Preprint. Tu BN**, Mas-Carrio E, Hodecek J***, Khelidj N**, Casartelli M, Rasmann S, Fumagalli L, Losapio G.
Glacier retreat reshapes trophic networks: interaction turnover outpaces species turnover over space-time
bioRxiv, April 15 2026
Ferrari A, Cilia G, Conti M*, Tu BT**, Khelidj N**, Losapio G, Polidori C. 2026
Unshared on the top: strong segregation of pathogen communities between western honey bees and wild bees in a retreating glacier foreland
Oikos, in press.
Steen B***, Morán-Ordóñez A***, Guisan A, Losapio G. 2026
Bioclimate overrules microtopography to inform on plant species’ sensitivity to climate change in proglacial ecosystems
Alpine Botany, in press.
Wan NF, ... Losapio G..., Scherber C GF. 2026
Plant diversity modifies multi-trophic interactions in croplands, grasslands and forests
Science Advances, 12, eaeb8680.
Simoncini A, Cantera I, ... Losapio G..., Ficetola GF. 2026
The global diversity and decline of glacier animals
PNAS, 123, 25, e2514455123.
Kambach S, Jandt U, ... Losapio G..., Bruelheide H. 2026
Habitat-specific trends in taxonomic, functional, and phylogenetic diversity in European plant communities over a century
Nature Communications, 17, 4208
Vujanović D, Knežević M, Đorđević A, Andrić A, Ranković Perišić M, Janković Milosavljević M, Veselić S, Losapio G, de Groot M, Vujić A, Radenković S. 2026
Larval diet breadth and wingspan mediate landscape–richness relationship in butterfly communities
Functional Ecology, 40, 624–635.
Conti M*, Dalpasso A, Nodari AM, Barzaghi B, Brambilla M, Cantera I, Ferrari A, Ficetola GF, Giachello S, Lo Parrino E, Messina V, Polidori C, Pozzi M, Redaelli S, Zerboni A, Losapio G, Falaschi M. 2026
Multitrophic effects of local management and landscape factors on taxonomic and functional diversity of managed grasslands
Agriculture, Ecosystems & Environment, 400, 110256, 1–13.
Balestra S*, Manuzi V*, Ceresa E*, Gatti P*, Pirttilahti RA*, Hodecek J***, Adatte T, Bomou B, Gabrieli P, Grand S, Losapio G. 2026
Long-term ecological impacts of Norway spruce plantations on biodiversity and microhabitat conditions
Ecosystems, 29, 34, 1–18.
Khelidj N**, Castilho A, Tu BN**, Charles C*, De Vere N, Grand S, Guisan A, Sanders I, Losapio G. 2026
Plant microhabitats mediate the effects of glacier retreat on soil microbial diversity and community assembly.
Applied Soil Ecology, 218, 1–13.
Losapio G, Mottet L*, Khelidj N**, Tu BN**, Cerabolini BEL, Grand S, de Vere N, Guisan A. 2025
Ecological indicators predict functional diversity dynamics following glacier retreat
Journal of Plant Ecology, 18, 6, 1–16.
Jiang F**, Fatichi S, Losapio G, Peleg N. 2025
Future glacier retreat and forest expansion in the Swiss Alps provide limited benefits for carbon sinks
Agricultural and Forest Meteorology, 372, 110682, 1– 14
Manuzi V*, Balestra S*, Gatti P*, Losapio G. 2025
Tree Monoculture Plantations Decrease Plant Diversity in the Italian Prealps
Ecological Solutions and Evidence, 6, e70067, 1-14.
[Data]
Whitley BS, Abermann J, Alsos IG, Biersma EM, Gårdman V, Høye6 TT, Jones L, Khelidj N**, Li Z, Losapio G, Pape T, Raundrup K, Schmitz P, Silva T, Wirta H, Roslin T, Ahlstrand NI, de Vere N. 2025
Harmonising digitised herbarium data to enhance biodiversity knowledge: major steps towards an updated checklist for the flora of Greenland
Plants, People, Planet, 1–15.
Losapio G, Lee JR, Fraser CI, Gillespie MAK, Kerr NR, Zawierucha K, Hamilton TL, Hotaling S, Kaufmann R, Kim O.-S., Klopsch C, Liu Y, Lukashanets D, Robinson SA, Brown LE. 2025
Impacts of deglaciation on biodiversity and ecosystem function
Nature Reviews Biodiversity, 1, 371-385.
Velasquez Casallas**, Khelidj N**, Moran-Ordoñez A***, Losapio G. 2025
Global impacts of glacier retreat on ecosystem services provided by soil and vegetation in mountain Regions: A literature review
Ecosystem Services, 73, 101730, 1-13.
[Data]
Charles C*, Khelidj N**, Mottet L*, Tu BN**, Adatte T, Bomou B, Faria M, Monbaron L, Reubi O, de Vere N, Grand S, Losapio G. 2025
The effects of glacier retreat on plant diversity and soil development.
Arctic, Antarctic, and Alpine Research, 57(1), 1-14.
Khelidj N**, Balestra S*, Caccianiga MS, Cerabolini BEL, Tampucci D, Losapio G. 2025
Plants' Contributions to People Shift With Glacier Extinction
Plant-Environment Interactions, 6, e70041, 1–14.
Cantera I, Giacchello S, Münkemüller T, Caccianiga M, Gobbi M, Losapio G, Marta S, Valle B, Zawierucha K, Thuiller W, Ficetola GF. 2025
Describing functional diversity of communities from environmental DNA
Trends in Ecology & Evolution, 40, 170–179.
Conti M*, Cerretti P, Ferrari A, Gabrieli P, Paone F, Polidori C, Sommaggio D, Losapio G. 2025
Glacier retreat decreases mutualistic network robustness over spacetime
Ecography, e07588, 1–11.
Alcántara JM, Verdú M, ... Losapio G, ... Zamora R. 2025
Key concepts and a world-wide look atplant recruitment networks
Biological Reviews, 100, 1127–1151.
Losapio G, Doussot B*, Araniti F, Bruno L, Guevara R, Dirzo R. 2024
Population persistence, phenotypic divergence and metabolic adaptation in yarrow (Achillea millefolium L.)
Ecosphere, 15, e70146.
Khelidj N**, Caccianiga M, Cerabolini BEL, Tampucci D, Losapio G. 2024
Glacier extinction homogenizes functional diversity via ecological succession.
Journal of Vegetation Science, 35, e13312.
Tu BN*, Khelidj N**, Cerretti P, de Vere N, Ferrari A, Paone F, Polidori C, Schmid J, Sommaggio D, Losapio G. 2024
Glacier retreat triggers changes in biodiversity and plant–pollinator interaction diversity.
Alpine Botany, 134, 171–182.
Losapio G, De Moraes CM, Dirzo R, Tscheulin T, Zuros N, Mescer MC. 2024
Habitat protection and removal of encroaching shrubs support the recovery of biodiversity and ecosystem functioning.
Conservation Science and Practice, 6, e13111.
Losapio G, De Moraes CM, Volker N, Tscheulin T, Zuros N, Mescher MC. 2024
The effects of shrub encroachment on arthropod communities depend on grazing history.
Global Ecology and Conservation, 50, e02819, 1–8.
Losapio G, Genes L, Knight CJ, McFadden TN, Pavan L. 2024
Monitoring and modelling the effects of ecosystem engineers on ecosystem functioning.
Functional Ecology, 38, 8-21.
[Plain language summary] [Blog post] [Press release]
Michalet R, Losapio G, Kikvidze Z, Brooker RW, Butterfield BJ, Callaway RM, Cavieres LA, Lortie CJ, Pugnaire FI, Schöb C. 2024
Assessing the accuracy of paired and random sampling for quantifying plant–plant interactions in natural communities.
Population Ecology, 66, 39-52.
Losapio G. 2023
Contextualizing the ecology of plant–plant interactions and constructive networks.
AoB Plants, 15, 1–9.
[Data]
Vujanović D, Losapio G, Mészáros M, Popov S, Ristić AM, Stojnić SM, Jović J, Vujić A. 2023
Forest and grassland habitats support pollinator diversity more than wildflowers and sunflower monoculture.
Ecological Entomology, 48, 421–432.
Verdú M, ..., Losapio G, ... Zamora R. 2023.
RecruitNet: A global database of plant recruitment networks.
Ecology, 10, e3923.
[Data]
Vujanovic D, Losapio G, Milic S, Milic D. 2022.
The impact of multiple species invasion on soil and plant communities increases with invasive species co-occurrence.
Frontiers in Plant Science, 13, 975824.
Genes L, Losapio G, Donatti CI, Guimarães PRJ, Dirzo R. 2022.
Frugivore population biomass, but not density, affects seed dispersal interactions in a hyper-diverse frugivory network.
Frontiers in Ecology and Evolution, 10, 794723.
Pugnaire FI, Losapio G, Schöb C. 2021.
Species interactions involving cushion plants in high-elevation environments under a changing climate.
Ecosistemas, 30, 2186.
Qu T, Du X, Peng Y, Guo W, Zhao C, Losapio G. 2021.
Invasive species allelopathy decreases plant growth and soil microbial activity.
PLoS ONE, 16, e0246685.
Losapio G, Schöb C, Staniczenko PPA, Carrara F, Palamara GM, De Moraes CM, Mescher MC, Brooker RW, Butterfield BJ, Callaway RM, Cavieres LA, Kikvidze Z, Lortie CJ, Michalet R, Pugnaire FI, Bascompte J. 2021.
Network motifs involving both competition and facilitation predict biodiversity in alpine plant communities.
PNAS, 118, e2005759118.
Losapio G, Norton Hasday E, Espadaler X, Germann C, Ortiz-Sánchez FJ, Pont A, Sommaggio D, Schöb C. 2021.
Facilitation and biodiversity jointly drive mutualistic networks.
Journal of Ecology, 109, 2029–2037.
Losapio G, Cerabolini BEL, Maffioletti C, Tampucci D, Gobbi M, Caccianiga M. 2021.
The consequences of glacier retreat are uneven between plant species.
Frontiers in Ecology and Evolution, 8, 616562.
[Data & R Code] [Blog post] [Infographic]
O'Brien MJ, Carbonell EP, Losapio G, Schlüter PM, Schöb C. 2021.
Foundation species promote local adaptation and fine-scale distribution of herbaceous plants.
Journal of Ecology, 109, 191–203.
Losapio G, Schmid B, Bascompte J, Michalet R, Cerretti P, Germann C, Haenni J-P, Neumeyer R, Ortiz J, Pont AC, Rousse P, Schmid J, Sommaggio D, Schöb C. 2021.
An experimental approach to assessing the impact of ecosystem engineers on biodiversity and ecosystem functions.
Ecology, 102, e03243. * On the cover *
Meyer JM, Leempoel K, Losapio G, Hadly EA. 2020.
Molecular Ecological Network Analysis: An effective conservation tool for the assessment of biodiversity, trophic interactions, and community structure.
Frontiers in Ecology and Evolution, 8, 360.
Pescador DS, Iriondo JM, Losapio G, Escudero A. 2020.
The assembly of plant–patch networks in Mediterranean alpine grasslands.
Journal of Plant Ecology, 14, 273–280.
Losapio G, Schöb C. 2020.
Pollination interactions reveal direct costs and indirect benefits of plant-plant facilitation for ecosystem engineers.
Journal of Plant Ecology, 13, 107–113. * On the cover *
Losapio G, Montesinos-Navarro A, Saiz H. 2019.
Perspectives for ecological networks in plant ecology.
Plant Ecology and Diversity, 12, 87–102.
O'Brien MJ, Tavares de Menezes LF, Bråthen KA, Losapio G, Pugnaire FI. 2019.
Facilitation mediates species presence beyond their environmental optimum.
Perspectives in Plant Ecology, Evolution and Systematics, 38, 24-30.
Losapio G, Fortuna MA, Bascompte J, Schmid B, Michalet R, Neumeyer R, Baur H, Castro L, Cerretti P, Germann C, Haenni J-P, Klopfstein S, Muller A, Ortiz J, Pont AC, Rousse P, Schmid J, Schwarz M, Sommaggio D, Schöb C. 2019.
Plant interactions shape pollination networks via nonadditive effects.
Ecology, 100, e02619.
Losapio G, De la Cruz M, Escudero A, Schmid B, Schöb C. 2018.
The assembly of a plant network in alpine vegetation.
Journal of Vegetation Science, 29, 999-1006.
Robinson SVJ, Losapio G, Henry GHR. 2018.
Flower-power: Flower diversity is a stronger predictor of network structure than insect diversity in an Arctic plant–pollinator network.
Ecological Complexity, 36, 1–6.
Losapio G, Pugnaire FI, O'Brien MJ, Schöb C. 2018.
Plant life history stage and nurse age change the development of ecological networks in an arid ecosystem.
Oikos, 128, 1390-1397.
Tian D, Jian L, Ma S, Fang W, Schmid B, Xu L, Zhu J, Li P, Losapio G, Jing X, Zheng C, Shen H, Zhu B, Fang J. 2017.
Effects of nitrogen deposition on soil microbial communities in temperate and subtropical forests in China.
Science of the Total Environment, 607-608, 1367-1375.
Losapio G, Schöb C. 2017.
Resistance of plant-plant networks to biodiversity loss and secondary extinctions following simulated environmental changes.
Functional Ecology, 31, 1145-1152.
[Data] [R code] [Lay summary]
Gobbi M, Ballarin F, Brambilla M, Compostella C, Isaia M, Losapio G, Maoletti C, Seppi R, Tampucci D, Caccianiga M. 2017.
Life in harsh environments: carabid and spider trait types and functional diversity on a debris-covered glacier and along its foreland.
Ecological Entomology, 42, 838-848.
Losapio G, Gobbi M, Marano G, Avesani D, Boracchi P, Compostella C, Pavesi M, Schöb C, Seppi R, Sommaggio D, Zanetti A, Caccianiga M. 2016.
Feedback effects between plant and insect communities along a primary succession gradient.
Arthropod-Plant Interactions, 10, 485-495.
Losapio G, Jordan F, Caccianiga M, Gobbi M. 2015.
Structure-dynamic relationship of plant-insect networks along a primary succession gradient on a glacier foreland.
Ecological Modelling, 314, 73-79.
* MSc student; ** PhD student
* Master student; ** PhD student; *** Postdoc
18. Manuzi V *, Balestra S *, Gatti P*, Losapio G. 2025. Data from: Tree Monoculture Plantations Decrease Plant Diversity in the Italian Prealps, Manuzi et al. 2025, Ecological Solutions and Evidence (v1.1). Zenodo. https://doi.org/10.5281/zenodo.15550910
17. Losapio G. 2024. Population persistence, phenotypic divergence and metabolic adaptation in yarrow (Achillea millefolium L.) along a climate gradient, CA, 1920 to 2023 ver 1.
Environmental Data Initiative https://doi.org/10.6073/pasta/88aad2d5c2677ed759d836f3f58564b4
16. Charles C *, Grand S, Losapio G. 2024. Data from Charles et al. 2024 ``Plant–Soil interactions drive the development of novel ecosystems after glacier retreat''.
Zenodo https://doi.org/10.5281/zenodo.10730935
15. Losapio G et al. 2021. Data from: Facilitation and biodiversity jointly drive mutualistic networks.
Dryad https://doi.org/10.5061/dryad.bzkh1897m
14. Losapio G et al. 2021. Data and R code for: An experimental approach to assessing the impact of ecosystem engineers on biodiversity and ecosystem functions.
Dryad https://doi.org/10.5061/dryad.dz08kprw0
13. Losapio G, Cerabolini BEL, Caccianiga M. 2020. Data of plant species distribution, traits, and soil in proglacial environments.
Stanford Digital Repository. https://purl.stanford.edu/hp416gs2665
12. O'Brien M et al. 2020. Data from: Foundation species promote local adaptation and fine-scale distribution of herbaceous plants.
Dryad https://doi.org/10.5061/dryad.xksn02vcs
11. Meyer J et al. 2020. Molecular ecological network analyses: An effective conservation tool for the assessment of biodiversity, trophic interactions, and community structure.
Dryad https://doi.org/10.5061/dryad.k6djh9w4q
10. Losapio G 2020. Data and R code from: Pollination interactions reveal direct costs and indirect benefits of plant-plant facilitation for ecosystem engineers.
Dryad https://doi.org/10.5061/dryad.g7m364t
9. Losapio G, Karlis G, Tscheulin T, Mescher M, De Moraes C. 2019. Plant–insect interactions data of ecological networks' response to land-use change and species invasion.
ETH Research Collection https://doi.org/10.3929/ethz-b-000318944
8. Losapio G, et al. 2019. Data from: Plant interactions shape pollination networks via nonadditive effects.
Dryad Digital Repository https://doi.org/10.5061/dryad.84rs783
7. Losapio G, Tscheulin T, De Moraes C, Mescher M. 2018. Diversity data of plant community response to land-use change and species invasion.
ETH Research Collection. https://doi.org/10.3929/ethz-b-000309353
6. Losapio G, Kawanstan I, Karlis G, Tscheulin T, De Moraes C, Mescher M. 2018. Biomass data of plant community response to land-use change and species invasion.
ETH Research Collection https://doi.org/10.3929/ethz-b-000311948
5. Losapio G, Nickels V, Tscheulin T, De Moraes C, Mescher M. 2018. Abundance and diversity data of arthropod community response to land-use change and species invasion.
ETH Research Collection https://doi.org/10.3929/ethz-b-000311956
4. Losapio G, Pulido H, Sims J, Mescher M, De Moraes C. 2018. Metabolomic data (GC-MS) of plants in overgrazed and invaded ecosystems.
ETH Research Collection https://doi.org/10.3929/ethz-b-000309354
3. Losapio G, De la Cruz M, Escudero A, Schmid B & Schob C. 2018. Data from: The assembly of a plant network in alpine vegetation.
ETH Research Collection https://doi.org/10.3929/ethz-b-000287871
2. Losapio G, Pugnaire FI, O'Brien MJ & Schob C. 2018. Data from: Plant life history stage and nurse age change the development of ecological networks in an arid ecosystem.
Dryad Digital Repository.https://doi.org/10.5061/dryad.db40gj8
1. Losapio G & Schöb C. 2017. Data from: Resistance of plant{plant networks to biodiversity loss and secondary extinctions following simulated environmental changes.
Dryad Digital Repository https://doi.org/10.5061/dryad.35q01
- Antonelli A, ..., Losapio G, ... 2026. State of the World’s Plants and Fungi 2026. Royal Botanic Gardens, Kew.
- Balestra S *, Losapio G. 2025. I ghiacciai si ritirano: impatti socio-ecologici. In Scienza Arte Tecnologia alleate per l'ambiente (ed. Streit-Bianchi M). DOI: 10.53136/97912218187893; Pagine: 29-35; Editore: Aracne.
- Losapio G. 2021. Creating your own environment: Ecosystems sustained by cushion plants. Bull Ecol Soc Am, 102, e01833.
- Losapio G. 2017. Plant interaction networks. Spatial dynamics, robustness and scaling up to pollinators. PhD Dissertation, University of Zurich. zora
- Losapio G. 2016. Plant interaction networks' response to environmental changes. Plant Science Center Newsletter, 29, 6.
