Scientific papers and book chapters

2019 and in press

  • Deakin Z, Hamer K, Sherley RB, Bearhop S, Bodey TW, Clark B, Grecian WJ, Gummery M, Lane J, Morgan G, Morgan L, Phillips RA, Wakefield ED and Votier SC. In press. Sex differences in migration and demography of a wide-ranging seabird, the northern gannet. Marine Ecology Progress Series. DOI: 10.3354/meps12986. [LINK].
  • Gianuca D, Votier SC, Pardo D, Wood AG, Sherley RB, Ireland L, Choquet R, Pradel R, Townley S, Forcada J, Tuck GN and Phillips RA. In press. Sex-specific effects of fisheries and climate on the demography of sexually dimorphic seabirds. Journal of Animal Ecology. DOI: 10.1111/1365-2656.13009. [LINK].
  • Crawford RJM, Sydeman WJ, Thompson SA, Sherley RB and Makhado AB. 2019. Food habits of an endangered seabird indicate recent poor forage fish availability. ICES Journal of Marine Science. DOI: 10.1093/icesjms/fsz081. [LINK].
  • Campbell KJ, Steinfurth A, Underhill LG, Coetzee JC, Dyer BM, Ludynia K, Makhado AB, Merkle D, Rademan J, Upfold L and Sherley RB. 2019. Local forage fish abundance influences foraging effort and offspring condition in an Endangered marine predator. Journal of Applied Ecology 56: 1751–1760. [LINK].
  • Ropert-Coudert Y, Chiaradia A, Ainley D, Barbosa A, Boersma PD, Brasso R, Dewar M, Ellenberg U, Garcia Borboroglu P. Emmerson L, Hickcox R, Jenouvrier S, Kato A, McIntosh RR, Lewis P, Ramirez F, Ruoppolo V, Ryan PG, Seddon PJ, Sherley RB, Vanstreels RET, Waller L, Woehler EJ and Trathan PN. 2019. Happy Feet in a hostile world? The future of penguins depends on proactive management of current and predictable threats. Frontiers in Marine Science 6: 248. [PDF].
  • Provencher JF, Borrelle S, Sherley RB, Avery-Gomm S, Hodum P, Bond A, Major HL, McCoy K, Crawford R, Merkel F, Votier SC, Hatfield J, Reynolds M, Spatz D and Mallory ML. 2019. Seabirds. In: Sheppard CRC (Ed.). World Seas: An Environmental Evaluation. Volume III: Ecological Issues and Environmental Impacts. Academic Press, Cambridge, MA, USA. ISBN: 9780128050521. [LINK].


  • Gaglio D, Sherley RB, Cook TR, Ryan PG and Flower T. 2018. The costs of kleptoparasitism: a study of mixed-species seabird breeding colonies. Behavioral Ecology 29: 939–947. [LINK]. Featured on the Issue's Front Cover.
  • Gaglio D, Sherley RB, Ryan PG and Cook TR. 2018. A non-invasive approach to estimate the energetic requirements of an increasing seabird population in a perturbed marine ecosystem. Scientific Reports 8: 8343. [PDF].
  • Gaglio D, Cook TR, Sherley RB and Ryan PG. 2018. How many can you catch? Factors influencing the occurrence of multi-prey loading in provisioning Greater Crested Terns. Ostrich 89: 145–149. [LINK].
  • Payo-Payo A, Sanz-Aguilar A, Gaglio D, Sherley RB, Cook TR, Altwegg R and Ryan PG. 2018. Survival estimates for the greater crested tern Thalasseus bergii in southern Africa. African Journal of Marine Science 40: 43–50. [LINK]. Featured on the Issue's Front Cover.
  • Gaglio D, Cook TR, McInnes A, Sherley RB and Ryan PG. 2018. Foraging plasticity in seabirds: a non-invasive study of the diet of greater crested terns breeding in the Benguela Region. PLoS ONE 13: e0190444. [PDF].
  • Sherley RB, Barham BJ, Barham PJ, Campbell KJ, Crawford RJM, Grigg J, Horswill C, McInnes A, Morris TL, Pichegru L, Steinfurth A, Weller F, Winker H and Votier SC. 2018. Bayesian inference reveals positive but subtle effects of experimental fishery closures on marine predator demographics. Proceedings of the Royal Society B: Biological Sciences 285: 20172443. [PDF].


  • Brisson-Curadeau É, Bird D, Burke C, Fifield DA, Pace P, Sherley RB and Elliott KH. 2017. Seabird species vary in behavioural response to drone census. Scientific Reports 7: 17884. [PDF].
  • Crawford R, Ellenberg U, Frere E, Hagen C, Baird K, Brewin P, Crofts S, Glass J, Mattern T, Pompert J, Ross K, Kemper J, Ludynia K, Sherley RB, Steinfurth A, Suazo CG, Yorio P, Tamini L, Mangel JC, Bugoni L, Jimenez-Uzcategui G, Simeone A, Luna-Jorquera G, Gandini P, Woehler EJ, Pütz K, Dann P, Chiaradia A and Small C. 2017. Tangled and drowned: A global review of penguin bycatch in fisheries. Endangered Species Research 34: 373–396. [PDF].
  • Sherley RB. 2017. ​​​​​​​PhD jobs: Revamp funding structures. Nature 551: 440. [LINK].
  • Sydeman WJ, Thompson SA, Anker-Nilssen T, Arimitsu M, Bennison A, Bertrand S, Boersch-Supan P, Boyd C, Bransome N, Crawford RJM, Daunt F, Furness R, Gianuca D, Gladics A, Koehn L, Lang J, Logerwell E, Morris TL, Phillips EM, Provencher J, Punt AE, Saraux C, Shannon L, Sherley RB, Simeone A, Wanless RM, Wanless S and Zador S. 2017. Best Practices for Assessing Forage Fish Fisheries - Seabird Resource Competition. Fisheries Research 194: 209–221. [LINK].
  • Votier SC and Sherley RB. 2017. Quick guide Seabirds. Current Biology 27: R448–R450. [LINK].
  • Sherley RB, Botha P, Underhill LG, Ryan PG, van Zyl D, Cockcroft AC, Crawford RJM, Dyer BM and Cook TR. 2017. Defining ecologically relevant scales for spatial protection with long-term data on an endangered seabird and local prey availability. Conservation Biology 31: 1312–1321. [LINK]. Featured on the Issue's Front Cover.
  • Morten JM, Parsons NJ, Schwitzer C, Holderied MW and Sherley RB. 2017. Body condition as a quantitative tool to guide hand-rearing decisions in an endangered seabird. Animal Conservation 20: 471–479. [LINK].
  • Sherley RB, Ludynia K, Dyer BM, Lamont T, Makhado AB, Roux J-P, Scales KL, Underhill LG and Votier SC. 2017. Metapopulation tracking juvenile penguins reveals an ecosystem-wide ecological trap. Current Biology 27: 563–568. [PDF]
  • Gaglio D, Cook TR, Connan M, Ryan PG and Sherley RB. 2017. Dietary studies in birds: testing a non-invasive method using digital photography in seabirds. Methods in Ecology and Evolution 8: 214–222. [LINK].


  • Weller F, Sherley RB, Shannon LJ, Jarre A, Stewart T, Scott L, Altwegg R, Cecchini L-A, Crawford RJM, Geldenhuys D, Ludynia K and Waller LJ. 2016. Penguins’ perilous conservation status calls for complementary approach based on sound ecological principles: reply to Butterworth et al. (2015). Ecological Modelling 337: 1–3. [LINK]
  • Weller F, Sherley RB, Waller LJ, Ludynia K, Geldenhuys D, Shannon LJ, and Jarre A. 2016. System dynamics modelling of the Endangered African penguin populations on Dyer and Robben islands, South Africa. Ecological Modelling 327: 44–56. [LINK]
  • Sherley RB, Winker H, Altwegg R, van der Lingen CD, Votier SC and Crawford RJM. 2015. Bottom-up effects of a no-take zone on endangered penguin demographics. Biology Letters 11: 20150237. [LINK]
  • Gaglio D, Cook T and Sherley RB. 2015. Egg morphology of Swift Terns in South Africa. Ostrich 86: 287–289. [LINK]
  • Gaglio D, Sherley RB and Cook T. 2015. Insects in the diet of the Greater Crested Tern Thalasseus bergii bergii in Southern Africa. Marine Ornithology 43: 131–132. [PDF]
  • Sherley RB, Waller LJ, Strauss V, Geldenhuys D, Underhill LG and Parsons NJ. 2014. Hand-rearing, release and survival of African penguin chicks abandoned before independence by moulting parents. PLoS ONE 9(10): e110794. [PDF]
  • Sherley RB, Abadi F, Ludynia K, Barham BJ, Clark AE and Altwegg R. 2014. Age-specific survival and movement among major African Penguin Spheniscus demersus colonies. Ibis 156: 716–728. [LINK]
  • Ludynia K, Waller LJ, Sherley RB, Abadi F, Galada Y, Geldenhuys D, Crawford RJM, Shannon L and Jarre A. 2014. Processes influencing the population dynamics and conservation of African penguins at Dyer Island, South Africa. African Journal of Marine Science 36: 253–267. [LINK]
  • Weller F, Cecchini L-A, Shannon L, Sherley RB, Crawford RJM, Altwegg R, Scott L, Stewart T and Jarre A. 2014. A system dynamics approach to modelling multiple drivers of the African penguin population on Robben Island, South Africa. Ecological Modelling 277: 38–56. [LINK]
  • Sherley RB, Barham PJ, Barham BJ, Crawford RJM, Dyer BM, Leshoro TM, Makhado AB, Upfold L and Underhill LG. 2014. Growth and decline of a penguin colony and the influence on nesting density and reproductive success. Population Ecology 56: 119–128. [LINK]


  • Bonato M, Evans MR, Hasselquist D, Sherley RB, Cloete SWP and Cherry IC. 2013. Ostrich chick humoral immune responses and growth rate are predicted by parental immune responses and paternal colouration. Behavioral Ecology and Sociobiology 67: 1891–1901. [LINK]
  • Sherley RB, Ludynia K, Lamont T, Roux JP, Crawford RJM and Underhill LG. 2013. The initial journey of an endangered penguin: implications for seabird conservation. Endangered Species Research 21: 89–95. [LINK]
  • Sherley RB, Underhill LG, Barham BJ, Barham PJ, Coetzee JC, Crawford RJM, Dyer BM, Leshoro TM and Upfold L. 2013. Influence of local and regional prey availability on breeding performance of African penguins Spheniscus demersus. Marine Ecology Progress Series 473: 291–301. [LINK]
  • Sherley RB, Barham BJ, Barham PJ, Leshoro TM and Underhill LG. 2012. Artificial nests enhance the breeding productivity of African Penguins (Spheniscus demersus) on Robben Island, South Africa. Emu 112: 97–106. [LINK]
  • Sherley RB, Ludynia K, Underhill LG, Jones R and Kemper J. 2012. Storms and heat limit the nest success of Bank Cormorants: implications of future climate change for a surface-nesting seabird in southern Africa. Journal of Ornithology 153: 441–455. [LINK]
  • de Villiers MS, Mecenero S, Sherley RB, Heinze E, Leshoro TM, Merbold L, Nordt A, Parsons NJ, and Peter H-U. 2010. Introduced European rabbits (Oryctolagus cuniculus) and domestic cats (Felis catus) on Robben Island: Population trends and management recommendations. South African Journal of Wildlife Research 40: 139–148. [LINK]
  • Sherley RB, Burghardt T, Barham PJ, Campbell N and Cuthill IC. 2010. Spotting the difference: towards fully-automated population monitoring of African penguins Spheniscus demersus. Endangered Species Research 11: 101–111. [PDF]
  • Underhill LG, Sherley RB, Dyer BM and Crawford RJM. 2009. Interactions between snakes and seabirds on Robben, Schaapen and Meeuw Islands, Western Cape province, South Africa. Ostrich 80: 115–118. [LINK]


Popular press and other articles

  • Gaglio D and Sherley RB. 2017. Just snap it! Using digital cameras to discover what birds eat. Methods in Ecology and Evolution Blog ( Posted 5 January 2017. [LINK]
  • Sherley RB. 2016. Unusual foraging behaviour of two introduced mammals following degradation of their island habitat. Biodiversity Observations 7.29: 1–10. [PDF]
  • Gaglio D and Sherley RB. 2014. Nasty neighborhood: kleptoparasitism and egg predation of swift terns by Hartlaub's gulls. Ornithological Observations 5: 131–134. [PDF]
  • Sherley RB. 2014. Adult survival declines as African Penguin population plummets. British Ornithologists' Union Blog. Posted 25 August 2014. [LINK]
  • Rueda E and Sherley RB. 2013. First observations of Black-winged Stilt Himantopus himantopus on Robben Island. Ornithological Observations 4: 24–26. [PDF]
  • Schwitzer C, Simpson N, Roestorf M and Sherley RB. 2013. The African Penguin Chick Bolstering Project: A One Plan approach to integrated species conservation. WAZA Magazine 14: 23–26.
  • Sherley RB and Robinson KJ. 2012. First observation: Pied Avocet at Robben Island. Ornithological Observations 3: 100–101. [PDF]
  • Sherley RB. 2012. Catching up on the secret lives of juvenile penguins. Environment 10: 64–66.
  • Sherley RB, Dyer BM, Underhill LG and Leshoro TM. 2011. Birds occurring or breeding at Robben Island, South Africa, since 2000. Ornithological Observations 2: 69–100. [PDF]
  • Sherley RB. 2011. Research highlights: Revealing the Secret Lives of African Penguins. Environment 7: 14–16.


PhD thesis

  • Sherley RB. 2010. Factors influencing the demography of Endangered seabirds at Robben Island, South Africa: Implications and approaches for management and conservation. PhD Thesis, University of Bristol, U.K. [PDF] (31.2 MB)


Please feel free to contact me for reprints of any of my

papers not available for download here.