People

Associate Professor, Chonnam National University, 2023-Present

Assistant Professor, Chonnam National University, 2019-2023

Post-doc. Michigan State University, 2017-2019

Ph.D. University of Massachusetts, Amherst 2017

M.S. Chungnam National University 2011

B.S. Chungnam National University 2009

Editor in Plant Disease, The Plant Pathology Journal, Mycobiology, Research in Plant Disease, and The Korean Journal of Mycology 

Email: hksang@jnu.ac.kr

More information: https://mimb.jnu.ac.kr/mimb/5343/subview.do#none 

B.S., Chonnam National University 2020

Email: pswfuture7@gmail.com

1. Park SW†, An W†, Choi S, Choi H, Choi JH, Park S, Jang HR, Back CG, Hong SW, Sang H*.  2024. Evaluation of fungicidal control of garlic leaf blight using mobile spraying equipment.Korean J. Pestic. Sci. 28(3): 291-297 (†Equal contribution)

2. Ma J†, Park SW†, Kim G, Kim CS, Chang HX, Chilvers MI*, Sang H*.  2024. Characterization of SsHog1 and Shk1 using efficient gene knockout systems through repeated protoplasting and CRISPR/Cas9 ribonucleoprotein approaches in Sclerotinia sclerotiorum. Journal of Agricultural and Food Chemistry 72:4237-4245

3. Choi H†, Park SW†, Oh J, Kim CS, Sung GH*, Sang H*. 2023. Efficient disruption of CmHk1 using CRISPR/Cas9 ribonucleoprotein delivery in Cordyceps militaris. FEMS Microbiology Letters 370:fnad072 

4. Park S-W†, Paul NC†, Lee K-H, Han GH, Kim H-J*, Sang H*. 2023. Identification and Characterization of Foliar Fungi Associated with Beach Vitex (Vitex rotundifolia L.). Forests 14, 220

5. Paul NC†, Park SW†, Liu H, Choi S, Ma J, MacCready JS, Chilvers MI, Sang H*. 2021. Plant and Fungal Genome Editing to Enhance Plant Disease Resistance Using the CRISPR/Cas9 System. Frontiers in Plant Science 12:700925 (†Equal contribution)

B.S., Chonnam National University 2021

Email: sde982002@gmail.com

1. Choi S†, Son D†, Jang HR, Park S, Liu H, Lee HG, Sang H*. Ipflufenoquin, Novel Dihydroorotate Dehydrogenase Inhibitor: Efficacy Against Strawberry Diseases and Docking Simulations via AlphaFold2 and Autodock Vina. Preprint at https://doi.org/10.1101/2024.12.17.628804 (†Equal contribution)

2. Kim G, Son D, Choi S, Liu H, Nam Y, Sang H*. 2023. Monitoring of benzimidazole resistance in Botrytis cinerea Isolates from strawberry in Korea and development of detection method for benzimidazole resistance. The Plant Pathology Journal 39(6): 614-624

3. Choi S†, Son D†, Chilvers MI, Kim HJ, Sang H*. 2023. First report of Diaporthe eres causing leaf spot disease on Machilus thunbergii in Korea. Plant Disease 107: 1225

4. Sang H*, Chang HX, Choi S, Son D, Lee G, Chilvers MI*.  2022. Genome-wide transcriptional response of the causal soybean sudden death syndrome pathogen Fusarium virguliforme to a succinate dehydrogenase inhibitor fluopyram. Pest Management Science 78 (2): 530-540

B.S., Chonnam National University 2021

Email: maji2427@gmail.com

1. Ma J†, Park SW†, Kim G, Kim CS, Chang HX, Chilvers MI*, Sang H*.  2024. Characterization of SsHog1 and Shk1 using efficient gene knockout systems through repeated protoplasting and CRISPR/Cas9 ribonucleoprotein approaches in Sclerotinia sclerotiorum. Journal of Agricultural and Food Chemistry 72:4237-4245

2. Paul NC†, Park SW†, Liu H, Choi S, Ma J, MacCready JS, Chilvers MI, Sang H*. 2021. Plant and Fungal Genome Editing to Enhance Plant Disease Resistance Using the CRISPR/Cas9 System. Frontiers in Plant Science 12:700925 (†Equal contribution)

B.S., Chonnam National University 2021

Email: gaaheee0@gmail.com

1. Liu H, Lee G, Sang H*. 2024. Exploring SDHI fungicide resistance in Botrytis cinerea through genetic transformation system and AlphaFold model-based molecular docking. Pest Management Science 80:5954-5964

2. Lee G†, Choi H†, Liu H†, Han YH†, Paul NC, Han GH, Kim H, Kim PI, Seo SI, Song J, Sang H*. 2023. Biocontrol of the causal brown patch pathogen Rhizoctonia solani by Bacillus velezensis GH1-13 and development of a bacterial strain specific detection method. Frontiers in Plant Science 13:1091030

3. Sang H*, Chang HX, Choi S, Son D, Lee G, Chilvers MI*.  2022. Genome-wide transcriptional response of the causal soybean sudden death syndrome pathogen Fusarium virguliforme to a succinate dehydrogenase inhibitor fluopyram. Pest Management Science 78 (2): 530-540

M.S., Chonnam National University 2023

B.S., Chonnam National University 2021

Email: chhj3270@gmail.com

1. Liu H, Choi H, Paul NP, Ariyawansa HA, Sang H*. 2025. Discovering fungal communities in roots of Zoysia japonica and characterizing novel species and their antifungal activities. IMA Fungus 16: e138479

2. Park SW†, An W†, Choi S, Choi H, Choi JH, Park S, Jang HR, Back CG, Hong SW, Sang H*.  2024. Evaluation of fungicidal control of garlic leaf blight using mobile spraying equipment.Korean J. Pestic. Sci. 28(3): 291-297 (†Equal contribution)

3. Choi H†, Paul NC†, Kim HJ, Sang H*. 2023. Identification and fungicide sensitivity of Fusarium asiaticum causing seedling rot of Hinoki cypress in a Forest nursery in South Korea. Forest Pathology 53:e12837

4. Choi H†, Park SW†, Oh J, Kim CS, Sung GH*, Sang H*. 2023. Efficient disruption of CmHk1 using CRISPR/Cas9 ribonucleoprotein delivery in Cordyceps militaris. FEMS Microbiology Letters 370:fnad072 

5. Lee G†, Choi H†, Liu H†, Han YH†, Paul NC, Han GH, Kim H, Kim PI, Seo SI, Song J, Sang H*. 2023. Biocontrol of the causal brown patch pathogen Rhizoctonia solani by Bacillus velezensis GH1-13 and development of a bacterial strain specific detection method. Frontiers in Plant Science 13:1091030

B.S., Chonnam National University 2022

Email: psyangel0522@gmail.com

1.  Lee JG†, Ji JH†, Ryu S, Park JW, Paul NC, Park S, Im S, Han GH*, Sang H*. 2025, Control of sweet potato Fusarium wilt and Black rot by endophytic Bacillus velezensis with improved lipopeptide production. BMC Microbiology. Accepted. (†Equal contribution)

2. Choi S†, Son D†, Jang HR, Park S, Liu H, Lee HG, Sang H*. Ipflufenoquin, Novel Dihydroorotate Dehydrogenase Inhibitor: Efficacy Against Strawberry Diseases and Docking Simulations via AlphaFold2 and Autodock Vina. Preprint at https://doi.org/10.1101/2024.12.17.628804 (†Equal contribution)

3. Park SW†, An W†, Choi S, Choi H, Choi JH, Park S, Jang HR, Back CG, Hong SW, Sang H*.  2024. Evaluation of fungicidal control of garlic leaf blight using mobile spraying equipment.Korean J. Pestic. Sci. 28(3): 291-297 (†Equal contribution)

4. Yoon YS†, Lee JG†, Paul NC, Park S, Ryu S, Kim H, Im S, Sang H*, Han GH*. 2024. Biological control of a novel strain Bacillus velezensis CMML21-47 against sweet potato wilt and black rot diseases. Biological Control 195:105541

5. Lee JG, Paul NC, Park S, Kim HJ, Sang H*. 2023. First report of binucleate Rhizoctonia AG-G causing root rot of Japanese bay tree (Machilus thunbergii) in Korea. Plant Disease 107:2220

6. Paul NC, Park S, Liu H, Lee JG, Han GH, Kim H, Sang H*. 2021. Fungi Associated with Postharvest Diseases of Sweet Potato Storage Roots and In Vitro Antagonistic Assay of Trichoderma harzianum against the Diseases. Journal of Fungi 7: 927

B.S., Chonnam National University 2022

Email: wnrud2338@gmail.com

1.  Lee JG†, Ji JH†, Ryu S, Park JW, Paul NC, Park S, Im S, Han GH*, Sang H*. 2025, Control of sweet potato Fusarium wilt and Black rot by endophytic Bacillus velezensis with improved lipopeptide production. BMC Microbiology. Accepted. (†Equal contribution)

2. Yoon YS†, Lee JG†, Paul NC, Park S, Ryu S, Kim H, Im S, Sang H*, Han GH*. 2024. Biological control of a novel strain Bacillus velezensis CMML21-47 against sweet potato wilt and black rot diseases. Biological Control 195:105541 (†Equal contribution)

3. Lee JG, Paul NC, Park S, Kim HJ, Sang H*. 2023. First report of binucleate Rhizoctonia AG-G causing root rot of Japanese bay tree (Machilus thunbergii) in Korea. Plant Disease 107:2220

4. Paul NC, Park S, Liu H, Lee JG, Han GH, Kim H, Sang H*. 2021. Fungi Associated with Postharvest Diseases of Sweet Potato Storage Roots and In Vitro Antagonistic Assay of Trichoderma harzianum against the Diseases. Journal of Fungi 7: 927

M.S., Chonnam National University 2024

B.S., Jeonbuk National University 2017

Email: ja2hacho2@gmail.com

1. Park SW†, An W†, Choi S, Choi H, Choi JH, Park S, Jang HR, Back CG, Hong SW, Sang H*.  2024. Evaluation of fungicidal control of garlic leaf blight using mobile spraying equipment.Korean J. Pestic. Sci. 28(3): 291-297 (†Equal contribution)

M.S. Chungnam National University 2010

B.S. Chungnam National University 2008

Email: boramrhee@gmail.com

B.S., Chonnam National University 2024

Email: guri6634@gmail.com

B.S., Chonnam National University 2024

Email: mejina3925@gmail.com

1. Choi S†, Son D†, Jang HR, Park S, Liu H, Lee HG, Sang H*. Ipflufenoquin, Novel Dihydroorotate Dehydrogenase Inhibitor: Efficacy Against Strawberry Diseases and Docking Simulations via AlphaFold2 and Autodock Vina. Preprint at https://doi.org/10.1101/2024.12.17.628804 (†Equal contribution)

2. Park SW†, An W†, Choi S, Choi H, Choi JH, Park S, Jang HR, Back CG, Hong SW, Sang H*.  2024. Evaluation of fungicidal control of garlic leaf blight using mobile spraying equipment.Korean J. Pestic. Sci. 28(3): 291-297 (†Equal contribution)

B.S., Chonnam National University 2024

Email: 205480jnu@gmail.com

1. Choi S†, Son D†, Jang HR, Park S, Liu H, Lee HG, Sang H*. Ipflufenoquin, Novel Dihydroorotate Dehydrogenase Inhibitor: Efficacy Against Strawberry Diseases and Docking Simulations via AlphaFold2 and Autodock Vina. Preprint at https://doi.org/10.1101/2024.12.17.628804 (†Equal contribution)

B.S., Chonnam National University 2024

Email: mmsscc5931@gmail.com

Current Position: Associate Professor, Yangtze University, China (2025- )

Ph.D Chonnam National University, Korea 2025

M.S. Yangtze University, Jingzhou, China 2020

B.S. Yangtze University, Jingzhou, China 2017

Email: liuhaifeng141@gmail.com

1. Liu H†, Htun AA †, Aung SLL, Sang H, Deng JX*, Tao Y. 2025. Fungal species associated with tuber rot of Foshou yam (Dioscorea esculenta) in China. Journal of Fungi 11:380

2. Liu H†, Liu FY†, Ke HY, Shi QX, Deng JX*, Sang H*. 2025. Three novel species of Alternaria (Pleosporales, Pleosporaceae) from cereal crops (Poaceae) in China. MycoKeys 116: 167-183

3. Liu H, Choi H, Paul NP, Ariyawansa HA, Sang H*. 2025. Discovering fungal communities in roots of Zoysia japonica and characterizing novel species and their antifungal activities. IMA Fungus 16: e138479

4. Choi S†, Son D†, Jang HR, Park S, Liu H, Lee HG, Sang H*. Ipflufenoquin, Novel Dihydroorotate Dehydrogenase Inhibitor: Efficacy Against Strawberry Diseases and Docking Simulations via AlphaFold2 and Autodock Vina. Preprint at https://doi.org/10.1101/2024.12.17.628804 (†Equal contribution)

5. Liu H, Lee G, Sang H*.  2024. Exploring SDHI fungicide resistance in Botrytis cinerea through genetic transformation system and AlphaFold model-based molecular docking. Pest Management Science 80:5954-5964

6. Kim G, Son D, Choi S, Liu H, Nam Y, Sang H*. 2023. Monitoring of benzimidazole resistance in Botrytis cinerea Isolates from strawberry in Korea and development of detection method for benzimidazole resistance. The Plant Pathology Journal 39(6): 614-624

7. Liu H, Park S, Sang H*. 2023. Identification and fungicide control of Bipolaris sorokiniana causing leaf spot and blight on common hop (Humulus lupulus) in Korea. Plant Disease 107: 2939-2943 

8. Lee G†, Choi H†, Liu H†, Han YH†, Paul NC, Han GH, Kim H, Kim PI, Seo SI, Song J, Sang H*. 2023. Biocontrol of the causal brown patch pathogen Rhizoctonia solani by Bacillus velezensis GH1-13 and development of a bacterial strain specific detection method. Frontiers in Plant Science 13:1091030

9. Paul NC, Park S, Liu H, Lee JG, Han GH, Kim H, Sang H*. 2021. Fungi Associated with Postharvest Diseases of Sweet Potato Storage Roots and In Vitro Antagonistic Assay of Trichoderma harzianum against the Diseases. Journal of Fungi 7: 927

10. Paul NC†, Park SW†, Liu H, Choi S, Ma J, MacCready JS, Chilvers MI, Sang H*. 2021. Plant and Fungal Genome Editing to Enhance Plant Disease Resistance Using the CRISPR/Cas9 System. Frontiers in Plant Science 12:700925 (†Equal contribution)

Current Position: Postdoctoral Researcher, Michigan State University (2025- )

Ph.D, Chonnam National University 2025

B.S., Chonnam National University 2020

Email: sungu425@gmail.com

1. Choi S†, Son D†, Jang HR, Park S, Liu H, Lee HG, Sang H*. Ipflufenoquin, Novel Dihydroorotate Dehydrogenase Inhibitor: Efficacy Against Strawberry Diseases and Docking Simulations via AlphaFold2 and Autodock Vina. Preprint at https://doi.org/10.1101/2024.12.17.628804 (†Equal contribution)

2. Park SW†, An W†, Choi S, Choi H, Choi JH, Park S, Jang HR, Back CG, Hong SW, Sang H*.  2024. Evaluation of fungicidal control of garlic leaf blight using mobile spraying equipment.Korean J. Pestic. Sci. 28(3): 291-297 (†Equal contribution)

3. Kim G, Son D, Choi S, Liu H, Nam Y, Sang H*. 2023. Monitoring of benzimidazole resistance in Botrytis cinerea Isolates from strawberry in Korea and development of detection method for benzimidazole resistance. The Plant Pathology Journal 39(6): 614-624

4. Choi S†, Son D†, Chilvers MI, Kim HJ, Sang H*. 2023. First report of Diaporthe eres causing leaf spot disease on Machilus thunbergii in Korea. Plant Disease 107: 1225

5. Ma H, Choi S, Sang H*, Kim H-J*. 2023. First report of root rot of Dendropanax trifidus caused by Fusarium oxysporum in Korea. The Korean Journal of Mycology 51, 51-56

6. Sang H*, Chang HX, Choi S, Son D, Lee G, Chilvers MI*.  2022. Genome-wide transcriptional response of the causal soybean sudden death syndrome pathogen Fusarium virguliforme to a succinate dehydrogenase inhibitor fluopyram. Pest Management Science 78 (2): 530-540

7. Paul NC†, Park SW†, Liu H, Choi S, Ma J, MacCready JS, Chilvers MI, Sang H*. 2021. Plant and Fungal Genome Editing to Enhance Plant Disease Resistance Using the CRISPR/Cas9 System. Frontiers in Plant Science 12:700925 (†Equal contribution)

8. Choi S†, Paul NC†, Lee KH, Kim HJ*, Sang H*. 2021. Morphology, Molecular Phylogeny, and Pathogenicity of Neofusicoccum parvum, Associated with Leaf Spot Disease of a New Host, the Japanese Bay Tree (Machilus thunbergii). Forests 12: 440 (†Equal contribution)

Current Position: Researcher, Center for Industrialization of Agricultural and Livestock Microorganisms (2025- )

M.S. Chonnam National University 2025

B.S., Chonnam National University 2023

Email: awm2454@gmail.com

1. Park SW†, An W†, Choi S, Choi H, Choi JH, Park S, Jang HR, Back CG, Hong SW, Sang H*.  2024. Evaluation of fungicidal control of garlic leaf blight using mobile spraying equipment.Korean J. Pestic. Sci. 28(3): 291-297 (†Equal contribution)