Grasslands are dominated by a rich mix of grasses and forbs (non-grass plants); herbaceous and non-woody plants. They have low precipitation rates and are maintained by fire, which suppresses tree saplings. This biome becomes prominent during the Miocene epoch (23-5 Mya) due to a cooling, drying climate on Earth. Grasslands have very fertile soil; therefore, they have been extensively converted to agriculture. When the soils of grasslands are compromised, these biomes frequently become deserts.
Grasslands are sometimes also called a prairie, meadow, pasture, range, plain, steppe, pampas, sward, llano, or the veldt
Warm to hot summers, with cold, windy winters
Precipitation in grasslands is relatively low, usually occurring in the late spring and early summer: 50.8 to 127 cm/yr (20-50 in/yr)
Precipitation is the major abiotic factor defining this biome, along with the presence of fire to suppress woody saplings
The temperature range is very large over the course of the year
Summer temperatures can be well over 38°C (100°F),
Winter temperatures can be as low as -40°C (-40°F).
The entire biome has an average annual temperature between -10 and 20°C
North American Midwest, Eurasia, Argentina, Uruguay, South African veld
Above: A Whittaker Graph showing where grasslands occupy this abiotic space, based on precipitation and temperature
Some of the world's most fertile soils
They are sometimes referred to as mollisols (dark, thick, nutrient-rich horizon)
Thick, woven root systems of grasses hold nutrients in place
There are members of the Fabaceae in grasslands that fix nitrogen, adding to the fertility of the soil
Low rainfall prevents erosion and loss of nutrients
Fires constantly return organic material to the soils
Perennial grasses and perennial forbs*
Sod-forming and bunch grasses (Poaceae)
Composites (Asteraceae) and legumes (Fabaceae) are common
*"Forb" is a term that is short for euphorbia but refers to all non-grass plants in a grassland.
Rhizome-bearing plants, such as grasses
Basal meristems, to avoid large herbivores
Fire-resistance in plants
Bulliform cells in leaves
Wind-pollination in grasses
Large showy flowers in "forbs"
The origin of grasslands is linked to the evolution of grasses (Poaceae)
The oldest evidence of grasses comes from 100–97 mya (Cenomanian) from phytoliths in the fossilized dung (coprolites) of long-necked dinosuars called titanosaurs
Early grasses may have been marsh-edge plants, and not forming grassland biomes at this point
It is assumed that grasslands, as a dry, fertile biome, became prominent around 30 million years ago with the cooling and drying of the Earth (Oligocene Epoch)
The origin of the grassland biome gave rise to large herds of mammals during the past 30 million years (e.g., elephants, horses, camels, deer, rhinos, etc.) in Asia, Africa, and North America.
Prairie strips can rapidly improve soil health (Phys.org 10Dec2025)
└Prairie strips improve many measures of soil health in nearly a decade (Dutter et al., 2025)
Increasing plant diversity in agricultural grasslands boosts yields, reducing reliance on fertilizer (Phys.org 4Dec2025)
└Multispecies grasslands produce more yield from lower nitrogen inputs across a climatic gradient (O'Malley et al., 2025)
What meadows reveal about the future: Spatial data can predict biodiversity loss before species disappear (Phys.org 26Nov2025)
└Congruent direction but different magnitude of biodiversity response to land-use intensification in space and time (Neuenkamp et al., 2025)
Prescribed burning helps store forest carbon in big fire-resistant Sierra Nevada trees (Phys.org 17Nov2025)
└Carbon costs of different pathways for reducing fire hazard in the Sierra Nevada (Zhu et al., 2025)
Two woody meadows planted a decade ago to see what would thrive. Now the concept is popular across Australia (The Conversation, 13Nov2025)
Grassland degradation shifts biodiversity's role from plants to soil microbes (Phys.org 11Nov2025)
└Grassland degradation alters plant and soil biodiversity–multifunctionality relationships (Gao et al., 2025)
Reviving South Africa’s grasslands: Eastern Cape villagers explain the challenges they face (The Conversation 17Sep2025)
Bison herds 'reawaken' Yellowstone's prairies (Phys.org 31Aug2025)
└Geremia et al. (2025) Yellowstone's free moving large bison herds provide a glimpse of their past ecosystem function
Grassland loss and woody plant encroachment drive decline of golden-shouldered parrot (Phys.org 24Jul2025)
└Crowley et al. (2025) Woody plant encroachment drives the decline of a grassland bird: The fate of golden-shouldered parrot (Psephotellus chrysopterygius) nests
Nitrogen in forests and grasslands overestimated, new global analysis shows (Cary Institute of Ecosystem Studies 16Jul2025)
└Ely (2025) Global terrestrial nitrogen fixation and its modification by agriculture
Unexpectedly deep roots in plants unearth new questions about soil carbon storage (Phys.org 17Jun2025)
└Malhotra et al. (2025) Continental-scale relationships of fine root and soil carbon stocks hold in grasslands but not forests
Long-term study reveals native seeding controls annual—but not perennial—invasive plants in sand grassland restoration (Phys.org 17Jun2025)
└Sáradi et al. (2025) Restorative seeding controls annual invasive species, but perennials can thrive in the long term despite treatments in sand grassland restoration
Grassland ecosystems show abrupt changes when nitrogen fertilization exceeds key threshold (Phys.org 16Jun2025)
└Bagousse-Pinguet et al. (2025) Thresholds of functional trait diversity driven by land use intensification
Invasive South African Gazania poses huge risk in Australia (Phys.org 4Jun2025)
└Adnan et al. (2025) Seeds of success: seed biology and germination response of Gazania weed in Australia
Research shows how solar arrays can aid grasslands during drought (ScienceDaily 2Jun2025)
└Sturchio & Knapp (2025) Evidence of photovoltaic aridity mitigation in semi-arid grasslands.
Extreme drought can disrupt grassland stability, weakening dominant grasses' influence (Phys.org 27May2025)
└Luo et al. (2025) Extreme Drought Increases the Temporal Variability of Grassland Productivity by Suppressing Dominant Grasses
Faced with drought, fertilizer helps grasslands grow strong (Phys.org 19May2025)
└Bondaruk et al. (2025) Aridity modulates grassland biomass responses to combined drought and nutrient addition
Nutrient additions amplify grassland response to rainfall, study finds (Phys.org 17 Apr2025)
└Fay et al. (2025) Interactions among nutrients govern the global grassland biomass–precipitation relationship
A long wait for pollinators: Restored grasslands need 75+ years for full biodiversity recovery, study finds (Phys.org 13Mar2025)
└Hirayama et al. (2025) Long-term management is required for the recovery of pollination networks and function in restored grasslands
Grasslands are responding to climate change almost in real time, according to research (Phys.org 16Oct2024)
└Zhu et al. (2024) Rapid shifts in grassland communities driven by climate change
Grassland study shows that elevated levels of CO₂ nearly tripled species losses attributed to nitrogen pollution (Phys.org 16Oct2024)
└Reich et al. (2024) High CO2 dampens then amplifies N-induced diversity loss over 24 years
A rare orchid survives on a few tracts of prairie (Phys.org 24Aug2024)
Danish grasslands are behaving strangely and researchers don't know why (Phys.org 21Feb2024)
└ Damgaard (2024) Decline in large-seeded species in Danish grasslands over an eight-year period
Plant diversity stabilizes soil temperature (Phys.org 6Dec2023)
└Huang et al. (2023) Enhanced stability of grassland soil temperature by plant diversity
Where the bison could roam (New York Times 10Jan2023)
Reintroducing bison on tallgrass prairie doubles plant diversity (Successful Farming 1Sep2022)
Reconstructing Two Centuries of Midwest Prairie Fire History (In Defense of Plants 28Nov2021)