1. Abhinav Banerjee, Saanya Yadav, Vedanth Shree Vidwath, Simanta Kalita, Sarit S Agasti, Himanshu Joshi*, Mahipal Ganji*. Proximity‐Based Super‐Resolution Imaging Enabled by DNA Base‐Stacking Interactions. Small, 2026.
2. Ramkumar Balaji, Himanshu Joshi*, Basant K Patel*. Elucidating the conformational dynamics of the mitochondrial localization signal, M3, of TDP-43 and accessing potential inhibitors using molecular docking and simulation. Computational Biology and Chemistry, 123, 109029, 2026.
3. Rishi Raj, Yiao Jiang, Rahul Kumar Jha, Eva Marie Y Moresco, Himanshu Joshi, Zhao Zhang, Bruce Beutler. Structural insights into GM4951 as a lipid droplet GTPase regulating hepatic lipid metabolism. Nature Communications 16, 11458, 2025.
4. Nikhil Tuti, Rashmi Ranjan Khatua, Saanya Yadav, Sharan Shanmuga Vuppaladadium Rathnam, Jitender Jangra, Subha Narayan Rath, Himanshu Joshi*, Faiz Ahmed Khan*, Anindya Roy*. Targeting DNA termini with a small-molecule natural product analogue: A DNA end-binder (DEB) to augment DNA double-strand breaks, Nucleic Acids Research, 53(19), 2025.
5. Pooja Dahiya, Ankit Singh, Soumya Aggarwal, Priyanka Yadav, Himanshu Joshi, Manoj Munde. Protamine and Zinc Binding-Induced Conformational Effects on Insulin Analogs: An Integrated Experimental and Computational Study. Physical Chemistry Chemical Physics, 27 (36), 19149-19162, 2025.
6. Sri Amruthaa Sankaranarayanan, Dokkari Nagalaxmi Yadav, Saanya Yadav, Aditya Srivastava, Sai Rachana Pramatha, Venkata Rao Kotagiri, Himanshu Joshi*, Aravind Kumar Rengan. Tailoring Phage Nanosomes for Enhanced Theranostic Properties of Near Infrared Dyes. Langmuir, 40 (32), 16743–16756, 2024.
7. Hyeonji Oh, Yu-Ming Tu, Laximicharan Samineni, Sophie De Respino, Behzad Mehrafrooz, Himanshu Joshi, Lynnicia Massenburg, Horacio Lopez-Marques, Nada Elessawy, Woochul Song, Harekrushna Behera, Raman Dhiman, Veda Sheersh Boorla, Kartik Kher, Yi-Chih Lin, Costas Maranas, Aleksei Aksimentiev, Benny D. Freeman, Manish Kumar. Dehydrated Biomimetic Membranes with Skin-like Structure and Function. ACS Applied Materials Interfaces, 16(16), 20865–20877, 2024.
8. Vini D. Meshram, Ramkumar Balaji, Preethi Saravanan, Yashashwini Subbamanda, Waghela Deeksha, Akarsh Bajpai, Himanshu Joshi, Anamika Bhargava, Basant K. Patel. Computational insights into the mechanism of EGCG’s binding and inhibition of the TDP-43 aggregation. Chemical Biology & Drug Design, 2024.
9. Jie Shen, Deepa R. Zhongyan Li, Hyeonji Oh, Harekrushna Behera, Himanshu Joshi, Manish Kumar, Aleksei Aksimentiev, Huaqiang Zeng. Sulfur‐Containing Foldamer‐Based Artificial Lithium Channels. Angewandte Chemie International Edition, e202305623, 2023.
10. Akshit Prajapatia, Aditya Srivasatavaa, Sri Amruthaa Sankaranarayanana, Monika Pebama, Himasree Buddhirajua, Paloma Patraa, Priyanka Yadav, Himanshu Joshi and Aravind Kumar Rengan. High-Affinity DNA Nanomatrix: A Platform Technology for Synergistic Drug Delivery and Photothermal Therapy. ACS Macro Letters, 12 (2), 255-262, 2023.
11. Yi Li, Christopher Maffeo, Himanshu Joshi, Aleksei Aksimentiev, Brice Menard, and Rebecca Schulman. Leakless end-to-end transport of small molecules through micron-length DNA nanochannels. Science Advances, 8(36), 1-9, 2022.
1. Diana Morzy, Himanshu Joshi, Sarah E. Sandler, Aleksei Aksimentiev, Ulrich F Keyser, "Membrane Activity of a DNA-Based Ion Channel Depends on the Stability of Its Double-Stranded Structure" Nano Letters, 21 (22), 9789–9796, 2021.
2. Dan Qiao*, Himanshu Joshi, H. Zhu, F. Wang, Y. Xu, J. Gao, F. Huang, A. Aksimentiev and J. Feng. “Synthetic macrocycle nanopore for potassium-selective transmembrane transport”. Journal of the American Chemical Society, 143 (39), 15975-15983, 2021. [Featured in cover image]
3. Sarah E. Ochmann, Himanshu Joshi, Ece Büber, Henri G Franquelim, Pierre Stegemann, Barbara Saccà, Ulrich F Keyser, Aleksei Aksimentiev, Philip Tinnefeld. “DNA Origami Voltage Sensors for Transmembrane Potentials with Single-Molecule Sensitivity”. Nano Letters, 21 (20), 86348641, 2021.
4. Diana Morzy, Roger Rubio-Sánchez, Himanshu Joshi, Aleksei Aksimentiev, Lorenzo Di Michele, Ulrich F Keyseriana. “Cations Regulate Membrane Attachment and Functionality of DNA Nanostructures”. Journal of the American Chemical Society, 2021. DOI:10.1021/jacs.1c0016
5. Sioned F. Jones, Himanshu Joshi, Stephen J. Terry, Jonathan R. Burns, Aleksei Aksimentiev, Ulrike S. Eggert, and Stefan Howorka. “Understanding the fundamental interaction of hydrophobic DNA duplexes with artificial and biological membranes”. Journal of the American Chemical Society, 2021. DOI:10.1021/jacs.0c13235.
6. Arundhati Roy, Jie Shen, Himanshu Joshi, Woochul Song, Yu-Ming Tu, Ratul Chowdhury, Ruijuan Ye, Ning Li, Changliang Ren, Manish Kumar, Aleksei Aksimentiev, and Huaqiang Zeng. “Foldamer-Based Ultraperme- able and Highly Selective Artificial Aquaporins that Exclude Protons”. Nature Nanotechnology, 2021.
7. Kristina Hubner, Himanshu Joshi, Aleksei Aksimentiev, Fernando D. Stefan, Philip Tinnefeld, and Guillermo P. Acuna. “Determining the inplane orientation and binding mode of single fluorescent dyes in DNA origami structures”. ACS Nano, 15(3), 5109–5117, 2021.
8. David Winogradoff, Pin-Yi Li, Himanshu Joshi, Lauren Quednau, Christopher Maffeo,and Aleksei Aksimentiev. Chiral Systems Made from DNA, Advanced Science 8 (5), 200311 2021.
9. Adnan Choudhary, Himanshu Joshi, Han-Yi Chou, Kumar Sarthak, James Wilson, Christopher Maffeo and Aleksei Aksimentiev. “High- Fidelity Capture, Threading and Infinite-Depth Sequencing of Single DNA Molecules with a Double-Nanopore System”. ACS Nano, 14 (11), 15566-15576, 2020.
10. Diana Sobota, Himanshu Joshi, Alexander Ohmann, Aleksei Aksimentiev, and Ulrich F Keyser. Tailoring interleaflet lipid transfer with a DNA-based synthetic enzyme. Nano Letters, 20(6):4306–4311, 2020.
11. Arundhati Roy*, Himanshu Joshi*, Ruijuan Ye, Jie Shen, Feng Chen, Aleksei Aksimentiev, and Huaqiang Zeng. Polyhydrazide-based organic nanotubes as efficient and selective artificial iodide channels. Angewandte Chemie International Edition, 59(12):4806–4813, 2020.
12. Prabhat Tripathi, Liang Shuai, Himanshu Joshi, Hirohito Yamazaki, William H Fowle, Aleksei Aksimentiev, Hicham Fenniri, and Meni Wanunu. Rosette nanotube porins as ion selective transporters and single-molecule sensors. Journal of the American Chemical Society, 142(4):1680–1685, 2020
13. Woochul Song, Himanshu Joshi, Ratul Chowdhury, Joseph S. Najem, Yue xiao Shen, Chao Lang, Codey B. Henderson, Yu-Ming Tu, Megan Farell, Megan E. Pitz, Costas D. Maranas, Paul S. Cremer, Robert J. Hickey, Stephen A. Sarles, Jun li Hou, Aleksei Aksimentiev, and Manish Kumar. Artificial water channels enable fast and selective water permeation through water-wire networks. Nature Nanotechnology, 15:73–79, 2020.
14. Alexander Ohmann, Kerstin Go ̈pfrich, Himanshu Joshi, Rebecca F Thompson, Diana Sob- ota, Neil A Ranson, Aleksei Aksimentiev, and Ulrich F Keyser. Controlling aggregation of cholesterol-modified DNA nanostructures. Nucleic Acids Research, 47(21):11441–11451, 2019.
15. Supriyo Naskar, Himanshu Joshi, Banani Chakraborty, Nadrian C Seeman, and Prabal K Maiti. Atomic structures of RNA nanotubes and their comparison with DNA nanotubes. Nanoscale, 11(31):14863–14878, 2019.
16. Ramkumar Balasubramanian, Sohini Pal, Himanshu Joshi, Anjana Rao, Akshay Naik, Manoj Varma, Banani Chakraborty, and Prabal K Maiti. DNA translocation through hybrid bilayer nanopores. Journal of Physical Chemistry C, 123(18):11908-11916, 2019.
17. Supriyo Naskar, Mounika Gosika, Himanshu Joshi, and Prabal K Maiti. Tuning the stability of DNA nanotubes with salt. Journal of Physical Chemistry C, 123(14):9461-9470, 2019.
18. Yiming Yang, Aaron Alford, Veronika Kozlovskaya, Shidi Zhao, Himanshu Joshi, Eunjung Kim, Shuo Qian, Volker Urban, Donald Cropek, Aleksei Aksimentiev, et al. Effect of temperature and hydrophilic ratio on the structure of poly (N-vinylcaprolactam)-block-poly (dimethylsiloxane)-block-poly (n-vinylcaprolactam) polymersomes. ACS Applied Polymer Materials, 1(4):722–736, 2019.
19. Patrick M Arnott, Himanshu Joshi, Aleksei Aksimentiev, and Stefan Howorka. Dynamic inter- actions between lipid-tethered DNA and phospholipid membranes. Langmuir, 34(49):15084– 15092, 2018. [Featured in cover image]
20. Himanshu Joshi and Prabal K Maiti. Structure and electrical properties of DNA nanotubes embedded in lipid bilayer membranes. Nucleic Acids Research, 46(5):2234-2242, 11, 2017. [Featured in cover image]
21. Himanshu Joshi, Dhiraj Bhatia, Yamuna Krishnan, and Prabal K Maiti. Probing the structure and in silico stability of cargo loaded DNA icosahedra using md simulations. Nanoscale, 9(13):4467–4477, 2017.
22. Dhiraj Bhatia, Senthil Arumugam, Michel Nasilowski, Himanshu Joshi, Christian Wunder, Val ́erie Chambon, Ved Prakash, Chlo ́e Grazon, Brice Nadal, Prabal K Maiti, Ludger Johannes, Benoit Dubertret, and Yamuna Krishnan. Quantum dot-loaded monofunctionalized DNA icosahedra for single-particle tracking of endocytic pathways. Nature Nanotechnology, 11(12):1112–1119, 2016.
23. Himanshu Joshi, Atul Kaushik, Nadrian C Seeman, and Prabal K Maiti. Nanoscale structure and elasticity of pillared DNA nanotubes. ACS Nano, 10(8):7780-7791, 2016.
24. Himanshu Joshi, Anjan Dwaraknath, and Prabal K Maiti. Structure, stability and elasticity of DNA nanotubes. Physical Chemistry - Chemical Physics, 17(2):1424-1434, 2015.