. 2003. Trophic strategies, niche partitioning, and patterns of ecological organization. Pp. 536-579 in Ecology of bats (T. H. Kunz and M. B. Fenton, ed.). University of Chicago Press, Chicago. pdf (4.6 Mb)

Willig, M.R., B.D. Patterson & R.D. Stevens. 2003. Patterns of range size, richness, and body size. Pp. 580-621 in Ecology of bats (T. H. Kunz and M. B. Fenton, ed.). University of Chicago Press, Chicago.
pdf (4.3 Mb).

Patterson, B. D., G. Ceballos, W. Sechrest, M. Tognelli, T. Brooks, L. Luna, P. Ortega, I. Salazar & B. E. Young. 2003. Digital Distribution Maps of the Mammals of the Western Hemisphere, version 1.0. NatureServe, Arlington, Virginia, USA.
shp files.

Components of lion manes (Fig. 1 of 2002 Canadian Journal of Zoology paper).
Neiburger, E.J. & B.D. Patterson. 2002. A forensic dental determination of serial killings by three African lions. General Dentistry 50:40-43. pdf (660 kb).

Kays, R.W. & B.D. Patterson. 2002. Mane variation in African lions and its social correlates. Canadian Journal of Zoology 80:471-478.
pdf (235 kb).

Patterson, B.D. 2002. On the continuing need for scientific collecting of mammals. Mastozoologí­a Neotropical  9:253-262.
pdf (280 kb).

Escalating estimates over the last half-century of the number of living mammal species
(from Diversity and Distributions, 2001)

Patterson, B.D. 2001. [Review] Prides: the lions of Moremi, by C. Harvey and P. Kat. Journal of Mammalogy 82:248-250. pdf (37 kb).

Patterson, B.D. & E.J. Neiburger. 2001. Lion with a sore tooth. Nature Australia 26(12):12. pdf (580 kb).

Patterson, B.D. 2001. Fathoming tropical biodiversity: the continuing discovery of Neotropical mammals. Diversity & Distributions 7:191-196. pdf (280 kb).

New York State Dental Journal, December 2000, containing article on Tsavo and Mfuwe man-eating lions

González, E.M. & B.D. Patterson. 2000. Un nuevo nombre genérico para Thomasomys pictipes Osgood, 1933 (Rodentia: Sigmodontinae). XIV Journadas Argentinas de Mastozoología, Salta, Argentina 1999: 34-35. pdf (380 kb)

Patterson, B.D. & W. Atmar. 2000. Analyzing species composition in fragments. Pp. 9-24 in Isolated vertebrate communities in the tropics (G. Rheinwald, ed.). Bonner Zoologische Monographen 46, Bonn, Germany, 400 pp.
pdf (970 kb).

Sutton, D.A. & B.D. Patterson. 2000. Geographic variation of the western chipmunks Tamias senex and Tamias siskiyou, with two new subspecies from California. Journal of Mammalogy 81: 299-316.
pdf (380 kb).

Patterson, B.D. 2000a. [Review] Walkers Mammals of the World, sixth ed. International Journal of Primatology 21:561-563.
pdf (345 kb).

Neiburger, E.J. & B.D. Patterson. 2000. Man eating lions...a dental link. Journal of the American Association of Forensic Dentists 24(7-9):1-3.
pdf (200 kb).

Patterson, B.D. 2000b. Patterns and trends in the discovery of new Neotropical mammals. Diversity & Distributions 6:145-151.
pdf (415 kb).

Neiburger, E.J. & B.D. Patterson. 2000. The man-eaters with bad teeth. New York State Dental Journal 66(10):26-29 + cover.
cover (370 kb) and pdf (870 kb).
Subpages (1): Some Prior publications
Bruce Patterson,
Aug 23, 2012, 10:52 AM