Anthocyanins are one group of the most abundant pigments in plants. Their biosynthesis is strongly regulated by different environmental factors controlling gene expression. Protein complexes consisting of WD40-bHLH-MYB (WBM) are have been identified to regulate the biosynthetic pathway of anthocyanins. We have established red pap1-D cells of Arabidopsis thaliana as a platform to understand accurate mechanisms of cell specificity of anthocyanin biosynthesis and elucidate regulation mechanisms by different factors