Plant ecology is a sub discipline of ecology which studies the distribution and abundance of plants, the effects of environmental factors upon the abundance of plants, and the interactions among and between plants and other organisms. Examples of these are the distribution of temperate deciduous forests in North America, the effects of drought or flooding upon plant survival, and competition among desert plants for water, or effects of herds of grazing animals upon the composition of grasslands.
Tropical plant community on Diego Garcia
A global overview of the Earth's major vegetation types is provided by O.W. Archibold.He recognizes 11 major vegetation types: tropical forests, tropical savannas, arid regions (deserts), Mediterranean ecosystems, temperate forest ecosystems, temperate grasslands, coniferous forests, tundra (both polar and High Mountain), terrestrial wetlands, freshwater ecosystems and coastal/marine systems. This breadth of topics shows the complexity of plant ecology, since it includes plants from floating single-celled algae up to large canopy forming trees.
One feature that defines plants is photosynthesis. One of the most important aspects of plant ecology is the role plants have played in creating the oxygenated atmosphere of earth, an event that occurred some 2 billion years ago. It can be dated by the deposition of banded iron formations, distinctive sedimentary rocks with large amounts of iron oxide. At the same time, plants began removing carbon dioxide from the atmosphere, thereby initiating the process of controlling Earth's climate. A long term trend of the Earth has been toward increasing oxygen and decreasing carbon dioxide, and many other events in the Earth’s history, like the first movement of life onto land, are likely tied to this sequence of events.
The study of plants and vegetation is complicated by their form. First, most plants are rooted in the soil, which makes it difficult to observe and measure nutrient uptake and species interactions. Second, plants oftenreproduce vegetatively, that is asexually, in a way that makes it difficult to distinguish individual plants. Indeed, the very concept of an individual is doubtful, since even a tree may be regarded as a large collection of linked meristems. Hence, plant ecology and animal ecology have different styles of approach to problems that involve processes like reproduction, dispersal and mutualism. Some plant ecologists have placed considerable emphasis upon trying to treat plant populations as if they were animal populations, focusing on population ecology. Many other ecologists believe that while it is useful to draw upon population ecology to solve certain scientific problems, plants demand that ecologists work with multiple perspectives, appropriate to the problem, the scale and the situation.
Plants, like most life forms, require relatively few basic elements: carbon, hydrogen, oxygen, nitrogen, phosphorus and sulphur; hence they are known as CHNOPS life forms. There are also lesser elements needed as well, frequently termed micronutrients, such as magnesium and sodium. When plants grow in close proximity, they may deplete supplies of these elements and have a negative impact upon neighbours. In many cases (perhaps most) the negative effects upon neighbours arise from competition for light, with larger plants shading smaller plants. In other cases, there may be competition below ground for water, nitrogen, or phosphorus. To detect and measure competition, experiments are necessary; these experiments require removing neighbours, and measuring responses in the remaining plants. Many such studies are required before useful generalizations can be drawn.
Overall, it appears that light is the most important resource for which plants complete, and the increase in plant height over evolutionary time likely reflects selection for taller plants to better intercept light. Many plant communities are therefore organized into hierarchies based upon the relative competitive abilities for light. In some systems, particularly infertile or arid systems, below ground competition may be more significant. Along natural gradients of soil fertility, it is likely that the ratio of above ground to below ground competition changes, with higher above ground competition in the more fertile soils. Plants that are relatively weak competitors may escape in time (by surviving as buried seeds) or in space (by dispersing to a new location away from strong competitors.)
In principle, it is possible to examine competition at the level of the limiting resources if a detailed knowledge of the physiological processes of the competing plants is available. However, in most terrestrial ecological studies, there is only little information on the uptake and dynamics of the resources that limit the growth of different plant species, and, instead, competition is inferred from observed negative effects of neighboring plants without knowing precisely which resources the plants were competing for. In certain situations, plants may compete for a single growth-limiting resource, perhaps for light in agricultural systems with sufficient water and nutrients, or in dense stands of marsh vegetation, but in many natural ecosystems plants may be co limited by several resources, e.g. light, phosphorus and nitrogen at the same time.[
There are therefore many details that remain to be uncovered, particularly the kinds of competition that arise in natural plant communities, the specific resource(s), the relative importance of different resources, and the role of other factors like stress or disturbance in regulating the importance of competition.
Mutualism is defined as an interaction "between two species or individuals that is beneficial to both". Probably the most widespread example in plants is the mutual beneficial relationship between plants and fungi, known asmycorrhizae. The plant is assisted with nutrient uptake, while the fungus receives carbohydrates. Some the earliest known fossil plants even have fossil mycorrhizae on their rhizomes.
The flowering plants are a group that has evolved by using two major mutualisms. First, flowers are pollinated by insects. This relationship seems to have its origins in beetles feeding on primitive flowers, eating pollen and also acting (unwittingly) as pollinators. Second, fruits are eaten by animals, and the animals then disperse the seeds. Thus, the flowering plants actually have three major types of mutualism, since higher plants also have mycorrhizae. Plants may also have beneficial effects upon one another, but this is less common. Examples might include "nurse plants" whose shade allows young cacti to establish. Most examples of mutualism, however, are largely beneficial to only one of the partners, and may not really be true mutualism. The term used for these more one-sided relationships, which are mostly beneficial to one participant, is facilitation. Facilitation among neighboring plants may act by reducing the negative impacts of a stressful environment Ingeneral; facilitation is more likely to occur in physically stressful environments than in favorable environments, where competition may be the most important interaction among species.Commensalism is similar to facilitation, in that one plant is mostly exploiting another. A familiar example is the ephiphytes which grow on branches of tropical trees, or even mosses which grow on trees in deciduous forests.
It is important to keep track of the benefits received by each species to determine the appropriate term. Although people are often fascinated by unusual examples, it is important to remember that in plants, the main mutualisms are mycorrhizae, pollination, and seed dispersal.