Selected Publications

• Gicaraya, V.A.G. & Española, C. P. (2026). Field observations and citizen science reveal ecological insights into rare and threatened parrots in the Philippines. Journal of Threatened Taxa 18(1): 28174–28185. https://doi.org/10.11609/jott.9812.18.1.28174-28185 

• Gicaraya, V. A. G., Española, C. P., Quijano, E. S., Soriano, K., Torio, E. S., & Naguit, D. R. D. (2025). Cavity characteristics explain the differences in realized nest niches among tree cavity-nesting birds in a lowland tropical forest in Luzon Island, Philippines. Annales Zoologici Fennici, 62(1), 51–64. https://doi.org/10.5735/086.062.0105

• Angeles,E.J.A., & Española, C.P. (2024). Earthworm Species Diversity across an Elevational Gradient in Mt. Sawtooth and its Neighboring Highlands in the Zambales Mountain Range, the Philippines. Philippine Journal of Science 153 No. 6A: 2003-2018. 

• Gicaraya, V. A. G., & Española, C. P. (2024). Breeding Biology of the Luzon Hornbill Penelopides manillae manillae (Aves: Bucerotidae) on Luzon Island, Philippines. Ornithological Science: 23 (2), 91-101. https://doi.org/10.2326/osj.23.91

• Gicaraya, V. A. G., & Española, C. P. (2024). Observations on the breeding biology of the Luzon flameback Chrysocolaptes haematribon (Wagler 1827) (Piciformes: Picidae) with notes on active nest defence against dollarbird Eurystomus orientalis (Linnaeus 1766) (Coraciiformes: Coraciidae). Natural History Sciences. https://doi.org/10.4081/nhs.2024.748

• Gicaraya, V. A. G., and Española, C. P. (2024). First description of the breeding biology and behaviour of the near threatened northern sooty woodpecker (Mulleripicus funebris) (Valenciennes 1826) (Piciformes: Picidae) in Luzon Island, Philippines. Natural History Sciences, 11(1). https://doi.org/10.4081/nhs.2024.701

• Gicaraya, V. A. G. and C. P. Española. 2023. Notes on the breeding biology and nesting behaviour of the Coleto Sarcops calvus in the wild. Journal of Asian Ornithology. 39: 26-29.

• Gicaraya, V. A. G. and C. P. Española. 2023. Brood parasitism of the Asian Koel (Eudynamys scolopacea) on the tree-cavity nesting Coleto (Sarcops calvus): first record in the Philippines. Ornithological Science 22 (2)

• Gan, J. L., A. M. Tiongco, T. A. Cervero, L. G. Ilustrisimo, S. R. Matipo, E. G. Doca and C. P. Española. 2023. Spring migration of diurnal raptors in Northern Luzon, Philippines along the oceanic flyway in 2015 and 2016. Journal of Raptor Research. https://doi.org/10.3356/JRR-22-63

• Li, Y., Xiao, J., Jiang, Y., Wright, S.J, Mayfield, M.M., Godoy, O., Alonso, A., Anderson-Teixeira, K.J., ..., Rodriguez, L.J.V., et al. 2026. Higher-order interactions enhance the latitudinal tree diversity gradient. Nature 653, 433–438. https://doi.org/10.1038/s41586-026-10434-6

• Phillips, O., Duque, A., Rodriguez, L., Zo-Bi, Irié Casimir. 2025. The forest space age needs eyes on the ground. Nature 641, 1101. doi: https://doi.org/10.1038/d41586-025-01643-6 (Correspondence)

• Palmieri, L., Rodriguez, L.J., vanNoort, S., Cruaud, A., Pereira, R.A.S. & Rasplus, J.-Y. 2026. From figs to acorns: Early specialization and host shifts shaped the global radiation of Curculionini weevils (Coleoptera: Curculionidae). Systematic Entomology 51(2), e70050. https://doi.org/10.1111/syen.70050

• Bascos, E.M.A., Fernando, E.S., Duya, M.V., Rodriguez, L.J.V. 2024. What's that smell? The Putrid Scent of Rafflesia consueloae, its Origin and Developmental Regulation. Flora 318: 152571. https://doi.org/10.1016/j.flora.2024.152571

• Latorre, J.E., Galindon, J.M.M., Bartolome, N.A., Duya, M.V., Rodriguez, L.J.V. 2024. Demography, Structure, and Composition of a Low-Disturbance Forest in Luzon, Philippines. Environment and Natural Resources Journal 22 (3): 270-282. DOI: 10.32526/ennrj/22/20230235

• Cancio, E.J.L., Rodriguez, L.J.V. 2024. Diel and vertical niche differentiation of frugivore communities feeding on Ficus nota (Blanco) Merr. Philippine Journal of Science 153(2): 639–648

• Bascos EMA, Fernando ES, Duya MV, Rodriguez LJV.  2021. Beginnings of a plant parasite: early development of Rafflesia consueloae inside its Tetrastigma host. Planta 254:61. https://doi.org/10.1007/s00425-021-03710-4

• Rodriguez LJV, Cruaud A, Rasplus J-Y. 2021. Low Sampling Effort and High Genetic Isolation Contribute to Underdocumented Diversity in Philippine Fig Wasps. Philippine Journal of Science 150 (S1): 173-180. https://philjournalsci.dost.gov.ph/publication/special-issues/biodiversity/104-vol-150-s1/1342-low-sampling-effort-and-high-genetic-isolation-contribute-to-underdocumented-diversity-in-philippine-fig-wasps

• Gigante ET, Lim EJ, Crisostomo KG, Cornejo P, Rodriguez LJ. 2021. Increase in humidity widens heat tolerance range of tropical Ceratosolen fig wasps. Ecological Entomology 46: 573-581. https://doi.org/10.1111/een.13003

• Rodriguez LJ, Bain A, Chou LS, Conchou L, Cruaud A, Gonzales R, Hossaert-McKey M, Rasplus JY, Tzeng HY, Kjellberg F. 2017. Diversification and spatial structuring in the mutualism between Ficus septica and its pollinating wasps in insular South East Asia. BMC Evolutionary Biology 17:207. https://doi.org/10.1186/s12862-017-1034-8

(Complete list of publications can be found on Google Scholar profile)

• Gan, J. L., Meyer, K. M., Pfeifer, M., Soroño, T. C., Torio, E. S., van Weerd, M., Gatan-Balbas, M. & Shirley, M. D. F. (2026). Retaining fruiting trees in the agricultural matrix is important for bird-mediated seed dispersal in a fragmented forest landscape. Ecological Modelling, 520, 111701. https://doi.org/10.1016/j.ecolmodel.2026.111701

• Morrison, K., Pottier, P., Pollo, P., Ricolfi, L., ...Gan, J. L….. & Nakagawa, S. (2025). MATES: A tool for appraising the completeness with which a meta-analysis has been reported. Environment International, 109935. https://doi.org/10.1016/j.envint.2025.109935

• Gan, J. L., Pfeifer, M., Soroño, T. C., Gicaraya, V. A., Torio, E. S., van Weerd, M., Gatan-Balbas, M., Shirley, M. D. F. (2025) Effect of local and landscape variables on bird communities in forest-agricultural mosaics in Northeast Luzon, Philippines. Biotropica, 57(5), e70076. https://doi.org/10.1111/btp.70076

• Gan, J. L., Grainger, M. J., Shirley, M. D. F., Davis, S., Watson, M., Dube, S., & Pfeifer, M. (2025). Effectiveness of perches in promoting bird-mediated seed dispersal for natural forest regeneration: a systematic review. Environmental Evidence, 14(1), 10. https://doi.org/10.1186/s13750-025-00363-8

• Gomez, S., English, H. M., Bejarano Alegre, V., Blackwell, P. G., Bracken, A. M., Bray, E., Evans, L. C., Gan, J. L.... & Börger, L. (2025). Understanding and predicting animal movements and distributions in the Anthropocene. Journal of Animal Ecology, 94(6), 1146-1164. https://doi.org/10.1111/1365-2656.70040

• Castro, J. C., Gan, J., & Longnecker, N. (2025). Participatory Exhibitions as An Awareness-raising Platform for Bird Window Collisions. CSID journal of infrastructure development, 8(1), 90-106. https://doi.org/10.7454/jid.v8.i1.1179

• Gan, J. L., Grainger, M. J., Shirley, M. D. F., & Pfeifer, M. (2023). How effective are perches in promoting bird-mediated seed dispersal for natural forest regeneration? A systematic review protocol. Environmental Evidence, 12(1), 1-11. https://doi.org/10.1186/s13750-023-00308-z

• Diaz, S. D. U., Gan, J. L., & Tapang, G. A. (2023). Acoustic indices as proxies for bird species richness in an urban green space in Metro Manila. PLoS ONE, 18(7), e0289001. https://doi.org/10.1371/journal.pone.0289001

• Gan, J. L., Tiongco, A. M., Cervero, M. T. A., Ilustrisimo, L. G., Matipo, S. R., Doca, E. G., & Española, C. P. (2023). Spring Migration of Diurnal Raptors in Northern Luzon, Philippines, along the Oceanic Flyway in 2015 and 2016. Journal of Raptor Research, 57(3), 1-8. https://doi.org/10.3356/JRR-22-63

(Complete list of publications can be found on Google Scholar profile)

• Batomalaque, G.A., Que, G.L.Q., Itong, T.A.B., Masanga, A.R.L., de Chavez, E.R.C. and Fontanilla, I.K.C. 2019. Cytochrome C Oxidase Subunit 1 (COI) Profile of the Philippine Helicostylinae (Gastropoda: Stylommatophora: Camaenidae). Philippine Journal of Science 148(S1): 1-13.

• Batomalaque, G.A. and Rosenberg, G. 2018. Three new species of Zaptyx (Gastropoda: Eupulmonata: Clausiliidae) from the Babuyan and Batanes Islands, Philippines. Archiv für Molluskenkunde 147 (1): 77–86.

• Ramos, D.A.E., Batomalaque, G.A. and Anticamara, J.A. 2018. Current status of Philippine Mollusk museum collections and research, and their implications on biodiversity science and conservation. Philippine Journal of Science 147(1):123-163.

• Batomalaque, G.A. 2016. Philippine helicostyline land snail diversity, distribution and areas of endemism: an effect of forest type? (COA Academic Grant Brief Report). American Conchologist 46(2): 36-40.

• De Chavez, E.R.C., Fontanilla, I.K.C., Batomalaque, G.A., and Chiba, S. 2015. A new Helicostyla species (Bradybaenidae: Helicostylinae) from Patnanungan Island, Philippines. Asia Life Sciences 24(1):39-47.

• Sosa, B.O. III, Batomalaque, G.A., and Fontanilla, I.K.C. 2014. An updated survey and biodiversity assessment of the terrestrial snail (Mollusca: Gastropoda) species in Marinduque, Philippines. Philippine Journal of Science 143(2):199-210.

• Pedales, R.D.C. and Batomalaque, G.A. 2014. An account of the accessioned collections of the UP Biology-Invertebrate Museum. Science Diliman 26(2):40-48.

• Batomalaque, G.A., Sales, P.R.L. and Fontanilla, I.K.C. 2013. DNA barcoding using cytochrome oxidase I (CO1) of pulmonate gastropods from Batan Island, Batanes, Philippines. Asia Life Sciences 22(2):341-357.

• Batomalaque, G.A., Arce, B.G.P., Hernandez, M.B.M. and Fontanilla, I.K.C. 2010. Survey and spatial distribution of shoreline malacofauna in Grande Island, Subic Bay. Philippine Journal of Science 139(1):149-159.

• Molina, J., Wicaksono, A., Michael, T.P., Kwak, S.H., Pedales, R.D., Joly‐Lopez, Z., Petrus, S., Mamerto, A., Tomek, B., Ahmed, S. and Maddu, V., 2023. The seed transcriptome of Rafflesia reveals horizontal gene transfer and convergent evolution: Implications for conserving the world's largest flower. Plants, People, Planet. 

• Manalili, M.D., Pedales, R., Dizon, E.M. and Fontanilla, I.K., 2022. Detection of bloodworm larvae (Diptera: Chironomidae) in the golden apple snail Pomacea canaliculata (Lamarck, 1819)(Gastropoda: Ampullariidae) in Metro Manila. Philippine Journal of Health Research and Development, 26(1), pp.1-7. 

• Molina, J., Nikolic, D., Jeevarathanam, J.R., Abzalimov, R., Park, E.J., Pedales, R., Mojica, E.R.E., Tandang, D., McLaughlin, W., Wallick, K. and Adams, J., 2022. Living with a giant, flowering parasite: metabolic differences between Tetrastigma loheri Gagnep.(Vitaceae) shoots uninfected and infected with Rafflesia (Rafflesiaceae) and potential applications for propagation. Planta, 255, pp.1-12. 

• Pedales, R.D. and Fontanilla, I.K.C., 2018. Forensic entomology in the Philippines: Establishing Baseline Data on the Forensically Important Blow Fly Species Chrysomya megacephala (Fabricius, 1794). Philippine Journal of Science, 147(1), pp.17-25.

• Molina, J., McLaughlin, W., Wallick, K., Pedales, R., Marius, V.M., Tandang, D.N., Damatac, A., Stuhr, N., Pell, S.K., Lim, T.M. and Novy, A., 2017. Ex Situ Propagation of Philippine Rafflesia in the United States: Challenges and Prospects. Sibbaldia: the Journal of Botanic Garden Horticulture, (15), pp.77-96.

• Pedales, R.D., Damatac, A.M., Limbo, C.A., Marquez, C.M.D., Navarro, A.I.B. and Molina, J., 2016. DNA Barcoding of Philippine Herbal Medicinal Products. Journal of AOAC International, 99(6), pp.1479-1489.

• Pedales, R.D. and Batomalaque, G.A., 2014. An Account of the Accessioned Collections of the UP Biology Invertebrate Museum. Science Diliman, 26(2).