Google Scholar ( Click )

2026.8.24. update

Peer reviewed journals

1. Tajima, H., Nishizawa, H., Kume, M., Okuyama J., Mitamura H., Doi H. (2026) Interspecific Differences in the Origin of Assimilated Diet Among Three Omnivorous Ghost Crab Species on Resource-Limited Sandy Beach Based on Carbon and Nitrogen Stable Isotope Analysis. Estuaries and Coasts 49: 186. https://doi.org/10.1007/s12237-026-01799-y 

2. Nishizawa H., Okuyama J., Murai H., Noguchi N., Tanita I. (2026) Green turtle foraging patterns revealed by seagrass meadow-centric time-lapse camera and drone surveys. Regional Studies in Marine Science 97: 105016. doi.org/10.1016/j.rsma.2026.105016.

3. Okuyama J., Nishizawa H., Sudo K., Shimabukuro H., Yoshida N., Tsukakoshi Y., Murai H., Sasano S., Tamaki M. (2026) Intensive grazing by green turtles depletes seagrass meadows and constrains turtle growth in Kume Island, Japan. Scientific Reports 16: 18394. doi.org/10.1038/s41598-026-49615-8

4. Candela T., Gaspar P., Bailey H., Wyneken J., Turla E., Noble-Trull T., Nel R., Kanghae H., Suraswadi P., Upanoi T., Okuyama J., Kawazu I., Maeda K., Mizuochi K., Ogino N., Vandeperre F., Mafalda Sousa A., Herguedas A., De Boer M., Dell’Amico F., Nunes A., Neves J., Gaspar I., Shillinger G.L.  (2026) Pioneering insights into the diving behavior of early-stage sea turtles revealed by novel marine miniaturized satellite tags. Scientific Reports 16: 16692. doi.org/10.1038/s41598-026-47239-6

5. Okuyama J., Kishida N., Nishizawa H., Obe Y., Mogi T., Koizumi T., Noda T., Yasuda T., Kawabata Y., Yokota T., Ichikawa K., Mitsunaga Y., Arai N. (2025) Gravid green turtles employ a mixed capital–income breeding strategy where food is abundant. Marine Ecology Progress Series  764: 175-188. https://doi.org/10.3354/meps14885

6. Noguchi N, Nishizawa H, Shimizu T, Okuyama J, Kobayashi S, Tokuda K, Tanaka H, Kondo S (2025) Efficient wildlife monitoring: Deep learning-based detection and counting of green turtles in coastal areas. Ecological Informatics 86: 103009. doi.org/10.1016/j.ecoinf.2025.103009

7. Okuyama J, Nakagawa M, Taneda T. (2024) A rapid increase in tropical species of groupers in the temperate waters, the Goto Islands, Japan. PLOS ONE 19(9): e0308715. doi: 10.1371/journal.pone.0308715. (Open Access) 

8. Hamabata T, Kameda K, Hayashi R, Okuyama J, Kawazu I and Nishizawa H (2024) Latest and comprehensive mitochondrial DNA haplotype data on green and hawksbill turtles collected in their habitats in Japan. Frontiers in Marine Science 11:1328101. doi: 10.3389/fmars.2024.1328101

9. Noda T, Koizumi T, Yukitake N, Yamamoto D, Nakaizumi T, Tanaka K, Okuyama J, Ichikawa K, Hara T. (2024) Animal-borne soundscape logger as a system for edge classification of sound sources and data transmission for monitoring near-real-time underwater soundscape. Scientific Reports 14:6394. doi:10.1038/s41598-024-56439-x

10. Yasuda T., Kinoshita J, Niino Y, Okuyama J. (2023) Vertical migration patterns linked to body and environmental temperatures in chub mackerel. Progress in Oceanography 213: 103017. DOI: 10.1016/j.pocean.2023.103017

11. Fujita K, Nishizawa H, Okuyama J*, Arita M, Takuma S, Narazaki T, Watabe A. (2023) Polymorphic foraging tactics in a marine reptile: insight from horizontal movement and dive behavior analyses. Marine Ecology Progress Series 707:115-129. DOI:10.3354/meps14258. * Corresponding author

12. Okuyama J., Nakagawa M. (2023) Seasonal fluctuation in the length-weight relationship of groupers in the Goto Islands, Japan. Thalassas: An International Journal of Marine Sciences 39: 27-34. DOI: 10.1007/s41208-022-00483-9

13. Kishida N., Okuyama J.*, Arita M., Kume N., Fujita K., Nishizawa H., Torisawa S. Mitsunaga Y. (2022) A validation of abstracted dive profiles relayed via the Argos satellite system: a case study of a loggerhead turtle. Animal Biotelemetry 10: 21. (Open Access)  * Corresponding author

14. Okuyama J., Watabe A., Takuma S., Tanaka K., Shirai K., Murakami-Sugihara N., Arita M., Fujita K., Nishizawa H., Narazaki T., Yamashita Y., Kameda K. (2022) Latitudinal cline in the foraging dichotomy of loggerhead sea turtles reveals the importance of East China Sea for priority conservation. Diversity and Distributions 28: 1568-1581. (Open Access) 

15. Okuyama J., Yamaguchi T. (2022) Individual spawning performance and mating pair combinations in captive grouper aggregations.  Environmental Biology of Fishes 105: 385-401.

16. Tabuki K., Nishizawa H., Abe O., Okuyama J., Tanizaki S. Utility of carapace images for long-term photographic identification of nesting green turtles. Journal of Experimental Marine Biology and Ecology 545: 151632. 2021.

17. Okuyama J., Benson S.R., Dutton P.H., Seminoff J.A. Changes in dive patterns of leatherback turtles with sea surface temperature and potential foraging habitats.  Ecosphere 12(2): e03365. 2021. (Open Access) 

18. Okuyama J., Ishii H., Tanizaki S., T. Suzuki, Abe O., Nishizawa H., Yano A., Tsujimura M., Ishigaki T., Ishigaki T., Kobayashi M., Yanagida H. Quarter-Century (1993–2018) Nesting Trends in the Peripheral Populations of Three Sea Turtle Species at Ishigakijima Island, Japan. Chelonian Conservation and Biology 19(1): 101-110. 2020.

19. Okuyama J., Shiozawa M., Shiode D. Heart rate and cardiac response to exercise during voluntary dives in captive sea turtles (Cheloniidae). Biology open 9 (2), bio049247. 2020. (Open Access)

20. Shiozawa M., Shiode D., Okuyama J., Hu F., Tokai T. Behavioural characteristics of loggerhead turtles (Caretta caretta) in a submerged bag net of a setnet observed in a bycatch simulation and the development of a turtle releasing device. Aquatic Conservation: Marine and Freshwater Ecosystems. 29(12), 2107-2115. 2019

21. Okuyama J., Shishidou, H., Hayashibara, T. Post-release horizontal and vertical behavior and philopatry of deepwater longtail red snapper Etelis coruscans around a bank. Fisheries Science, 85(2), 361-368. 2019.

22. Nanami A., Sato T., Kawabata Y.,  Okuyama J. Spawning aggregation of white-streaked grouper Epinephelus ongus: spatial distribution and annual variation in the fish density within a spawning ground. PeerJ, 5, 2017. e3000. (Open Access) 

23. Okuyama J., Seminoff J.A., Dutton P.H., Benson S.R.. Fine-scale monitoring of routine deep dives by gravid leatherback turtles during the internesting interval indicate a capital breeding strategy. Frontiers in Marine Science (Marine Megafauna section). 3:166. 2016.  doi: 10.3389/fmars.2016.00166. (Open Access)

24. Okuyama J., Nakajima K.,  Matsui K., Nakamura Y., Kondo K., Koizumi T., Arai N.. Application of a computer vision technique to animal-borne video data: Extraction of head movement to understand sea turtles’ visual assessment of surroundings. Animal Biotelemetry 3: 35. 2015.  (Open Access)

25. Noda T., Okuyama J., Kawabata Y., Mitamura H., Arai N.. Harvesting energy from the oscillation of aquatic animals: testing a vibration-powered generator for bio-logging data logger systems. Journal of Advanceｄ Marine Science and Technology Society 20: 37-43. 2014.

26. Okuyama J., Tabata R., Nakajima K., Arai N., Kobayashi M., Kagawa S. Surfacers change their dive tactics depending on the aim of the dive: evidence from simultaneous measurements of breaths and energy expenditure. Proceedings of the Royal Society B 281: :20140040. 2014. doi: 10.1098/rspb.2014.0040

27. Okuyama J., Kagawa S., Arai N. Random mate-searching behavior: Male sea turtles judge visually similar objects to females as a mating partner. Chelonian Conservation and Biology 13(2) :278-281. 2014.   

28. Nishizawa H., Naito Y., Suganuma H., Abe O., Okuyama J., Hirate K., Tanaka S, Inoguchi E., Narushima K., Kobayashi K., Ishii H., Tanizaki S., Kobayashi M., Goto A., Arai N. Composition of green turtle feeding aggregations along the Japanese Archipelago: implications for changes in composition with current flow. Marine Biology 160: 2671-2685. 2013.

29. Nishizawa H., Noda T., Yasuda T., Okuyama J., Arai N., Kobayashi M. Decision tree classification of behaviors in nesting process of green turtles (Chelonia mydas) from tri-axial acceleration data. Journal of Ethology 31: 315-322. 2013.

30. Okuyama J., Nakajima K., Noda T., Kimura S., Kamihata H., Kobayashi M., Arai N., Kagawa S., Kawabata Y., Yamada H. Ethogram of immature green turtles: behavioral strategy for somatic growth in large marine herbivores. PLoS ONE 8(6) e65783. 2013. (Open Access)

31. Noda T., Okuyama J., Koizumi T., Arai N., Kobayashi M. Monitoring attitude and dynamic acceleration of free-moving aquatic animals using a gyroscope. Aquatic Biology 16: 265-276. 2013.

32. Okuyama J., Kataoka K., Kobayashi M., Abe O., Yoseda K., Arai N. The regularity of dive performance in sea turtles: a new perspective from precise activity data. Animal Behaviour 84: 349-359. 2012.

33. Nishizawa H., Okuyama J., Abe O., Kobayashi M., Arai N. Mitochondrial DNA variation in hawksbill turtles (Eretmochelys imbricata) nesting in Ishigaki Island, Japan. Marine Turtle Newsletter 132: 1-2. 2012.

34. Nishizawa H., Abe O., Okuyama J., Kobayashi M., Arai N., Population genetic structure and natal philopatry of nesting green turtles (Chelonia mydas) in Yaeyama Islands, Japan. Endangered Species Research. 14: 141-148. 2011.

35. Okuyama J., Kitagawa T., Zenimoto K., Kimura S., Arai N., Yoshikazu Sasai, Hideharu Sasaki. Trans-Pacific dispersal of loggerhead turtle hatchlings inferred from numerical simulation modeling. Marine Biology 158: 2055-2063. 2011.

36. Nishizawa H., Okuyama J., Abe O., Kobayashi M., Yoseda K., Arai N. The difference of activity between in the sea and in the tank of green turtle (Chelonia mydas) juveniles. Coastal Marine Science 34: 7-12. 2010.

37. Kawabata Y., Okuyama J., Asami K., Yoseda K., Arai N., Effects of a tropical cyclone on the distribution change of hatchery-reared black-spot tuskfish Choerodon schoenleinii, determined by acoustic telemetry. Journal of Fish Biology 77: 627-642. 2010.

38. Okuyama J., Shimizu T., Abe O., Yoseda K., Arai N. Wild versus head-started hawksbill turtles Eretmochelys imbricata: post-release behavior and feeding adaptions. Endangered Species Research 10: 181-190. 2010.

39. Okuyama J., Kawabata Y., Naito Y., Arai N., Kobayashi M., Monitoring beak movements with an acceleration datalogger: a useful technique for assessing the feeding and breathing behaviors of sea turtles. Endangered Species Research 10: 39-45. 2010.

40. Nishizawa H., Okuyama J., Kobayashi M., Abe O., Arai N. Comparative phylogeny and historical perspectives on population genetics of the Pacific hawksbill (Eretmochelys imbricata) and green turtles (Chelonia mydas). Zoological Science 27: 14-18. 2010.

41. Okuyama J., Abe O., Nishizawa H., Kobayashi M., Yoseda K., Arai N. Ontogeny of the dispersal movement of green turtle (Chelonia mydas) hatchlings. Journal of Experimental Marine Biology and Ecology 379: 43-50. 2009.

42. Mitamura H., Uchida K., Miyamoto Y., Arai N., Kakihara T., Yokota T., Okuyama J., Kawabata Y., Yasuda T. Preliminary study on homing, site fidelity, and diel movement of black rockfish Sebastes inermis measured by acoustic telemetry. Fisheries Science 75: 1133-1140. 2009.

43. Kawabata Y., Okuyama J., Asami K., Yoseda K., Arai N. The post-release process of establishing stable home ranges and diel movement patterns of hatchery-reared black-spot tuskfish. Journal of Fish Biology 73: 1770-1782. 2008.

44. Kawabata Y., Okuyama J., Asami K., Yoseda K., Arai N. Post-release movement and diel activity patterns of hatchery-reared and wild black-spot tuskfish Choerodon schoenleinii determined by ultrasonic telemetry. Fisheries Science 73: 1146-1153. 2007.

45. 奥山隼一, Charuchinda M., 荒井修亮, 守屋和幸. タイ国ラヨーン県におけるシステムダイナミクスを用いた沿岸管理政策の提言, 海洋理工学会誌, 11(2). 7-19. 2006.

Books & Article 

1. 奥山隼一.  コラム11  "ウミガメ"、奥山隼一、河端雄毅.  "Ⅲ‒10 バイオテレメトリー"、「ビジュアル詳細 サンゴ礁の世界―多面的に観る・知る・科学する―」、 日本サンゴ礁学会(監修)／梅澤 有・鹿熊 信一郎・栗原 晴子・高橋 そよ・山下 洋・山野 博哉(編). 朝倉書店. 192 pp. (LINK)

2. アレン玉井 光江（文）, 村本 ちひろ（絵）,  奥山 隼一（ウミガメ監修）. THE SEA TURTLE. 東京書籍. 2024年7月16日刊行. ISBN: 978-4-487-81748-1. （LINK）

3. 奥山隼一、宍道弘敏、海老沢明彦、山口智史、中川雅弘．スジアラ地域個体群の資源特性に見られる緯度勾配から推察する温暖化が沿岸性魚類へ与える影響．水産海洋研究　87(3): 198-199. 2023.

4. 渡辺伸一、野田琢嗣、小泉拓也、依田憲、吉田誠、岩田高志、西澤秀明、奥山隼一、青木かがり、木村里子、坂本健太郎、高橋晃周、前川卓也、楢崎友子、三田村啓理、佐藤　克文．Biologging intelligent Platform（BiP）により実現するバイオロギングデータの共有と海洋の可視化．日本生態学会誌 73: 9-22. 2023.

5. 黒部ゆみ（写真・文）, 奥山隼一（監修）.  海に生きる！ ウミガメの花子.  偕成社. 2022年9月刊行. ISBN: 978-4-03-333740-1. (LINK) 

6. 奥山隼一.   水産研究・教育機構広報誌 おさかな瓦版 No.96「ブダイ」 2020年7月刊行　（PDF） 

7. 奥山隼一.   水産研究・教育機構広報誌 FRA news No.60「 サンゴ礁と水産」サンゴ礁にすむ魚類の生態　 2019年9月刊行　（PDF） 

8. 奥山隼一  . 旅する動物図鑑（全３巻）第3巻旅する海の生き物．「産まれた砂浜をわすれない アカウミガメ」pp. 28-29. 筑摩書房. 2019年2月

9. 奥山隼一.   水産研究・教育機構広報誌 FRA news No.57「 バイオロギング －海の生き物の行動を知る－ 」バイオロギングで観る水産生物、バイオロギングが未来をひらく　 2018年12月刊行　（PDF） 

10. 奥山隼一 「シリーズ新・生命の輪63：沖へ向かう子ガメに秘められた驚きの能力」自然保護 No. 555、2017年1・2月号、発行：公益財団法人日本自然保護協会

11. 奥山隼一. バイオロギング2 ―動物たちの知られざる世界を探る―（日本バイオロギング研究会 編）京都通信社. pp.192-197. 2016年. 

12. Okuyama J. & Arai N.  Head-start (re-introduction) program of captive-reared hawksbill turtles in the Yaeyama Islands, Okinawa, Japan. In: IUCN Global Re-introduction Perspectives: 2016. Further case-studies from around the globe Edited by Pritpal S. Soorae. 72-75. 2016.

13. 奥山隼一.   カメカメラでウミガメを観察する─理に適ったアオウミガメの生態． 『海は百面相（京都大学総合博物館 編）』  京都通信社. pp.146-148. 2013年10月．

14. Arai N. & Okuyama J., Bio-logging In: Field Informatics (Ed. Ishida T.) Splinger –Verlag, New York Inc. pp.21-38.  2012年8月.

15. 奥山隼一 「仔ガメの放流会はいかにあるべきか？」 海洋と生物 第190号（2010年10月号） 特集：最新・ウミガメ学．生物研究社．pp. 430-435 2010年．

16. 奥山隼一 「2章.水産生物.2-1.水産動物.2-2-1.種類.b.海棲爬虫類. 改訂・水産海洋ハンドブック（竹内俊郎、中田英昭、和田時夫、上田宏、有本貴文、渡辺終五、中前明 共編）生物研究社. pp. 85-86. 2010年

17. 奥山隼一. バイオロギング―最新科学で解明する動物生態学―（日本バイオロギング研究会 編）京都通信社. pp.18-21, 30, 32-35, 40-43, 97. 2009年

18. 奥山隼一. 自然を観察し、予測する. 「フィールド情報学入門―自然観察、社会参加、イノベーションのための情報学」（京都大学フィールド情報学研究会 編）. 共立出版．pp.5-7. 2009年

19. 荒井修亮 ＆ 奥山隼一. バイオロギング. 「フィールド情報学入門―自然観察、社会参加、イノベーションのための情報学」（京都大学フィールド情報学研究会 編）. 共立出版．pp.25-40. 2009年

Peer reviewed conference proceedings

1. Kawabata Y, Okuyama J, Naito Y, Arai N, Kobayashi M. Preliminary result of the relationship between the breathing frequency and dynamic body acceleration (DBA) of the hatchery-reared loggerhead turtle Carretta carretta. Proceedings of the 5th international Symposium on SEASTAR2000 and Asian Bio-logging Science. 11-14. 2010

2. Okuyama J, Nishizawa H., Abe O., Kobayashi M., Yoseda K., Arai N. Preliminary result of ontogenic change of wave orientation of green turtles in the initial growth stages 4th Int. Symp. SEASTAR2000 and Bio-logging Science 23-27 2009

3. Nishizawa H., Okuyama J., Abe O., Kobayashi M., Yoseda K., Arai N. Genitic diversity of the green turtle nesting population in Yaeyama Archipelago. Japan4th Int. Symp. SEASTAR2000 and Bio-logging Science 15-18 2009

4. Kataoka K., Okuyama J., Kobayashi M., Abe O., Yoseda K., Yasuda T., Arai N. Comparison of the activity budget between wild and reared juvenile hawksbill turtles4th Int. Symp. SEASTAR2000 and Bio-logging Science 19-22 2009

5. Kataoka K., Okuyama J., Kobayashi M., Abe O., Yoseda K., Arai N. Establishment of biologging approach of sea turtle in the neritic environment 3rd Int. Symp. SEASTAR2000 and Bio-logging Science 31-34 2007

6. Kawabata Y., Okuyama J., Asami K., Yoseda K., Ichikawa K., Arai N. Possibility of position estimates for bottom-dwelling aquatic animals in a small scale using ultrasonic coded transmitters and passive monitoring receivers 3rd Int. Symp. SEASTAR2000 and Bio-logging Science 83-87 2007

7. Okuyama J., Shimizu T., Abe O., Yoseda K., Arai N. A new technique for monitoring the grazing behavior of hawksbill turtles (Eretmochelys imbricata) using acceleration data loggersInt. Symp. SEASTAR2000 and Bio-logging Science 37-40 2005

8. Okuyama J., Boonhai K., Boonmee P., Mananunsap S., Charuchida M., Arai N. The regional management model for ecotourism planning in the Rayong coastal area, Thailand 4th South East Asia Sea Turtle Associative Cooperative Research 55-60 2004

9. Okuyama J.,  Boonhai K., Boonmee P., Mananunsap S., Charuchida M., Arai N. Development on Sea Turtle Ecotourism in Mannai Island 3rd South East Asia Sea Turtle Associative Cooperative Research 71-78 2003

Conference proceedings and others

1. Okuyama J., Kengo Kataoka, Kobayashi M., Abe O., Yoseda K., Arai N.. Behavioral pattern of juvenile hawksbill turtles at Yaeyama Islands, JAPAN. Proceedings of 28th International Symposium on Sea Turtle Biology and Conservation in press

2. Okuyama J., Shimizu T., Abe O., Yoseda K., Arai N. A new technique for monitoring grazing behavior of hawksbill turtles (Eretmochelys imbricata) using acceleration data loggers Proc. 25th Int. Symp. Sea Turtle Biology and Conservation 2008

3. Okuyama J., Arai N. Application possibility of agent-based simulation for consensus building on the establishment of marine protected area in Thailand IEEE3128: Proceedings of International Conference on Informatics Education and Research for Knowledge-Circulating Society 213-216 2008

4. Okuyama J., Abe O., Nishizawa H., Kobayashi M., Yoseda K., Arai N. Differences in orientation and swimming behavior between the initial growth stages of green turtles Proceedings of 27th International Symposium on Sea Turtle Biology and Conservation 53 2008

5. Okuyama J., Shimizu T., Abe O., Yoseda K., Arai N. Behavior and feeding adaptation of head-started hawksbill turtles (Eretmochelys imbricata) Proc. 26th Int. Symp. Sea Turtle Biology and Conservation 108 2006

6. Okuyama J., Boonhai K., Boonmee P., Mananunsap S., Charuchida M., Arai N. The current situation of tourism and the possibility of ecotourism in Rayong, Thailand Proc. 6th International Conference on the Environmental Management of Enclosed Coastal Seas 103-111 2005