Publications & Communications

Articles (peer reviewed)

Submitted

• Veuillen, L., Simioni, G., De Cáceres, M., Badel, E., Carrière, S. D., Cochard, H., Courbet, F., Doussan, C., Druel, A., Ladier, J., Prévosto, B., Raynal, K. and Martin-StPaul, N.: Thinning enhances hydraulic safety but not growth resistance to drought in Atlas cedar on the long term. Agricultural and Forest Meteorology, AGRFORMET-D-25-02073, submitted, 2025.

• Blondeel, H., Serrano-León, H., Bonal, D., Martin-StPaul, N., Grossiord, C., Guillemot, J., Schnabel, F., Boone, M., Decarsin, R., Druel, A., Godbold, D., Glenz, P., Gong, J., Hajek, P., Jactel, H., Koricheva, J., Meredieu, C., Mereu, S., Messier, C., Muys, B., Ponette, Q., Rewald, B., Sandén, H., Scherer-Lorenzen, M., Skiadaresis, G., Steurer, J., Van den Bulcke, J., Verheyen, K, Werner, R., Bauhus, J. and Baeten L.: Neighborhood diversity in hydraulic traits reduces physiological drought stress in trees. New Phytologist, submitted, 2025.

• Osy, C., Opfergelt, S., Druel, A. and Massonnet, F.: Trends, variability and drivers of the date of large-scale permafrost active layer freeze-back in the ERA5-Land dataset in Siberia, Russia. The Cryosphere, submitted, 2025. (Preprint available here)

2025

• Copie, A., Scotti-Saintagne, C., Lefèvre, F., Cochard, H., Delzon, S., Druel, A., Dumas, P.-J., Gounelle, D., Herbette, S., Jean, F., Mariotte, N., Mencuccini, M., Scotti, I. and Martin-StPaul, N.: Beyond Proxies: Towards ecophysiological indicators of drought resistance for forest management. Tree Physiology, Volume 45, Issue 9, tpaf090, https://doi.org/10.1093/treephys/tpaf090, 2025. (Available here)

• Serrano-León, H., Blondeel, H., Glenz, P., Steurer, J., Schnabel, F., Baeten, L., Guillemot, J., Martin-StPaul, N., Skiadaresis, G., Scherer-Lorenzen, M., Bonal, D., Boone, M., Decarsin, R., Druel, A., Godbold, D. L., Gong, J., Hajek, P., Jactel, H., Koricheva, J., Mereu, S., Ponette, Q., Rewald, B., Sandén, H., Bulcke, J. V., Verheyen, K., Werner, R. and Bauhus, J.: Multi-year drought strengthens positive and negative functional diversity effects on tree growth response. Global Change Biology, 31:e70394, https://doi.org/10.1111/gcb.70394, 2025. (Available here)

• Verfaillie, D., Chapron, E., Jomelli, V., Druel, A., Lacou, A., Nougarède, S., Guillou, V., Schimmelpfennig, I., Charton, J. and ASTER Team: Holocene glacier evolution in the Pyrenees based on 36Cl cosmic-ray exposure dating (Troumouse Cirque, Pyrenees National Park). Mediterranean Geoscience Reviews, https://doi.org/10.1007/s42990-025-00175-5, 2025. (Available here)

• Rouet, C., Davi, H., Druel, A., Fady, B., and Morin, X.: PDG-Arena: An ecophysiological model for characterizing tree-tree interactions in heterogeneous and mixed stands. Annals of Forest Science, 82, 8, https://doi.org/10.1186/s13595-025-01277-8, 2025. (Available here)

• Druel, A., Ruffault, J., Davi, H., Chanzy, A., Marloie, O., De Cáceres, M., Mouillot, F., François, C., Soudani, K., and Martin-StPaul, N.: Enhancing environmental models with a new downscaling method for global radiation in complex terrain. Biogeoscience, 22, 1-18, https://doi.org/10.5194/bg-22-1-2025, 2025. (Available here)

2024

• Decarsin, R., Guillemot, J., le Maire, G., Blondeel, H., Meredieu, C., Achard, E., Bonal, D., Cochard, H., Corso, D., Delzon, S., Doucet, Z., Druel, A., Grossiord, C., Torres-Ruiz, J.M., Bauhus, J.,  Godbold, D., Hajek, P., Jactel, H., Jensen, J., Mereu, S., Ponette, Q., Rewald, B., Ruffault, R., Sandén, H., Scherer-Lorenzen, M., Serrano-León, H., Simioni, G., Verheyen, K., Werner, R., and Martin-StPaul, N.: Tree drought-mortality risk depends more on intrinsic species resistance than on stand species diversity. Global Change Biology, 30-9, https://doi.org/10.1111/gcb.17503, 2024.  (Here)

• Blondeel, H., Guillemot, J., Martin-StPaul, N., Druel, A., Bilodeau-Gauthier, S., Bauhus, J., Grossiord, C., Hector, A., Jactel, H., Jensen, J., Messier, C., Muys, B., Serrano-León, H., Auge, H., Barsoum, N., Birhane, E., Bruelheide, H., Cavender-Bares, J., Chu, C., Cumming, J., Damtew, A., Eisenhauer, N., Ferlian, O, Fiedler, S., Ganade, G., Godbold, D., Gravel, D., Hall, J., Hölscher, D., Hulvey, K., Koricheva, J., Kreft, H., Lapadat, C., Liang, J., Liu, X., Meredieu, C., Mereu, S., Montgomery, R., Morillas, L., Nock, C., Paquette, A., Parker, J., Parker, W., Paterno, G., Perring, M., Ponette, Q., Potvin, C., Reich, P., Rentch, J., Rewald, B., Sandén, H., Sinacore, K., Standish, R., Stefanski, A., Tobin, P., Breugel, M., Vergara Fagundes, M., Weih, M., Williams, L., Zhou, M., Scherer-Lorenzen, M., Verheyen, K., and Baeten, L.: Tree diversity reduces variability in sapling survival under drought. Journal of Ecology, 112-5, 1164-1180, https://doi.org/10.1111/1365-2745.14294, 2024. (Available here)

2022

• Druel, A., Munier, S., Mucia, A., Albergel, C., and Calvet, J.-C.: Implementation of a new crop phenology and irrigation scheme in the ISBA land surface model using SURFEX_v8.1, Geosci. Model Dev., https://doi.org/10.5194/gmd-15-8453-2022, 2022. (Available here)

• Villani, M., Mauclet, E., Agnan, Y., Druel, A., Jasinki, B., Taylor, M., Schuur, E., and Opfergelt, S.: Mineral element recycling in topsoil following permafrost degradation and a vegetation shift in sub-Arctic tundra, Geoderma, https://doi.org/10.1016/j.geoderma.2022.115915, 2022. (Available here)

2020

• Masson, V., Bocher, E., Bucher, B., Chitu, Z., Christophe, S., Fortelius, C., Hamdif, R., Lemonsu, A., Perrels, A., Schaeybroeck, B.V.,  BenWichers, Schreur, B.W., Velea, L., Beddarn Y., Calvet, J.C., Delcloof, A., Druel, A., Duchêne, F., Dumas, G., Dumitrescu, A., Gautier,  J., Goret, M., Horttanainen, M., Kouznetsov, R., Le Bris, A., Lecorre, S., Le Roy, B., Palamarchuk, Y., Petit, G., Ruuhela, R., Saranko, O., Sofiev, M., Siljamo, P., Van de Vyver, H., van Velthoven, P., and Votsis., A.: The Urban Climate Services URCLIM project, Climate Services, 20, 100194, https://doi.org/10.1016/j.cliser.2020.100194, 2020. (Available here)

• Leibovici, D., Quegan, A., Comyn-Platt, E., Hayman, G., Val Martin, M., Guimberteau, M., Druel, A., Zhu, D., and Ciais, P.: Spatio-Temporal Variations and Uncertainty in Land Surface Modelling for High Latitudes: Univariate Response Analysis, Biogeosciences, 17, 1821–1844 , https://doi.org/10.5194/bg-17-1821-2020, 2020. (Available here)

2019

• Druel, A., Ciais, P., Krinner, G., and Peylin, P.: Modeling the vegetation dynamics of northern shrubs and mosses in the ORCHIDEE land surface model, J. of Adv. in Model. Earth Syst., 11, 2020–2035, https://doi.org/10.1029/2018MS001531, 2019. (Available here)

2017

• Druel, A., Peylin, P., Krinner, G., Ciais, P., Viovy, N., Peregon, A., Bastrikov, V., Kosykh, N., and Mironycheva-Tokareva, N.: Towards a more detailed representation of high-latitude vegetation in the global land surface model ORCHIDEE (ORC-HL-VEGv1.0), Geosci. Model Dev., 10, 4693-4722, https://doi.org/10.5194/gmd-10-4693-2017, 2017. (Available here)

2015

• Zhu, D., Peng, S. S., Ciais, P., Viovy, N., Druel, A., Kageyama, M., Krinner, G., Peylin, P., Ottlé, C., Piao, S. L., Poulter, B., Schepaschenko, D., and Shvidenko, A.: Improving the dynamics of Northern Hemisphere high-latitude vegetation in the ORCHIDEE ecosystem model, Geosci. Model Dev., 8, 2263-2283, https://doi.org/10.5194/gmd-8-2263-2015, 2015. (Available here)

2014

• Haury, J., Druel, A., Cabral, T., Paulet, Y., Bozec, M , and Coudreuse, J.: Which adaptations of some invasive Ludwigia spp. (Rosidae, Onagraceae) populations occur in contrasting hydrological conditions in Western France?, Hydrobiologia, 737:45, https://doi.org/10.1007/s10750-014-1815-7, 2014. (Here)

International Talks (refereed conference proceedings)

• Réunion des Sciences de la Terre (29e édition), Montpellier (France), 27-31 October 2025. Influence de la distribution racinaire sur les stress hydrique et l’utilisation de l’eau profonde par les arbres. Mondon, L., Martin-St Paul, N., Rickert, G., Druel, A., Özgen-Xian, I., Cochard, H., Pessel, M., Jougnot, D. and Carrière S. (Abstract)

• Xylem International Meeting (XIM6), Seville (Spain), 18-20 March 2025. From Species Identity to Stand Structure: Understanding Tree Drought Responses in Diverse Forests. Decarsin, R., Guillemot, J., le Maire, G., Blondeel, H., Meredieu, C., Achard, E., Bonal, D., Cochard, H., Corso, D., Delzon, S., Doucet, Z.,  Druel, A., Grossiord, C., Torres‐Ruiz, J. M., Bauhus, J., Godbold, D. L., Hajek, P., Jactel, H., Jensen, J., Mereu, S., Ponette, Boris Rewald, Q., Ruffault, J., Sandén, H., Scherer‐Lorenzen, M., Serrano‐León, H., Simioni, G., Verheyen, K., Werner, R. and Martin‐StPaul, N.

• Climat et impacts 2024, Paris-Saclay (France), 6-8 November 2024. Accessing the total soil available water, an essential step in assessing the vulnerability of our forests to climate change. Druel, A., Ruffault, J., Cochard, H., De Cáceres, M., Delpierre, N. and Martin-St Paul, N. (Virtual presentation)

• ICOS Science Conference 2024, Versailles (France), 10–12 September 2024. Quantifying the Total Water Available to trees through water fluxes measurements at 14 European forest sites. Delpierre, N., Joetzjer, E.,  Lafont, S., Berveiller, D., Carrière, S., Druel, A., François, C., Martin-StPaul, N., Simioni, G. and Limousin, J.-M. (Abstract)

• EGU General Assembly 2023, Vienna (Austria), 23–28 April 2023. Solicited and highlighted presentation: European forest vulnerability to hydraulic failure: an ecohydrological approach. Druel, A., Martin, N., Cochard, H., DeCaceres, M., Delzon, S., Mencuccini, M., Torres-Ruiz, J., and Ruffault, J. (Abstract) 

• INTECOLL 2022, Geneva (Switzerland), 28 August -2 September 2022. Downscaling radiation to improve high-resolution tree simulation. Druel, A., Martin, N., Davi, H., Ruffault, J., Mouillot, F., and Chanzy, A.

• EGU General Assembly 2022, Vienna (Austria), 23–27 May 2022. Does vegetation shift in Arctic tundra upon permafrost degradation influence mineral element recycling in the topsoil? Villani, M., Mauclet, E., Agnan, Y., Druel, A., Jasinki, B., Taylor, M., Schuur, E., and Opfergelt, S. (Abstract and Presentation)

• 1st GLASS/GHP Workshop on Irrigation, Online, 4-5 November 2021. Implementation of a new irrigation scheme in the ISBA Land Surface Model. Druel, A., Albergel, C., Munier, S., and Calvet, J.-C. (Presentation)

• EMS Annual Meeting 2019, Copenhagen (Denmark), 9-13 September 2019. Implementation of a new irrigation scheme in the ISBA Land Surface Model. Druel, A., Albergel, C., Munier, S., and Calvet, J.-C. (Abstract and presentation)

• SURFEX Users Workshop 2019, Toulouse (France), 18-19 March 2019. Irrigation with ECOCLIMAP Second Generation in SURFEX. Druel, A., Albergel, C., Munier, S., and Calvet, J.-C. (Presentation)

• AGU Fall Meeting 2016, San Francisco (USA), 12-16 December 2016. Consequences of introducing bryophytes and Arctic-shrubs in a land surface model with dynamical vegetation. Druel, A., Krinner, G., Peylin, P., Ciais, P., and Viovy, N. (Abstract)

• EGU General Assembly 2016, Vienna (Austria), 17–22 April 2016. Improvement of boreal vegetation modelling and climate interactions through the introduction of new bryophyte and artic-shrub plant functional types in a land surface model. Druel, A., Krinner, G., Peylin, P., Ciais, P., Viovy, N., Peregon, A., Kosykh, N., and Mironycheva-Tokareva, N. (Abstract and presentation)

• International Symposium on Aquatic Plants - Plants in hydrosystems, Poznan (Poland), 27-31 August 2012. Comparing aquatic and terrestrial forms of Ludwigia in western France: biomass and morphology. Haury, J., Druel, A., Cabral, T., Paulet, Y., Bozec, M., Coudreuse, J.  (Here)

International Posters (refereed conference proceedings)

• Xylem International Meeting (XIM6), Seville (Spain), 18-20 March 2025. Accessing the Total soil Available Water using plant hydraulic models and ecohydrological theory : a step toward accurate prediction of forest vulnerability to climate change. Druel, A., Ruffault, J., Cochard, H., De Cáceres, M., Delpierre, N., Joetzer, E., Mencuccini, M., Limousin, J.-M. and Martin-StPaul, N. (Poster)

• Xylem International Meeting (XIM6), Seville (Spain), 18-20 March 2025. Towards SurEau-Ecos v3.0: Developments and call for feedback to enhance plant hydraulic modelling. Rickert, G., Özgen-Xian, I., Druel, A., Ruffault, J., Pimont, F., Cochard, H., De Cáceres, M., Calderon-Morales, E. and Martin-StPaul, N. (Poster)

PhD thesis

Modelling of the boreal vegetation and its dynamics in the context of polar amplification of past and future climate change. University Grenoble Alpes, doctoral school Terre Univers Environnement. Thesis prepared at the  Institute of Environmental Geoscience (IGE, ex LGGE) and supervised by Philippe Peylin and Gerhard Krinner. Defense on January 23rd, 2017. (Thesis manuscript)

Research Reports

• Ruffault, J., Martin-StPaul, N., Druel, A., and Cochard, H. Présentation et applications de SurEau : un modèle pour prédire les effets des sécheresses extrêmes sur les arbres forestiers.  Forêt méditerranéenne, t. XLV, n°1, p69-74, 2024.

• Haury, J., Chotard, S., Astier, D., Noël, F., Cabral, T., Druel, A., Poirier, V., Bozec, M., Coudreuse, J., Saur, N., Linard, D., and Sanz-Casas, N.: Ecologie et gestion des espèces invasives dans les marais Nord-Loire 2010-2013. ProdINRA 295439 , 47p, hal-02798361, 2014.

• Druel, A., Bozec, M., Paulet, Y., Cabral, T., Moreau, C., and Haury, J.: Etude des formes terrestres de Jussie à grandes fleurs (Ludwigia grandiflora) dans le marais de Mazerolles en Loire Atlantique, Mise en place d'une expérience de lutte et de restauration prairiale. ProdINRA 295454 , 20p, hal-02811138, 2012. (Here)

• Druel, A., Cabral, T., Bozec, M., Paulet, Y., Coudreuse, J., Moreau, C., Haury, E., and Haury, J.: La Jussie. Etudes des caractéristiques biologiques et écologiques : différences d'architecture et de biomasse suivant les sites, les espèces et les régimes hydriques. Bilan des activités sur les biomasses et la phénométrie en Loire-Atlantique en 2011. ProdINRA  295455, 18p, hal-02805708, 2012.

• Haury, J., Cabral, T., Coudreuse, J., Bozec, M., Druel, A., Moreau, C., Paulet, Y., and Saur, N.: Plantes invasives dans les Marais Nord-Loire : bilan et analyse des expérimentations 2011. Agrocampus Ouest Rennes - DREAL Pays de la Loire , 2012.

• Haury, J., Moreau, C., Bozec, M., Druel, A., Paulet, Y., and Cabral, T.: Expérimentation de gestion raisonnée de la Renouée du Japon (Reynoutria japonica). Site du «Moulin à papier», à Saint Calais du Désert (53). Synthèse d'intervention 2011 et rappel des résultats 2010. ProdINRA 295459, 20p, hal-02810737, 2012.

• Druel, A., Haury, J. , Damien, J. P., Bozec, M. , Coudreuse, J., Moreau, C., Cabral, T., Paulet, Y. , Haury, E., Castel, M. and Thabot., S.: La Jussie - Etude de la colonisation des milieux temporairement inondés en Brière, étude des impacts de l'arrachage sur la flore et estimation de son efficacité, évaluation des principales interactions avec l'Ecrevisse de Louisiane et le Ragondin. ProdINRA 295470,  hal-02809996, 2011.