Anne Chao ( 趙蓮菊 )
Address:
Institute of Statistics and Data Science, National Tsing Hua University, Hsin-Chu, Taiwan 30043
Tel:
886-3-5715131 ext 33161
E-mail:
Website:
Google Scholar https://scholar.google.com/citations?user=P-rr9foAAAAJ&hl=en
Research Gate https://www.researchgate.net/profile/Anne_Chao
Github https://github.com/AnneChao
Education:
1973 B. S. (Mathematics), National Tsing Hua University, Taiwan
1977 Ph.D. (Statistics), University of Wisconsin-Madison, USA
Experience:
Visiting Assistant Professor, Department of Statistics, University of Michigan, USA (1977–1978);
Associate Professor (1978–1982), Professor (1982–1988), and Director (1986–1988), Institute of Applied Mathematics, National Tsing Hua University;
Visiting Scientist, Department of Biostatistics, Harvard University, USA (1985–1986);
Professor (1988–2022) and Director (1991–1992, 1996–1998), Institute of Statistics, National Tsing Hua University;
Distinguished Chair Professor (2006–2022), and College of Science Emerita Chair Professor (2022–), National Tsing Hua University
Mercator Fellow, German Research Foundation (DFG) BETA-FOR Research Unit, (2022–)
Adviser, Think Nature Company, Japan (2024–)
Editorship:
1994-2013 Associate Editor, Biometrics
1998-2011 Associate Editor, Journal of Agricultural, Biological, and Environmental Statistics
1991-96, 1999-2005 Associate Editor, Statistica Sinica
2001-2009 Associate Editor, Australian & New Zealand Journal of Statistics
2015- 2019 Associate Editor, Methods in Ecology and Evolution
Referees for various journals in biology, ecology, statistics, epidemiology and related sciences.
Honors and Awards:
1987 Elected Member of International Statistical Institute
1990-95 Taiwan National Science Council Award for Outstanding Research (傑出研究獎三次)
1995–01 Taiwan National Science Council Special Research Award (國科會特約研究獎兩次)
1996 Outstanding Teaching Award, National Tsing Hua University (傑出教學獎)
1997 Fellow, Institute of Mathematical Statistics
2001 ISI Citation Classic Award for two highly-cited papers
(Chao, 1984; Chao and Lee, 1992)
2003 Academic Award, Ministry of Education, Taiwan (教育部學術獎)
2005-2008 Taiwan National Chair Professor (國家講座)
2006-2022 Tsing Hua Distinguished Chair Professor (清華特聘講座)
2013 Lifetime Achievement Award, Statistical Society, Taiwan (統計學社終身成就獎 )
2017 Sun Yat-sen Chair in Ecology and Evolution, China (中國孫中山生態及演化講座 )
2022 Distinguished Fellow, International Biogeography Society
2022- College of Science Emerita Chair Professor, National Tsing Hua University
Theses Supervised:
24 Ph.D. Theses and > 100 Master Theses, Institute of Statistics, National Tsing Hua University
Major Research Areas:
Ecological statistics and biodiversity data analysis/estimation
Species-richness estimation and population size estimation
Biodiversity indices (species, phylogenetic, and functional diversities)
and their decomposition and estimation
Similarity and differentiation among biological communities
Statistical inferences and standardization for biological sampling data
Capture-recapture models and their applications
Software development for analyzing biodiversity analysis
Publications and Citations in Google Scholar:
Google Scholar citations (GS) and Web of Science citations (WoS) Accessed on Sept. 25, 2025
A complete list of 168 publications (with pdf and/or journal links) is given on Anne Chao’s website.
The total number of citations in Google Scholar is > 55,000 citations, with citations continuing to rise annually (exceeding 5,400 in 2024 alone)
Chao (1984) has been cited more than 6,400 times for a single paper; each of the two iNEXT papers (Chao et al. 2014; Hsieh, Ma and Chao 2016) has been cited over 4,500 times.
A total of ten papers have each been cited more than 1,000 times.
The eight most-cited papers are listed below (each with Google Scholar citations exceeding 1,700 and Web of Science citations exceeding 1,000)
1. Chao, A. (1984). Nonparametric estimation of the number of classes in a population. Scandinavian Journal of Statistics, 11, 265-270. [Citation 6428 (GS); 3958 (WoS)]
2. Hsieh, T. C., Ma, K. H. and Chao, A. (2016). iNEXT: An R package for rarefaction and extrapolation of species diversity (Hill numbers). Methods in Ecology and Evolution, 7, 1451-1456. [Citation 4743 (GS); 3287 (WoS)]
3. Chao, A., Gotelli, N. G., Hsieh, T. C., Sander, E. L., Ma, K. H., Colwell, R. K. and Ellison, A. M. (2014). Rarefaction and extrapolation with Hill numbers: a framework for sampling and estimation in species biodiversity studies. Ecological Monographs 84, 45-67. [Citation 4639 (GS); 3038 (WoS)]
4. Chao, A. (1987). Estimating the population size for capture-recapture data with unequal catchability. Biometrics, 43, 783-791. [Citation 3110 (GS); 1845 (WoS)]
5. Chao, A. and Jost. L. (2012) Coverage-based rarefaction: standardizing samples by completeness rather than by size. Ecology, 93, 2533-2547. (Cover article) [Citation 2479 (GS); 1600 (WoS)]
6. Colwell, R. K., Chao, A., Gotelli, N. J., Lin, S. Y., Mao, C. X., Chazdon, R. L., and Longino J. T. (2012). Models and estimators linking individual-based and sample-based rarefaction, extrapolation, and comparison of assemblages. Journal of Plant Ecology, 5, 3-21. [Citation 2383 (GS); 1520 (WoS)]
7. Chao, A., Chazdon, R. L., Colwell, R. K. and Shen, T.-J. (2005). A new statistical approach for assessing similarity of species composition with incidence and abundance data. Ecology Letters, 8, 148-159. [Citation 2259 (GS); 1547 (WoS)]
8. Chao, A. and Lee, S-M. (1992). Estimating the number of classes via sample coverage. Journal of American Statistical Association, 87, 210-217. [Citation 1771 (GS) 1038 (WoS)]
Plenary/Keynote Speeches Since 2012
2012 World Congress on Natural Resource Modeling (WCNRM), Brisbane, Australia;
2013 Eco-Stats Symposium, The University of New South Wales, Sydney, Australia;
2013 Institute of Mathematical Statistics, Asia Pacific Rim Meeting (IMS-APRM 2014), Taiwan;
2014 Graybill/ENVR Conference on Modern Statistical Methods for Ecology, Colorado State University, USA;
2016 British Ecological Society Annual Meeting, Liverpool, UK;
2018 Annual Conference of Ecological Society of Australia, Brisbane, Australia;
2019 International Congress of Conservation Biology (ICCB), Kuala Lumpur, Malaysia;
2022 Annual Taiwan Statistical Conference 統計學術研討會, Tamkang University, Taiwan;
2024 35th Congress of Animal Behavior and Ecology (動物行為暨生態研討會), Taiwan;
2025 54th Annual Meeting of the Ecological Society of Germany, Austria and Switzerland, Wurzburg, Germany
Numerous invited presentations at national and international conferences and scientific meetings
Major Software and Packages:
All software (with users’ guides) can be downloaded from Anne Chao’s website and Github; some (in bold below) are available from CRAN
iNEXT (iNterpolation/EXTrapolation): for taxonomic diversity;
iNEXT.4steps (iNterpolation/EXTrapolation — 4-step analysis including sample completeness, size- and coverage-based iNEXT, and evenness)
iNEXT.3D (iNterpolation/EXTrapolation for 3 dimensions of biodiversity, including taxonomic, phylogenetic, and functional diversity)
iNEXT.beta3D (iNterpolation/EXTrapolation with beta diversity for 3 dimensions of biodiversity)
iNEXT.link (iNterpolation/EXTrapolation for network/link data)
iNEXT.meta (iNterpolation/EXTrapolation for meta analysis)
CARE (CApture-REcapture): for capture-recapture and epidemiological data
SPADE (Species Prediction And Diversity Estimation): biodiversity data analyses and statistical inferences for species-richness, shared species-richness, Hill numbers, and various (dis)similarity indices
MF.beta4: statistical analysis based on ecosystem multi-functionality data (for German beta4 project)
iNEXT received GitHub’s recognition as an Arctic Code Vault Contributor (2020), with source code archived in the Arctic World Archive to preserve global open-source knowledge for future generations.