Peer-reviewed publications

(10) Pokharel SS, Chettri AK, Chatterjee S, Seshagiri PB, Sukumar R (2025) Physiological responses in free-ranging Asian elephant populations living across human-production landscapes. Scientific Reports 15 (1), 32365

(9) Takahashi K, Pokharel SS, Kinoshita K, Ishiwata K, Soeda Y (2024) A newly discovered woolly mammoth molar from off the Notsuke Peninsula in Hokkaido, Japan. Journal of Fossil Research 57 (1・2), 50-57

(8) Pokharel SS, Brown JL (2023) Physiological plasticity in elephants: highly dynamic glucocorticoids in African and Asian elephants. Conservation Physiology 11: coad088. doi.org/10.1093/conphys/coad088

(7) Pokharel SS, Sharma N, Sukumar R (2022) Viewing the rare through public lenses: insights into dead calf carrying and other thanatological responses in Asian elephants using YouTube videos. Royal Society Open Science 9(5):211740. doi.org/10.1098/rsos.211740

(6) Pokharel SS, Yoneda H, Yanagi M, Sukumar R, Kinoshita K (2021) The tail-tale of stress: an exploratory analysis of cortisol levels in the tail-hair of captive Asian elephants. PeerJ 9: e10445. doi.org/10.7717/peerj.10445

(5) Sharma N, Pokharel SS, Kohshima S, Sukumar R. 2020. Behavioural responses of free-ranging Asian elephants (Elephas maximus) towards dying and dead conspecifics. Primates: 61, 129–138. doi.org/10.1007/s10329-019-00739-8 (Equal first author)

(4) Pokharel SS, Seshagiri PB, Sukumar R (2020) Influence of the number of calves and lactating adult females in a herd on the adrenocortical activity of free-ranging Asian elephants. Wildlife Research: 46(8), 679-689. doi.org/10.1071/WR18163

(3) Hanya G, Tackmann J, Sawada A, Lee W, Pokharel SS, de Castro Maciel VG, Toge A, Kuroki K, Otsuka R, Mabuchi R, Liu J (2020) Fermentation Ability of Gut Microbiota of Wild Japanese Macaques in the Highland and Lowland Yakushima: In Vitro Fermentation Assay and Genetic Analyses. Microbial Ecology: 24, 1-6

(2) Pokharel SS, Seshagiri PB, Sukumar R (2019) Lower levels of glucocorticoids in crop-raiders: Diet quality as a potential ‘pacifier’ against stress in free-ranging Asian elephants in a human-production habitat. Animal Conservation: 22(2), 177-188. doi.org/10.1111/acv.12450

(1) Pokharel SS, Seshagiri PB, Sukumar R (2017) Assessment of season-dependent body condition scores in relation to faecal glucocorticoid metabolites in free-ranging Asian elephants. Conservation Physiology 5: cox039. doi.org/10.1093/conphys/cox039


(Reach out to me if you need PDF copies)

 

Conference proceedings (*peer-reviewed proceedings)

18. Pokharel SS (2025) A tale of the elephant who wandered beyond the ‘forest’. Living Together: Human–Wildlife Relationships in a Changing World

Exploring Ecological, Ethical, and Cultural Dimensions of Coexistence. Uehiro Research Center for Japan Environmental Studies (UJES), Research Institute for Humanity and Nature (RIHN) and Indian Institute of Science (IISc) Interactive Meeting. (Invited talk)

17. Pokharel SS (2024) Physiological adaptations in large mammals: Exploring the present and the past (proboscideans). Stress and aging: a Japan-India biological research collaboration to meet 21st century challenges. IISc-Kyoto University Joint Workshop/Symposium-2024, India. (Invited talk)

16. Pokharel SS, Sharma N (2024) Dead Calf Carrying in Asian Elephants. First International Symposium on Comparative Evolutionary Thanatology, Kyoto University, Japan. (Invited talk)

15. Pokharel SS, Brown JL (2023) Physiological plasticity in elephants. 8th International Conference of International Society of Wildlife Endocrinology, Jim Corbett National Park, India.*

14. Pokharel SS (2023) Physiological tunings: How elephants react to changing ecological contexts. 4th DST-SERB School on Chemical Ecology, India. (Invited public talk)

13. Pokharel SS, Brown JL, Kinoshita K, Seshagiri PB, Sukumar R (2021) Cortisol as a clue of physiological stress: Decoding in extant species and its insights into extinct species. VIII International Conference on Mammoths and their Relatives, India.*

12. Pokharel SS (2021) Stress and elephants: Physiological adaptation to the changing ecological milieu. The Evolution and Adaptation: Current Scenario conference, GITAM University India. (Invited panel talk)

11. Pokharel SS, Yoneda H, Kinoshita K (2019) The tail-tale of stress: an exploratory study to assess hair cortisol in Asian elephants. A paper presented at the 7th Conference of the International Society of Wildlife Endocrinology, Kruger National Park, hosted by the Mammal Research Institute of the University of Pretoria and South African National Parks.* (Best Presentation Award)

10. Pokharel SS, Seshagiri PB, Sukumar R (2019) Lower levels of stress in crop-raiding Asian elephants: diet as a potential ‘pacifier’. Symposium of Integrative Biology: Biodiversity in Asia, Center for Ecological Research, Kyoto University, Japan. (Invited panel talk)

9. Pokharel SS, Seshagiri PB, Sukumar R (2018) Influence of ecological correlates on the stress status of free-ranging adult female Asian elephants. A paper presented at the 8th International Seminar on Biodiversity and Evolution, Wildlife Research Centre, Kyoto University, Japan.

8. Pokharel SS, Seshagiri PB, Sukumar R (2017) Herd-size, lactational-status and landscape affect stress status of free-ranging adult female Asian elephants: variations in fecal glucocorticoid metabolites. A paper presented at the 6th Conference of the International Society of Wildlife Endocrinology (ISWE), Florida, USA.* (Awarded with the International Travel Grant-ISWE)

7. Pokharel SS, Seshagiri PB, Sukumar R (2016) Stress physiology of free-ranging Asian elephants. A poster presented at the 15th International Elephant and Rhino Conservation and Research Symposium, Wildlife Reserves Singapore, Singapore.*

6. Pokharel SS, Seshagiri PB, Sukumar R (2016) Influence of intrinsic and extrinsic stressors on the stress status of free-ranging Asian elephants. A paper presented at the International Seminar on Biodiversity and Evolution, Wildlife Research Centre, Kyoto University, Japan.

5. Pokharel SS, Sukumar R (2016) Chemical Communications in Elephants. A paper presented at the SERB Collaborations in Indian Chemical Ecology, National Centre for Biological Sciences (NCBS), India. (Invited talk)

4. Pokharel SS, Seshagiri PB, Sukumar R (2015) Ecological variations impact the stress status of free-ranging Asian elephants. A paper presented at the 5th Conference of International Society of Wildlife Endocrinology, Berlin, Germany.*

3. Pokharel SS, Seshagiri PB, Sukumar R (2015) Use of non-invasive tool to understand the stress status in free-ranging Asian elephants. A poster presented at the National workshop on Reproduction and Welfare of endangered animals in conservation breeding, Laboratory for the Conservation of Endangered Species (LaCONES-CCMB), India.*

2. Pokharel SS, Chaudhary S, Uniyal VP (2010) Assessment of avian diversity in the human-dominated urban areas. A paper presented at the International Conference on Cooling the Earth: Tactics for Restoring Climate Order and Saving the Living Plane, G.B. Pant University of Agriculture and Technology, India.

1. Pokharel SS (2010) Reviewing the probabilities of Glacier Lake Outburst Floodings in Nepal. A review presented at the International Conference on Climate Change and Sustainable Management of Natural Resources, Institute of Technology and Management-Gwalior, India. (Invited panel talk)


Popular articles

8. Pokharel SS (2025)  The Big Bulls of Bardia. The Nepali Times.

7. Pokharel SS (2024)  Making festivals elephant-friendly in Nepal. The Nepali Times.

6. Pokharel SS (2022) Talking of elephants: A tug of war. The Himalayan Times (Topics).

5. Pokharel SS (2022) Crossing the border: viewing through elephants’ lens. The Hindu (Science).

4. Pokharel SS (2020) Baby Bhunte Explains Causes of Stress Among Elephants and Ways to Assess It. The Times of India (The Weather Channel).

3. Pokharel SS (2018) When an elephant dies. Current Conservation.

2. Pokharel SS (2017) Learning from the Locals. Sanctuary Asia.

1. Pokharel SS, Pradhan NMB, Dahal S. (2016) Asian elephant ‘Haathi’-WWF Learning Portal. A pictorial book on the ecology of Asian elephants. WWF Nepal, Hariyo Ban Program, Nepal.


Invited Public Lectures


28. 2025  Elephants, stress, and Bardia: The future of elephant science in Nepal, Bardia National Park, Nepal

27. 2025  Are elephants adapting to the altered ecosystems? Kathmandu University, Nepal

26. 2024  Understanding the science behind elephant physiology. St. Berchmans College, India

25. 2024  Understanding the physiology of Asian elephants. Shree JCBM College, India

24. 2024  Rethinking wildlife research through elephants’ lens. Nepal Development Research Institute, Nepal

23. 2024  Science behind stress: Learning the physiology of elephants. Sree Narayana College, India

22. 2023  Why do elephants visit my village? ENVIS: Amrita Viswha Vidyapeetham, India

21. 2023  Understanding the physiological dimensions of Asian elephant ecology. University of Adelaide, Australia

20. 2023  Learning the stories and science of Asian elephants. Aichi Prefectural Zuiryo High School, JSPS Science Dialogue Lecture series, Japan

19. 2022  Learning the lesser-known science of Asian elephants. BNHS: Wildlife Week Special Talk, India

18. 2022  The Jumbo Science: Understanding the basics of elephant ecology. ENVIS: Government of India, India

17. 2021  Conservation, cortisol and elephants: Insights into the probable ‘stress’. Elephant Professional Lecture Series. Golden Triangle Asian Elephant Foundation, Thailand.

16. 2021  Solving conservation puzzles: Animals’ physiology as a clue. Society for Conservation Biology (SCB-Nepal chapter). Conservation Talk Series 4, Nepal

15. 2021  Opportunities in Wildlife Research. Career counselling lecture series. GB Pant University, India

14. 2021  Understanding restoration: Insights into elephant ecology. Nilamber Pitamber University, India

13. 2021  What we can learn from an elephant (Hindi talk). Heartfulness Organization- Youth Wing, India

12. 2021  Endangered species: The Asian elephants. World Organizations of Students and Youths (WOSY), India

11. 2021  Bhunte the baby elephant. Women in Nature Network Talk, USA

10. 2020  Effects of megaprojects on elephant ecology. Xavier Institute Talk Series, Nepal

9. 2020  Nijgadh disaster and elephant puzzles. Power Shift Nepal. Nepal

8. 2020  Career in Wildlife Science. Wonder Girls Academy, India

7. 2020  Journey to the elephant land. Kurukshetra University, India

6. 2020  Multiple lecture series: Elephant ecology and conservation. Kalinga Center for Rainforest Ecology, India

5. 2019  Know your elephants. Ministry of Environment and Forestry (ENVIS) and IISc, India

4. 2017  Elephants among us. Green Skill Development Program by Government of India (ENVIS), India

3. 2017  Elephant physiology. Ashoka Trust for Research in Ecology and the Environment (ATREE), India

2. 2017  What do tigers and elephants tell us? Brigade School-Bangalore, India

 1. 2016  Poetry and elephants. Friends of Elephant Society, India