Ground Tissues

Simple Tissues

  • Ground tissues, sometimes called simple tissues, since they are composed of only 1 cell type

  • Ground tissues make up the bulk mass and strengthening elements of the plant body

  • There are three main types of ground tissues in plants:

Parenchyma

  • Parenchyma serves as the bulk material and storage area for plants

Collenchyma

  • Collenchyma is a tissue used for flexible support in stems or petioles

Sclerenchyma

  • Sclerenchyma is a tissue used for rigid support in stems or petioles

Plant tissue shapes

Above: Shapes and relative sizes of plant tissues, including wall thickenings

Parenchyma

  • From para meaning "beside", and chyma meaning "in-filling", referring to the nature of parenchyma to fill-in around the other tissues

Function

  • Parenchyma serves as the main storage tissue for plants

  • It also serves as the bulk structural tissue of roots, stems, and leaves

Features

  • These cells are usually thin-walled, usually lacking secondary walls

    • In some cases of stress, plants can add lignin to parenchyma, strengthening those areas (called lignified parenychma)

  • They are isodiametric in shape; they are an equal diameter in all directions

  • Parenchyma cells are alive at maturity conducting cellular processes, unlike xylem and sclerenchyma

  • Parenchyma is the most abundant cell type in a plant

Above [x400]: Magnified parenchyma cells, with a couple cells showing a pinkish nucleus. Note the large size, thin wall, and nearly circular shape

Parenchyma sub-types

Aerenchyma

  • Parenchyma tissues with abundant air spaces are called aerenchyma

  • The spongy mesophyll of a leaf, where gas exchange occurs near stomata, is a common site of this tissue

  • Many aquatic plants possess aerenchyma in stems to shuttle oxygen to roots for respiration in anoxic, water-logged soils

Chlorenchyma

  • Parenchyma tissues with many chloroplasts are called chlorenchyma

  • The palisade layer of a leaf, where light interception occurs for photosynthesis, is a common site of this tissue

  • Many herbaceous stems also exhibit chlorenchyma to contribute to the photosynthesis of the plant

Collenchyma

  • From kolle meaning "glue", and chyma meaning "in-filling", referring to the flexible but supportive nature of collenchyma

Function

  • Collenchyma provides flexible support in stems or petioles

Features

  • Collenchyma has an irregularly-thickened primary wall

  • These cells are found under the epidermis (i.e. centrifugal direction), in herbaceous stems and some leaves

  • Many plants in the Lamiaceae, with square stems, use collenchyma for stem support

  • Collenchyma cells are alive at maturity conducting cellular processes, unlike xylem and sclerenchyma

Above [x400]: Magnified collenchyma cells, just under the epidermis. Note the irregular-thickening of the cell wall

Sclerenchyma

  • From skleros meaning "hard", and chyma meaning "in-filling", referring to the rigid or hardened nature of sclerenchyma

Function

  • Sclerenchyma fibers provide rigid support in stems

  • Sclerenchyma sclereids provide hardened protection in fruit pits and leaves

Features

  • Cells have evenly-thickened secondary walls

  • These cells have a very small lumen (interior space), compared to xylem cells

  • Sclerenchyma cells are dead at maturity

  • Many herbaceous stems use sclerenchyma tissues to strengthen weak areas, such as around primary phloem

  • The strength of bamboo, comes purely from sclerenchyma, and not from wood!

Above [1,000x]: Magnified sclerenchyma cells, called fibers. Note the evenly-thickened wall, small lumen, and small size

Sclerenchyma sub-types

Fibers

  • Fibers are long and thin in shape

  • They are evenly-thickened in cross-section with a small or no lumen

  • Fibers are found longitudinally in stems, roots, and leaves

Sclereids

  • Sclereids are roughly the same length and width

  • These cells are found in "pockets" of fruits, seeds, stems, and leaves

  • Sclereids create strength and/or a deterrent to herbivores that may damage embryos, stems, and leaves while feeding