Modeling of ecological networks based on these evolutionary faunas suggest that increasing Phanerozoic diversity and ecological complexity, and increasing intensity of biotic interactions result in increasing network complexity. However, it has been suggested that early Paleozoic community structure may be remarkably similar to that of modern communities, and that trophic organization may not have undergone any significant changes since the Cambrian.
This project aims to test whether Mesozoic escalatory trends affected ecosystem structure and function, using marine food webs from the Jurassic and Cretaceous. Differences in community dynamics among these ecosystems will be quantified using food web structure, models of community stability dynamics, and a mathematical model of secondary extinction that accounts for the uncertainty associated with paleontological data.
Food webs are being reconstructed using the Paleobiology Database and published literature, and numerous museum collections. Trophic interactions are inferred from relevant autecological literature, functional morphology, habitat, species associations, or living analog species. Changes in ecosystem dynamics will be measured as community stability after minor perturbation, and resistance to the propagation of secondary extinctions in stochastically generated species level food web networks.