Ground Tissues
Simple Tissues
Ground tissues, sometimes called simple tissues, since they are composed of only 1 cell type
Ground tissues make up the bulk mass and strengthening elements of the plant body
There are three main types of ground tissues in plants:
Parenchyma
Parenchyma serves as the bulk material and storage area for plants
Collenchyma
Collenchyma is a tissue used for flexible support in stems or petioles
Sclerenchyma
Sclerenchyma is a tissue used for rigid support in stems or petioles
Above: Shapes and relative sizes of plant tissues, including wall thickenings
Parenchyma
From para meaning "beside", and chyma meaning "in-filling", referring to the nature of parenchyma to fill-in around the other tissues
Function
Parenchyma serves as the main storage tissue for plants
It also serves as the bulk structural tissue of roots, stems, and leaves
Features
These cells are usually thin-walled, usually lacking secondary walls
In some cases of stress, plants can add lignin to parenchyma, strengthening those areas (called lignified parenychma)
They are isodiametric in shape; they are an equal diameter in all directions
Parenchyma cells are alive at maturity conducting cellular processes, unlike xylem and sclerenchyma
Parenchyma is the most abundant cell type in a plant
Above [x400]: Magnified parenchyma cells, with a couple cells showing a pinkish nucleus. Note the large size, thin wall, and nearly circular shape
Parenchyma sub-types
Aerenchyma
Parenchyma tissues with abundant air spaces are called aerenchyma
The spongy mesophyll of a leaf, where gas exchange occurs near stomata, is a common site of this tissue
Many aquatic plants possess aerenchyma in stems to shuttle oxygen to roots for respiration in anoxic, water-logged soils
Chlorenchyma
Parenchyma tissues with many chloroplasts are called chlorenchyma
The palisade layer of a leaf, where light interception occurs for photosynthesis, is a common site of this tissue
Many herbaceous stems also exhibit chlorenchyma to contribute to the photosynthesis of the plant
Collenchyma
From kolle meaning "glue", and chyma meaning "in-filling", referring to the flexible but supportive nature of collenchyma
Function
Collenchyma provides flexible support in stems or petioles
Features
Collenchyma has an irregularly-thickened primary wall
These cells are found under the epidermis (i.e. centrifugal direction), in herbaceous stems and some leaves
Many plants in the Lamiaceae, with square stems, use collenchyma for stem support
Collenchyma cells are alive at maturity conducting cellular processes, unlike xylem and sclerenchyma
Above [x400]: Magnified collenchyma cells, just under the epidermis. Note the irregular-thickening of the cell wall
Sclerenchyma
From skleros meaning "hard", and chyma meaning "in-filling", referring to the rigid or hardened nature of sclerenchyma
Function
Sclerenchyma fibers provide rigid support in stems
Sclerenchyma sclereids provide hardened protection in fruit pits and leaves
Features
Cells have evenly-thickened secondary walls
These cells have a very small lumen (interior space), compared to xylem cells
Sclerenchyma cells are dead at maturity
Many herbaceous stems use sclerenchyma tissues to strengthen weak areas, such as around primary phloem
The strength of bamboo, comes purely from sclerenchyma, and not from wood!
Above [1,000x]: Magnified sclerenchyma cells, called fibers. Note the evenly-thickened wall, small lumen, and small size
Sclerenchyma sub-types
Fibers
Fibers are long and thin in shape
They are evenly-thickened in cross-section with a small or no lumen
Fibers are found longitudinally in stems, roots, and leaves
Sclereids
Sclereids are roughly the same length and width
These cells are found in "pockets" of fruits, seeds, stems, and leaves
Sclereids create strength and/or a deterrent to herbivores that may damage embryos, stems, and leaves while feeding