Selmeston has been a settlement since Mesolithic times. Early humans would have been attracted to this area because of its natural geographical features - and in particular, its water supply. It sits along a spring line, fed by the winter rains along the South Downs, effectively a huge block of porous chalk. Nearly every house built in the village before the 1960s has some form of well close by and it was only in the mid 1950s that running water was laid on to each property. There are no street lights or pavements and this gives the village some of its character, which is enhanced by large verges which are planted with wild flowers and daffodils by residents.
In the 1930s local archaeologists began to find flints in 'sandpits' to the south of Selmeston Church. These turned out to be 'dwelling pits' used by Mesolithic Man, yielding a total of 6,400 stones. Some of the stones were 'potboilers', heated red hot and then dropped into jugs of water to heat up. Others were sharpened flints, used as spears and arrow-heads by the hunter-gatherers. When the huts were discarded, the pits filled up with drifting sand.
A collection of flints from the Selmeston sandpits can be seen in the Barbican House Museum in Lewes.