研究業績  

著書

1. 和田 哲, 2025. 繁殖生態にも戦略あり　ホンヤドカリ属の多様な生き方. 今すぐ見つけに行きたくなる ヤドカリ探索図鑑 第５章 ヤドカリを学ぶ, 緑書房, pp. 170-175.

2. 和田 哲, 2020. 種内変異と種内関係.  海岸動物の生態学入門〜ベントスの多様性に学ぶ 第３章, 海文堂, pp. 39-62.

3. 和田 哲, 2020. 個体群動態.  海岸動物の生態学入門〜ベントスの多様性に学ぶ 第６章, 海文堂, pp. 103-120.

4. 和田 哲, 2018. 動物が好きな人へ：行動生態学をヤドカリで紹介する.  海をまるごとサイエンス〜水産科学の世界へようこそ〜 第６章, 海文堂, pp. 66-76.

査読付き学術誌

1. Iyoda, T., Kido, Y., Yasuda, C. I., Wada, S. (accepted) Sexual differences in seasonal changes in major cheliped size and use pattern during dyadic interactions in the hermit crab Pagurus filholi. Plankton and Benthos Research.

2. Tomiyoshi, H. and S. Wada 2025. Effect of neighboring prey species on antipredator response of the marine gastropod. Plankton and Benthos Research 20: 205-209.

3. Nakayama, D., C. I. Yasuda and S. Wada 2025. Use of social information about novel food by juvenile solitary forktongue goby, Chaenogobius annularis. Ethology 2025: 0:e13569. 

4. Nagasaka, R., H. Yamada and S. Wada 2025. Fish yawn: interpopulation variation in yawning characteristics of juvenile white-spotted char Salvelinus leucomaenis. Journal of Ethology https://doi.org/10.1007/s10164-025-00844-w

5. Gushiken, H. and S. Wada 2024. “Standing on tiptoe”: A novel foraging technique in the starfish Patiria pectinifera. Plankton and Benthos Research 19: 84-87.

6. Yamada, H. and S. Wada 2024. Phenotype-dependent downstream dispersal under ordinary flow conditions in juvenile white-spotted char. Ecological Research 39: 330-340. 

7. Igarashi, K. and S. Wada 2024. Tonic immobility in a marine isopod: the effects of body size, sex, and colour morph. Journal of the Marine Biological Association of the United Kingdom 104: e17. 

8. Yamada, H. and S. Wada 2023. Interpopulation variation of behavioural and morphological traits that affect downstream displacement of the juvenile white-spotted charr Salvelinus leucomaenis. Journal of Fish Biology 102: 1168-1176.

9. Yamada, H. and S. Wada 2023. Fish yawn: the state-change hypothesis in juvenile white-spotted char Salvelinus leucomaenis. Journal of Ethology 41: 111-117.

10. 深澤藍子・和田 哲 2022. ヨツハモガニPugettia quadridens (De Haan, 1837) の北海道函館湾からの初記録 (First record of Pugettia quadridens (De Haan, 1837) from Hakodate Bay, Japan). Cancer 31: 49-52. 

11. Yamakami, R. and S. Wada 2022. Stomach contents of the deep-water whelks Buccinum kashimanum and Neptunea sp. (Caenogastropoda: Buccinidae) off the coast of Cape Esan, southwestern Hokkaido, Japan, in the western North Pacific Ocean. Venus 80: 40-45.

12.  Yamakami, R. and S. Wada 2021. Prey preferences of juvenile whelk Neptunea arthritica (Gastropoda: Buccinidae) in Hakodate Bay, Hokkaido, Japan. Plankton and Benthos Research 16: 257-265.

13. Yamada, H. and  S. Wada 2021. Morphological evolution reduces downstream displacement in juvenile landlocked salmon. Evolution 75: 1850-1861.

14. 津崎尚也・村上万龍・石原(安田)千晶・和田 哲 2021. 北海道南部に生息するヤドカリ2種における貝殻利用パターンの種間変異. Cancer 30: 35-48.

15. Yamakami, R. and S. Wada 2021. Prey utilization by Neptunea arthritica (Caenogastropoda: Buccinidae): predation on gastropods and size-related variation. Journal of Molluscan Studies 87: eyab016.

16. Kido, Y. and S. Wada 2020. Males display "inverse rapping" as a mating behavior to receptive females in the hermit crab Pagurus nigrofascia. Plankton & Benthos Research 15: 279-288.

17. Tamechika, M., K. Matsuno, S. Wada and Y. Yusa 2020. Different effects of mating group size as male and as female on sex allocation in a simultaneous hermaphrodite. Ecology and Evolution 10: 2492-2498.

18. 長谷川稜太・山田寛之・石原千晶・和田 哲, 2020. イワナの稚魚の個性に見られる生息地間変異. 魚類学雑誌 67: 11-24.

19. 山田寛之・榎本 尊・和田 哲, 2019. 北海道南部亀川に生息するイワナの稚魚における行動・形態の支流間比較. 魚類学雑誌 66: 221-225.

20. 木戸結菜・伊與田朋未・石原(安田)千晶・和田 哲, 2019. ヨモギホンヤドカリの性的対立：オスによる交尾前ガードの試みに対するメスの拒絶行動. Cancer 28: 7-15.

21. 小島早智・石原(安田)千晶・和田 哲, 2016. ヨモギホンヤドカリの交尾前ガードペアにみられる同類交配と闘争経験に基づいたオスの配偶者選択. Cancer 25: 17-24.

22. 加我夏美・石原(安田)千晶・和田 哲, 2016. ホンヤドカリPagurus filholiのオス間闘争：闘わないオスが多い. Cancer 25: 9-16.

23. Matsuo, K., D. Tanikawa, C.I. Yasuda and S. Wada, 2015. Sex-related differences in size, function and regeneration of the major cheliped in the hermit crab Pagurus filholi. Marine Ecology 36: 1391-1399. doi: 10.1111/maec.12238

24. Yasuda C. I. and S. Wada, 2015. Regeneration of major cheliped after the mating season in the hermit crab Pagurus middendorffii. Crustacean Research 44: 29-38.

25. Hasaba, Y., C. I. Yasuda and S. Wada, 2015. Reason for being single: some males do not guard receptive females in the hermit crab Pagurus filholi. Animal Behaviour 107: 1-5. doi:10.1016/j.anbehav.2015.06.003

26. Yasuda, C. I., K. Matsuo and S. Wada, 2015. Previous mating experience increases fighting success during male-male contests in the hermit crab Pagurus nigrofascia. Behavioral Ecology and Sociobiology 69: 1287-1292. doi: 10.1007/s00265-015-1941-x

27. Matsuo, K., C.I. Yasuda and S. Wada, 2014. Effect of cheliped loss due to autotomy on shell selection behavior in the hermit crab Pagurus middendorffii. Crustacean Research 43: 41-46.

28. Yasuda, C.I., K. Matsuo, Y. Hasaba and S. Wada, 2014. Hermit crab, Pagurus middendorffii, males avoid the escalation of contests with familiar winners. Animal Behaviour 96: 49-57.

29. Yasuda, C.I., K. Matsuo and S. Wada, 2014. Rapid regeneration of major cheliped related to its function in male-male contests in the hermit crab Pagurus middendorffii. Plankton and Benthos Research 9: 122-131.

30. 古屋野太一・伊藤 篤・和田 哲, 2013. マガキガイStrombus luhuanusの繁殖生態：漁獲圧の異なる２地域間の比較 (Reproductive ecology of the strawberry conch Strombus luhuanus: comparison between two sites differing in the degree of fishery exploitation). 日本ベントス学会誌 68: 1-6.

31. Ojima, H. and S. Wada, 2013. Contrasting anti-predatory responses in the intertidal periwinkle Littorina sitkana: effects of chemical cue, body size and time of day. Plankton and Benthos Research 8: 38-45.

32. Suzuki, Y., C. Yasuda, F. Takeshita and S. Wada, 2012. Male mate choice and male-male competition in the hermit crab Pagurus nigrofascia: importance of female quality. Marine Biology 159: 1991-1996. doi: 10.1007/s00227-012-1986-6.

33. 谷川大介・安田千晶・鈴木祐太郎・和田 哲, 2012. ホンヤドカリPagurus filholiのオス間闘争におけるオスの体サイズと配偶相手の質の影響 (Effects of male size and mate quality on male-male contest in the hermit crab Pagurus filholi). 日本ベントス学会誌 67: 15-19.

34. Yasuda, C., F. Takeshita and S. Wada, 2012. Assessment strategy in male-male contest of the hermit crab Pagurus middendorffii. Animal Behaviour 84: 385-390.

35. Takeshita F. and S. Wada, 2012. Morphological comparison of the second gnathopod in males of four caprellid species (Amphipoda: Caprellidae). Journal of Crustacean Biology 32: 673-676.

36. Yasuda, C., Y. Suzuki, and S. Wada,  2011. Function of the major cheliped in male-male competition of the hermit crab Pagurus nigrofascia. Marine Biology 158: 2327-2334. doi: 10.1007/s00227-011-1736-1

37. Takeshita, F., R. C Lombardo, S. Wada and Y. Henmi, 2011. Increased guarding duration reduces growth and offspring number in females of the skeleton shrimp Caprella penantis. Animal Behaviour 81: 661-666. doi: 10.1016/j.anbehav.2010.12.017

38. Wada, S., Y. Arashiro, F. Takeshita and Y. Shibata, 2011. Male mate choice in hermit crabs: prudence by inferior males and simple preference by superior males. Behavioral Ecology 22: 114-119. doi: 10.1093/beheco/arq183

39. Hasegawa, H., S. Wada, M. Aoki and K. Wada, 2011. Regional variation in preference for gastropod shell species in the hermit crab Pagurus filholi. Journal of the Marine Biological Association of the United Kingdom 91: 893-896. doi: 10.1017/S0025315410000561

40. 吉井健二・竹下文雄・和田 哲, 2009. ヨモギホンヤドカリにおける繁殖と成長: オスの成長にとって交尾前ガード行動はコストとなるか？ (Reproduction and growth in the hermit crab Pagurus nigrofascia: Do males incur costs for growth in precopulatory guarding?). 日本ベントス学会誌 64: 25-31.

41. Hasegawa, H., S. Wada, M. Aoki and K. Wada, 2009. Regional variation in shell utilization pattern of the hermit crab Pagurus filholi. Plankton and Benthos Research 42: 72-76. pdf

42. Wada, S., T. Oba, K. Nakata and A. Ito, 2008. Temporal allocation pattern between reproduction and growth within a breeding season of the hermit crab Pagurus nigrivittatus. Crustacean Research 37: 67-73.

43. Wada, S., T. Oba, K. Nakata and A. Ito, 2008. Factors affecting the interval between clutches in the hermit crab Pagurus nigrivittatus. Marine Biology 154: 501-507. doi: 10.1007/s00227-008-0944-9 abstract/full-text HUSCAP(free draft of this paper)

44. Oba, T., S. Wada and S. Goshima, 2008. Shell partitioning of two sympatric hermit crabs, Pagurus middendorffii and P. brachiomastus, in northeastern Hokkaido, Japan. Journal of the Marine Biological Association of the United Kingdom 88: 103-109. doi: 10.1017/S0025315408000015 abstract/full-text (free access)

45.  Wada, S., A. Ito and A. Mima, 2007. Evolutionary significance of prenuptial molting in female Pagurus hermit crabs. Marine Biology 152: 1263-1270. doi:10.1007/s00227-007-0773-2 abstract/full-textHUSCAP(free draft of this paper)

46. Goshima, S., S. Minouchi, K. Yoshino and S. Wada, 2006. Size assortative mating by the hermit crab Pagurus filholi (Decapoda: Anomura: Paguridae). Biology of Anomura II. Crustacean Research, Special Issue 6: 87-94.

47. Sakamoto, R., A. Ito and S. Wada, 2006. Combined effect of risk type and activity rhythm on anti-predator response of the shore crab Gaetice depressus (Crustacea: Grapsidae). Journal of the Marine Biological Association of the United Kingdom 86: 1401-1405. doi:10.1017/S0025315406014433 abstract/full-text HUSCAP(free draft of this paper)

48. Ito, A. and S. Wada, 2006. Intrasexual copulation and mate discrimination by Nodilittorina radiata (Gastropoda: Littorinidae). Journal of Ethology 24: 45-49. doi: 10.1007/s10164-005-0159-1 abstract/full-text

49. Ito, A. and S. Wada, 2005. Effect of food availability on reproduction and growth in the intertidal sea hare Aplysia juliana. Benthos Research 60: 11-16.

50. Wada, S., A. Mima and A. Ito, 2005. Reproductive phenology of sympatric hermit crabs in temperate Japan. Journal of the Marine Biological Association of the United Kingdom 85: 889-894. doi:10.1017/S0025315405011859 abstract/full-text

51. 伊藤 篤・古屋野太一・岡本侑樹・大野正夫・和田 哲, 2005. イセエビプエルルス幼生の着底場所選択 (Habitat preference in puerulus postlarvae of Japanese spiny lobster Panulirus japonicus). 水産増殖 53: 23-30.

52. Sato, T., M. Ashidate, S. Wada and S. Goshima, 2005. Effects of male mating frequency and male size on ejaculate size and reproductive success of female spiny king crab, Paralithodes brevipes. Marine Ecology Progress Series 296: 251-262. doi: 10.3354/meps296251 abstract/full-text

53. 樋口真理可・古屋野太一・伊藤 篤・和田 哲, 2004. マガキガイStrombus luhuanusの捕食者回避行動 (Predator avoidance of the conch snail Strombus luhuanus). 水産増殖 52: 251-258.

54. 古屋野太一・和田 哲, 2004. 土佐湾内の浦ノ内湾におけるマガキガイの密度に対する漁獲の影響 (Effect of fishery on density of the strawberry conch, Strombus luhuanus in Uranouchi Inlet, Tosa Bay, Japan). 水産増殖 52: 129-132.

55. Wada, S., and A. Mima, 2003. Reproductive characters of the hermit crab Pagurus proximus Komai 2000 in Hakodate Bay, southern Hokkaido, Japan. Crustacean Research 32: 73-78. full-text

56. Mima, A., S. Wada and S. Goshima, 2003. Antipredator defence of the hermit crab Pagurus filholi induced by predatory crabs. Oikos 102: 104-110. doi:10.1034/j.1600-0706.2003.12361.x abstract/full-text

57. Mima, A., H. Ohmori, S. Wada and S. Goshima, 2002. Exposure to predators affects shell preference of the hermit crab Pagurus filholi. Benthos Research 57: 85-89.

58. Kobayashi, T., S. Wada and H. Mukai, 2002. Extended maternal care for juveniles observed in Parallorchestes ochotensis Brandt, 1851 (Amphipoda: Gammaridae). Journal of Crustacean Biology 22: 135-142. abstract/full-text

59. Wada, S., 2001. Reproductive characters and population structure of the subtidal hermit crab Pagurus ochotensis. Benthos Research 56: 43-46.

60. Goshima, S., M. Kanazawa, K. Yoshino and S. Wada, 2000. Maturity in male stone crab Hapalogaster dentata and its application for fishery management. Journal of Crustacean Biology 20: 641-646. abstract/full-text

61. Wada, S., M. Ashidate, K. Yoshino, T. Sato and S. Goshima, 2000. Effects of sex ratio on spawning frequency and mating behavior of the spiny king crab, Paralithodes brevipes. Journal of Crustacean Biology 20: 479-482. abstract/full-text

62. Wada, S., H. Kitaoka, and S. Goshima, 2000. Reproduction of the hermit crab Pagurus lanuginosus and comparison of reproductive traits among sympatric hermit crabs. Journal of Crustacean Biology 20: 474-478. abstract/full-text

63. Goshima, S., M. Ikegawa, T. Sonoda, and S. Wada, 1999. Seasonal vertical migration within sediment in the brackish water clam Corbicula japonica. Benthos Research 54: 87-97.

64. Wada, S., 1999. Environmental factors affecting sexual size dimorphism in the hermit crab Pagurus middendorffii. Journal of the Marine Biological Association of the United Kingdom 79: 953-954. doi:10.1017/S0025315499001150 abstract/full-text

65. Wada, S., K. Tanaka, and S. Goshima, 1999. Precopulatory mate guarding in the hermit crab, Pagurus middendorffii (Decapoda: Paguridae): effects of population parameters on male guarding duration. Journal of Experimental Marine Biology and Ecology 239: 289-298. doi:10.1016/S0022-0981(99)00045-3 abstract/full-text

66. Wada, S., K. Ishizaki, H. Kitaoka, and S. Goshima, 1999. Shell utilization of the hermit crab Pagurus lanuginosus: sexual differences and specific comparisons. Benthos Research 54: 9-16.

67. Goshima, S., T. Kawashima, and S. Wada, 1998. Mate choice by males of the hermit crab Pagurus filholi: Do males assess ripeness and/or fecundity of females? Ecological Research 13: 151-161. abstract/full-text

68. Wada, S., M. Ashidate, and S. Goshima, 1997. Observations on the reproductive behavior of the spiny king crab Paralithodes brevipes (Anomura: Lithodidae). Crustacean Research 26: 56-61. full-text

69. Wada, S., H. Ohmori, S. Goshima, and S. Nakao, 1997. Shell-size preference of hermit crabs depends on their growth rate. Animal Behaviour 54: 1-8. doi:10.1006/anbe.1996.0319 abstract/full-text

70. Wada, S., T. Sonoda, and S. Goshima, 1996. Temporal size covariation of mating pairs of the hermit crab Pagurus middendorffii (Decapoda: Anomura: Paguridae) during a single breeding season. Crustacean Research 25: 158-164. full-text

71. Goshima, S., S. Wada, and H. Ohmori, 1996. Reproductive biology of the hermit crab Pagurus nigrofascia (Anomura: Paguridae). Crustacean Research 25: 86-92. full-text

72. Ohmori, H., S. Wada, S. Goshima, and S. Nakao, 1995. Effects of body size and shell availability on the shell utilization pattern of the hermit crab Pagurus filholi (Anomura: Paguridae). Crustacean Research 24: 85-92. full-text

73. Goshima, S., K. Ito, S. Wada, M. Shimizu, and S. Nakao, 1995. Reproductive biology of the stone crab Hapalogaster dentata (Anomura: Lithodidae). Crustacean Research 24: 8-18. full-text

74. Wada, S., S. Goshima, and S. Nakao, 1995. Reproductive biology of the hermit crab Pagurus middendorffii Brandt (Decapoda: Anomura: Paguridae). Crustacean Research 24: 23-32. full-text

その他

1. Yamakami, R. and S. Wada, 2021. Assemblages of mollusks (Gastropoda and Bivalvia) on the intertidal platform around Kattoshi, Hakodate Bay, Hokkaido. Bulletin of Fisheries Sciences, Hokkaido University 71: 1-14.

2. 石原（安田）千晶・和田 哲, 2018. テナガホンヤドカリに見られる繁殖期後の大鋏脚の再生. Cancer 27: 1-5.

3. 和田 哲・守田安祐美・石原（安田）千晶, 2018. 甲殻類はどのような情報を利用して行動を決定するのか？：ヤドカリの交尾前ガード行動を中心に. 海洋と生物 40: 106-113.

4. Morita, A. and S. Wada, 2018. Antenna length in the hermit crab Pagurus middendorffii: sexual dimorphism and effect on two types of male-male competition. Bulletin of Fisheries Sciences, Hokkaido University 68: 27-32.

5. 為近昌美・石原（安田）千晶・和田 哲, 2016.ミネフジツボの脱皮前後の行動. Cancer 25: 25-27.

6. 松尾謙人・石原（安田）千晶・和田 哲, 2015. 右鋏脚の自切によるテナガホンヤドカリの捕食者回避行動. Cancer 24: 21-23.

7. Wada, S., C.I. Yasuda, and C. McLay, 2014. Sexual size dimorphism in two endemic hermit crabs, Pagurus traversi and P. novizealandiae, in New Zealand. Bulletin of Fisheries Sciences, Hokkaido University 64: 31-35.

8. 阿部峻太・田代 慧・竹下文雄・和田 哲, 2012. 函館市近郊に生息するニホンザリガニにおける繁殖形質の個体群間比較. Bulletin of Fisheries Sciences, Hokkaido University 62: 29-41.

9. 松政正俊・倉田健悟・和田 哲・M. Grygier, 2010. 汽水域の生態プロセスを解く (1). 日本ベントス学会誌 65: 27.

10. 和田 哲, 2008. 函館近辺の海岸でみられるヤドカリたちの見分け方と繁殖生態. Oshimanography 15: 17-18.

11. 赤松克司・和田 哲, 2003. イセエビの摂餌行動に対する光条件の影響. Bulletin of Marine Sciences and Fisheries, Kochi University 22: 61-64.

12. 高橋真智子・樋口真理可・和田 哲, 2003. ヨロイイソギンチャクAnthopleura uchidaiとヒメイソギンチャクA. asiaticaにおける分布、繁殖、攻撃行動の比較. Bulletin of Marine Sciences and Fisheries, Kochi University 22: 53-60.

13. 和田 哲・伊藤 篤, 2000. アラレタマキビにおけるつま先立ち行動. Bulletin of Marine Sciences and Fisheries, Kochi University 20: 15-24.

14. 和田 哲, 2000. ヤドカリ類の生活史進化 (総説). Bulletin of Marine Sciences and Fisheries, Kochi University 20: 1-14.

15. Wada, S., 2000. Seasonal growth pattern and the effect of gastropod shells on sexual growth rates in the hermit crab Pagurus middendorffii. Bulletin of Faculty of Fisheries, Hokkaido University 51: 1-11.HUSCAP (free pdf)