Index number theory

The apples and oranges theorem for price indices Economics Letters 2009

A note on different approaches to index number theory (with Roy van der Weide) American Economic Review 2008

Multilateral indices: conflicting approaches? (with John Quiggin) Review of Income and Wealth 2007 

An impossibility theorem concerning multilateral international comparison of volumes Econometrica 2002

Some papers fit in multiple categories. I tried to put them in the category where they fit the most.

Evolution 

commitment

Evolution and the ultimatum game (with Aslıhan Akdeniz) Games and Economic Behavior 2023

The evolution of morality and the role of commitment (with Aslıhan Akdeniz) Evolutionary Human Sciences 2021 YouTube1 YouTube2 YouTube3 

Predictably angry—Facial cues provide a credible signal of destructive behavior (with Boris van Leeuwen, Charles Noussair, Theo Offerman, Sigrid Suetens & Jeroen van de Ven) Management Science 2018.

repeated games

Repeated games with partner choice (with Christopher Graser, Takako Fujiwara-Greve and Julián García) PLoS Computational Biology 2025. 

Repeated prisoner's dilemmas with errors: How much subgame perfection, how much forgiveness, and how much cooperation? (with Christopher Graser) Tinbergen Institute Discussion Paper 2024

Repeated games with endogenous separation (with Segis and Luis Izquierdo) Working paper 2021

No winning strategy in the Iterated Prisoner’s Dilemma: Game Theory and Simulated Evolution (with Julián García) In Adaptive and Learning Agents Workshop at AAMAS 2019.

In and out of equilibrium II: Evolution in repeated games with discounting and complexity costs (with Julián García) Games and Economic Behavior 2019.

In and out of equilibrium I: Evolution of strategies in repeated games with discounting (with Julián García) Journal of Economic Theory 2016

Direct reciprocity in structured populations (with Julián García, David Rand & Martin Nowak) Proceedings of the National Academy of Sciences 2012

the Price equation

What is the price of using the Price equation in ecology? (with Pierrick Bourrat, William Godsoe, Pradeep Pillai, Tarik Gouhier, Werner Ulrich & Nicolas Gotelli) Oikos 2023

The problem with the Price equation Philosophical Transactions of the Royal Society B 2020

On the use of the Price equation Journal of Theoretical Biology 2005

Hamilton's rule

The general version of Hamilton’s rule eLife 2025 YouTube Referee reports and replies for an earlier submission

Can Hamilton’s rule be violated? eLife 2018

Hamilton's rule (with Benjamin Allen, Moshe Hoffmann, Burton Simon & Carl Veller) Journal of Theoretical Biology 2017

Hamilton's missing link Journal of Theoretical Biology 2007

group selection

The cancellation effect at the group level (with Aslıhan Akdeniz) Evolution 2020.

It takes grouping and cooperation to get sociality (with Julián García and Leticia Avilés) Journal of Theoretical Biology 2010

In love and war: altruism, norm formation, and two different types of group selection (with Astrid Hopfensitz) Journal of Theoretical Biology 2007

Why kin and group selection models may not be enough to explain human other-regarding behaviour Journal of Theoretical Biology 2006

group selection & inclusive fitness

A simple model of group selection that cannot be analyzed with inclusive fitness (with Shishi Luo & Burton Simon) Journal of Theoretical Biology 2014

Group selection and inclusive fitness are not equivalent; the Price equation vs. models and statistics (with Julián García, Maus Sabelis & Martijn Egas) Journal of Theoretical Biology 2012

A rule is not a rule if it changes from case to case (a reply to Marshall's comment) Journal of Theoretical Biology 2011

Group selection, kin selection, altruism and cooperation: when inclusive fitness is right and when it can be wrong Journal of Theoretical Biology 2009

(evil) green beards

Evil green beards: Tag recognition can also be used to withhold cooperation in structured populations (with Julián García & Arne Traulsen). Journal of Theoretical Biology 2014

intuition vs deliberation

A general evolutionary framework for the role of intuition and deliberation in cooperation (with Stephan Jagau) Nature Human Behaviour 2017

evolutionary game theory 

Multi-player games on the cycle (with Martin Nowak) Journal of Theoretical Biology 2012.

Robustness against indirect invasions Games and Economic Behavior 2012

The replicator dynamics with n players and population structure Journal of Theoretical Biology 2011

Evolution in games with a continuous action space (with Peter Spreij) Economic Theory 2009

miscellaneous 

Fisher’s fundamental theorem of natural selection, dynamic sufficiency, and the necessity of higher moments. Journal of Theoretical Biology 2018

Interpretations arising from Wrightian and Malthusian fitness under strong frequency dependent selection (with Bin Wu, Chaitanya Gokhale, Long Wang & Arne Traulsen) Ecology and Evolution 2013.

book chapters

Human cooperation among kin and close associates may require enforcement of norms by third parties (with Sarah Mathew and Rob Boyd) in: Cultural Evolution: Society, Technology, Language, and Religion 2013

Does it pay to be good? Competing evolutionary explanations of pro-social behaviour The moral brain: Essays on the evolutionary and neuroscientific aspects of morality 2009.

commentaries

The group selection–inclusive fitness equivalence claim: not true and not relevant Evolutionary Human Sciences 2020

Review of: S. Bowles, H. Gintis (2011) A cooperative species: human reciprocity and its revolution Journal of Economic Literature 2012

News and Views: Selection for positive illusions (with Martin Nowak) Nature 2011

Call for a return to rigour in models (with Julián García, Maus Sabelis & Martijn Egas) Nature 2010.