Members and research topics (2026) / メンバーとその研究内容（2026年度）
Faculty member / 教 員
● Shinya Yamamoto（Prof.) yamamoto.shinya.3u [at] kyoto-u.ac.jp
Comparative cognitive & behavioral science on the evolution of sociality with various viewpoints from physiology, psychology, behavioral ecology, etc.
Keywords: social intelligence, group-mindedness, empathy, prosociality, cooperation, culture, social network, drone
Animal species: chimpanzees, bonobos, horses, dogs, etc (both in the wild and captivity)
山本真也（教授）yamamoto.shinya.3u [at] kyoto-u.ac.jp
ヒトを含めた動物の社会性の進化。専門は比較認知行動学。ホルモン・認知・行動・社会・生態など幅広く扱っています。
キーワード：社会的知性、集団性、おもいやり、共感、協力、文化、社会ネットワーク、ドローン
主な対象種：チンパンジー・ボノボ・ウマ・イヌ・ヒト
● Saho Takagi (Hakubi Assistant Prof.)
高木佐保（白眉特定助教）
● Nachiketha Sharma (Hakubi Assitant Prof. from July）Elephant cognition and behavior in the wild and captivity. nachisrnachi [at] gmail.com
ナチケータ・シャルマ（白眉特定助教）ゾウの認知・行動研究：フィールド研究と飼育下実験. nachisrnachi [at] gmail.com
● Hoi-Lam Jim (Program-Specific Assistant Professor) Social cognition in Asian elephants. jim.hoilam.6k [at] kyoto-u.ac.jp
ジム・ホイラム（特定助教 ）アジアゾウの社会的認知. jim.hoilam.6k [at] kyoto-u.ac.jp
Secretary / 秘書
● Satomi Kondo 近藤智美
Postdocs / 研究員
● Miki Kamatani (JSPS research fellow PD) Social bonding in horses. mm.kamatani [at] gmail.com
鎌谷美希 (日本学術振興会 特別研究員PD)ウマにおける社会的絆の形成. mm.kamatani [at] gmail.com
● Kumi Shinoda (JSPS research fellow PD) Morphological, perceptual, and cognitive changes during the process of dog de-domestication. shinophilax93 [at] gmail.com
篠田公美（日本学術振興会 特別研究員PD）イヌの脱家畜化プロセスにおける形態・知覚・認知的変化の解明. shinophilax93 [at] gmail.com
● Piao Yige (Spoon, 朴弋戈) (JSPS International Research Fellow) Social learning process and cultural transmission in chimpanzees and orangutans. spoonasf [at] gmail.com
ピャオ・イグェ（学振外国人特別研究員）チンパンジーとオランウータンにおける社会的学習プロセスと文化伝達. spoonasf [at] gmail.com
● Asuka Tosaka (Researcher)
戸坂明日香 (研究員）
● Madoka Hattori (Researcher) Bonding mechanisms between cats and humans. hattori.madoka.46p [at] st.kyoto-u.ac.jp
服部円（研究員）ネコとヒトとの絆形成メカニズムの解明. hattori.madoka.46p [at] st.kyoto-u.ac.jp
Technical staff / 技術スタッフ
● Yoshiko Yamane 山根美子
Graduate students / 大学院生
● Lin Chaoyu (林昭妤) (D3) Vocal Communication and Social Relationships in Captive Mixed-Species Flamingo Flocks.
リン・ショウヨ (D3) 飼育下の混合種フラミンゴ群における音声コミュニケーションと社会関係の研究. linchaoyu_mk [at] yahoo.co.jp
● Christen Lin (林佳翰) (D3) Empathy and emotional contagion in chimpanzees/bonobos. linchristen [at] gmail.com
クリステン・リン (D3) チンパンジー・ボノボにおける共感性および情動伝染. linchristen [at] gmail.com
● China Kobayashi (D3) Multi-dimensional sociality of horses. 2011.1b16 [at] gmail.com
小林知奈 (D3) ウマの多次元的な社会性の解明. 2011.1b16 [at] gmail.com
● Ayumi Ogawa (D2) Coordinated action for joint commitment in horse joint action. ogawa.ayumi.35x [at] st.kyoto-u.ac.jp
小川あゆみ (D2) ウマの共同行動における共同コミットメントのための調整行動. ogawa.ayumi.35x [at] st.kyoto-u.ac.jp
● Mayu Shimada (D1) Prestige: the evolution of social status. shimada.mayu.48u [at] st.kyoto-u.ac.jp
島田真優 (D1) プレステージ型社会的地位の進化. shimada.mayu.48u [at] st.kyoto-u.ac.jp
● Kaito Kimura (D1)
木村開人 (D1)
● Hamid Khan (M1) Emotional and behavioral regulations in Japanese macaques. hamidkhan081403 [at] gmail.com
ハミド・カーン (M1) ニホンザルの情動および行動制御 hamidkhan081403 [at] gmail.com
Visiting reseachers & students / 客員・訪問研究員、インターン生
● Dr. Cedric Sueur (2026 IFoHS Sabbatical Fellow, from University of Strasbourg, France)
● Evan Gurkin (2026 intern from University College London, England) Field experiments on self-control with Koshima monkeys
● Jade Primel (2026 intern from ENS, France) Social network and interaction around a blind chimpanzee at Kumamoto Sanctuary
● Rifa Rahma (2026 intern from Indonesia) Orangutan personality at Borneo Orangutan Survival Foundation in Kalimantan, Indonesia
● Applen Tahu (2026 intern from IISER Pune, India) Mutual grooming among lemurs at JMC
● Arthur Potereau (2026 intern from University of Nantes, France) Survey on mutual grooming across various animal taxa
● Danielle Wood (2026 intern from Max-Planck-Institute of Animal Behavior, Germany) Supporting experiments with elephants in Thailand
● Julien Paulet (2026 JSPS summer program from University of Strasbourg, France) AI individual identification of free-ranging Japanese monkeys
Almuni (students/postdocs etc) / 過去の在籍者
● Sofia Vilela (2021-2026 PhD) Empathy-related phenomena in Asian elephants. vpsofia.a [at] gmail.com
ソフィア・ヴィレラ（2021-2026 博士）アジアゾウにおける共感性. vpsofia.a [at] gmail.com
● Shimei Shirasawa (2023-2026 Master's) Resaerch plan to measure emotional states in free-ranging Japanese macaques using an infared thermal camera. shirasawa.shimei.75x [at] st.kyoto-u.ac.jp
白澤子銘（2023-2026 修士）野生のニホンザルにおいて赤外線カメラを用いた情動伝染の観察. shirasawa.shimei.75x [at] st.kyoto-u.ac.jp
● Liu Liu (2024-2026 Master's) Reunion responses in chimpanzees and bonobos. fibbin4fibbin [at] gmail.com
リウ・リウ (2024-2026 修士) 離合集散社会を築くチンパンジー・ボノボにおける再開時の社会交渉. fibbin4fibbin [at] gmail.com
● Yu Kaigaishi (2021-2025 JSPS research fellow PD, researcher) Inter-group variation in social tolerance among free-ranging Japanese macaques. kaigaraseki [at] gmail.com
貝ヶ石優 (2021-2025 日本学術振興会 特別研究員PD・研究員) 野生ニホンザルにおける寛容性の集団間変異. kaigaraseki [at] gmail.com
● Ena Onishi (2019-2025 Master's, PhD; Current position: Kyoto University Office of Research Acceleration) Synchronization / contagion of urination in chimpanzees. onishi.ena.84m [at] st.kyoto-u.ac.jp
大西絵奈 (2019-2025 修士・博士、現：京大総合研究推進本部) チンパンジーにおける排尿の同期・伝染. onishi.ena.84m [at] st.kyoto-u.ac.jp
● Madoka Hattori (2021-2024 PhD course; Current position: Lecturer at Kyoto City University of Arts) Bonding mechanisms between cats and humans. hattori.madoka.46p [at] st.kyoto-u.ac.jp
服部円（2021-2024 博士後期、現：京都市芸術大学講師）ネコとヒトとの絆形成メカニズムの解明. hattori.madoka.46p [at] st.kyoto-u.ac.jp
● James Brooks (2019-2024 Master's, PhD, JSPS International Research Fellow; Current position: Postdoc researcher at German Primate Center) The evolution of group-mindedness in great apes and domestic species. jamesgerardbrooks [at] gmail.com
ジェームズ・ブルークス (2019-2024 修士・博士・学振外国人特別研究員、現：ドイツ霊長類研究センター) 類人猿・家畜動物における集団性の進化. jamesgerardbrooks [at] gmail.com
● Monamie Ringhofer (2017-2021 researcher; Current position: Senior Lecturer at Teikyo University of Science) Social behavior and cognition in feral/domestic horses. monamie.ringhofer [at] gmail.com
リングホーファー 萌奈美（2017-2022 研究員、現：帝京科学大学講師）野生・飼育下ウマの社会行動と社会的認知を中心に研究。monamie.ringhofer [at] gmail.com
● Akiho Muramatsu (2020-2022 researcher; Current position: Assistant Professor at Akita Prefectural University) Number concept and working memory in chimpanzees / Social cognition in macaque species. akiho.muramatsu [at] gmail.com
村松明穂（2020-2022 研究員、現：秋田県立大学助教）チンパンジーにおける数の概念と作業記憶 / マカクにおける社会的認知．akiho.muramatsu [at] gmail.com
● Gema Palacino-Gonzalez (2021-2023 PhD course) The role of joint commitment and coordination during cooperative activities in horses. palacinogonzalez.gema [at] gmail.com
ヘマ・パラシノ (2021-2023 博士後期課程) ウマの協力・共同行動における2個体間の協調的相互作用. palacinogonzalez.gema [at] gmail.com
● Sok Hwan Lee (Brian) (2021-2023 Master's course) The evolution of prestige: effect of a novel skill in social rank of chimpanzees and bonobos. seokwhan94 [at] gmail.com
イ・ソクファン（ブライアン）(2021-2023 修士課程) 新しい知識・能力が社会的地位に与える影響の比較認知研究. seokwhan94 [at] gmail.com
Almuni (visiting researchers/ interns) / 過去の在籍者（訪問研究者・インターンほか）
● Dr. Cedric Sueur (2023, 2024, 2025 from University of Strasbourg, France)
● Dr. Federico Rosano (2025 from University of California, San Diego, US)
● Dr. Dan Schofield (2024 from University of Oxford, UK)
● Dr. Crickette Sanz (2019 from Washington University in St. Louis, US)
● Stijn Van Den Bossche (2025 intern from Ghent University, Belgium) Video analyses on dog behavior
● Theodora Icarre (2025 intern from ENS, France) Playback experiments with feral horses at Cape Toi
● Omer Yinon (2025 intern from Tel Aviv University, Israel) Playback experiments with chimpanzees at KS
● Nicolas Goppold (2025 intern from Germany) Playback experiments with chimpanzees at KS
● Anushka Vispute (2025 intern from the US) Playback experiments with chimpanzees at KS; Analysis of road crossing in wild chimpanzees
● Gourav Mandi (2025 intern from IISER Berhampur, India) Drone projects with wild elephants in India and a survey on elephant-human conflicts
● Emily Sands (2024 Intern from ENS, France) Deep learning for automatic facial detection and recognition in Japanese macaques in Shodoshima.
● Maxim Herbrich (2024 Intern from U of Strasbourg, France) Ecological and social factors on huddling behaviour in Japanese macaques in Shodoshima.
● Julien Paulet (2023 Intern from U of Strasbourg, France) Deep learning for automatic facial detectoin and recognition in Japanese macaques in Koshima.
● Chloé Gouyet (2019 intern from Sorbonne Université Paris 13, France) Long-term stability/change of social networks in captive chimpanzees at KS.
● Isabel Clark (2019 intern from Duke University, the US) Playback experiments and observation of chimpanzee social interactions at KS.