Publications

Publications are listed chronologically below, along with a copy of each text. You can find short summaries of each publication here. The copyright of these articles is with their respective publishers. 

In Press

• Alhajeri BH. Cranial Shape Variation in Kangaroo Mice (Microdipodops): Taxonomic Insights and Ecological Adaptations. Animal Biology. doi: https://doi.org/10.1163/15707563-bja10170. PDF. 

• Alhajeri BH. Ecological Drivers of Cranial Evolution in Bristly Ground Squirrels (Tribe Xerini). Organisms Diversity & Evolution. doi: https://doi.org/10.1007/s13127-025-00683-0. PDF. 

2025

• Alhajeri BH. Taterillus cranial shape variation is linked to allometry, geography, and climate (Gerbillinae, Rodentia). Mammal Research. 70: 577–593. DOI: https://doi.org/10.1007/s13364-025-00817-x. PDF.

• Alhajeri BH. Interspecific cranial variation in coarse-haired pocket mice (Chaetodipus): a geometric morphometric analysis within a phylogenetic framework. Mammalian Biology. 105: 449–466. DOI: https://doi.org/10.1007/s42991-025-00490-2. PDF.

• Fourcade Y, Alhajeri BH. Global phylogenetic and functional structure of rodent assemblages. Ecography. e07534. DOI: https://doi.org/10.1002/ecog.07534. PDF.

• Alhajeri BH. Alaqeely R, Alhaddad H. Geometric Morphometrics of Silky Pocket Mice (Perognathus; Perognathinae; Rodentia) Crania Reveals New Insights Into Their Variation, Evolution, and Taxonomy. Zoological Journal of the Linnean Society. 203: zlae029. DOI: https://doi.org/10.1093/zoolinnean/zlae029. PDF.

• Alhajeri BH, Steppan SJ. Cranial Variation Across Spiny Pocket Mice (Heteromys, Liomys) in New Phylogenetic and Taxonomic Perspectives. Zoology. 168: 126238.  DOI: https://doi.org/10.1016/j.zool.2025.126238. PDF.

• Akbar KM, Alhajeri BH, Alhaddad H. Sequencing KRT71 as a candidate gene for hair shape variation in dromedary camels. Kuwait Journal of Science. 52: 100364. DOI: https://doi.org/10.1016/j.kjs.2025.100364. PDF.

• Alhajeri BH. Cranial Variation in Species and Subspecies of Kangaroo Rats (Dipodomys; Dipodomyinae, Rodentia) According to Geometric Morphometrics. Integrative Zoology. 20: 108–134. DOI: https://doi.org/10.1111/1749-4877.12824. PDF.

2024

• Akbar KM, Alhajeri BH, Alhaddad H. Fiber Characteristics of the Dromedary Camel in the Arabian Peninsula. Small Ruminant Research. 235: 107276. DOI: https://doi.org/10.1016/j.smallrumres.2024.107276. PDF.

• Alhaddad H, Alnughaimish A, Alhajeri D, Alhajeri BH. Towards phenotyping adaptive traits in camels: a study of the response of blood cells to a hypotonic environment. PLOS ONE. 19: e0298336. DOI: https://doi.org/10.1371/journal.pone.0298336. PDF.

• Alhaddad H, Powell BB, Pinto LD, Sutter N, Brooks SA, Alhajeri BH. Geometric morphometrics of face profile across horse breeds and within Arabian horses. Journal of Equine Veterinary Science. 132: 104980. DOI: https://doi.org/10.1016/j.jevs.2023.104980. PDF.

2023

• Alhajeri BH, Hasan Z, Alhaddad H. Cranial differences in three-toed jerboas (Dipodinae, Dipodidae, Rodentia) according to recent taxonomic revisions. Current Zoology. 69: 475–490. DOI: https://doi.org/10.1093/cz/zoac057. PDF.

• Alhajeri BH. Cranial geometric morphometrics of jumping mice (Genera: Eozapus, Napaeozapus, and Zapus; Zapodinae, Rodentia): Implications for subspecies conservation. Journal of Mammalian Evolution. 30: 713–734 . doi: https://doi.org/10.1007/s10914-023-09666-4. PDF.

• Fourcade Y, Alhajeri BH. Environmental correlates of body size influence range size and extinction risk: A global study in rodents. Global Ecology and Biogeography. 32: 206–217. DOI: https://doi.org/10.1111/geb.13622. PDF.

2022

• Dashti Z, Alhaddad H, Alhajeri BH. Skull variation in populations of the Indian gerbil Tatera indica (Gerbillinae, Rodentia) sampled across its broad geographic range. Vertebrate Zoology. 72: 1077–1098. DOI: https://doi.org/10.3897/vz.72.e90474. PDF.

• Dashti Z, Alhaddad H, Alhajeri BH. A geometric morphometric analysis of geographic variation in the Cape Short-eared gerbil, Desmodillus auricularis (Rodentia: Gerbillinae). Mammalia. 86: 615–631 . DOI: https://doi.org/10.1515/mammalia-2022-0019. PDF.

• Steppan SJ, Meyer AA, Barrow LN, Alhajeri BH, Al-Zaidan ASY, Gignac PM, Erickson GM. Phylogenetics And The Evolution Of Terrestriality In Mudskippers (Gobiidae: Oxudercinae). Molecular Phylogenetics and Evolution. 169: 107416. DOI: https://doi.org/10.1016/j.ympev.2022.107416. PDF.

• Alhajeri BH. Desmodilliscus braueri crania compared to Pachyuromys duprasi (Desmodilliscini, Gerbillinae, Rodentia). Mammalia. 86: 77–87 . DOI: https://doi.org/10.1515/mammalia-2021-0036. PDF.

• Alhajeri BH. Geometric differences between the crania of Australian hopping mice (Notomys, Murinae, Rodentia). Australian Mammalogy. 44: 24–38. DOI: https://doi.org/10.1071/AM20067. Postprint.

2021

• Maraqa T, Alhajeri BH, Alhaddad H. FGF5 missense mutation is associated with dromedary hair length variation. Animal Genetics. 52: 848–856.  DOI: https://doi.org/10.1111/age.13132. Postprint.

• Alaqeely R, Alhajeri BH, Almathen F, Alhaddad H. Mitochondrial Sequence Variation, Haplotype Diversity, and Relationships Among Dromedary Camel-Types. Frontiers in Genetics. 12: 723964. DOI: https://doi.org/10.3389/fgene.2021.723964. PDF.

• Alaskar H, Alaqeely R, Alhajeri BH, Alhaddad H. The enigma of camel-types: localities, utilities, names, and breed statuses. Journal of Camelid Science. 14: 22–34. link: http://isocard.net/en/journal/detail/145. PDF.

• Alhajeri BH, Alhaddad H, Alaqeely R, Alaskar H, Dashti Z, Maraqa T. Camel breed morphometrics: current methods and possibilities. Transactions of the Royal Society of South Australia. 145: 90–111. DOI: https://doi.org/10.1080/03721426.2021.1889347. PDF.

• Alhajeri BH. A Geometric Morphometric Analysis of Geographic Mandibular Variation in the Dwarf Gerbil Gerbillus nanus (Gerbillinae, Rodentia). Journal of Mammalian Evolution. 28: 469–480. DOI: https://doi.org/10.1007/s10914-020-09530-9. PDF.

• Alhajeri BH. A morphometric comparison of the cranial shapes of Asian dwarf hamsters (Phodopus, Cricetinae, Rodentia). Zoologischer Anzeiger. 292: 184–196. DOI: https://doi.org/10.1016/j.jcz.2021.04.001. Postprint.

• Alhajeri BH. Cranial variation in allactagine jerboas (Allactaginae, Dipodidae, Rodentia): a geometric morphometric study. Zoological Research. 42: 182–194. DOI: https://doi.org/10.24272/j.issn.2095-8137.2020.302. PDF.

2020

• Alhajeri BH, Fourcade Y, Upham NS, Alhaddad H. A global test of Allen’s rule in rodents. Global Ecology and Biogeography. 29: 2248–2260. DOI: https://doi.org/10.1111/geb.13198. PDF.

• AlAskar H, Alhajeri BH, Almathen F, Alhaddad H. Genetic diversity and population structure of dromedary camel-types. Journal of Heredity. 111: 405–413. . DOI: https://doi.org/10.1093/jhered/esaa016. Preprint.

• Alhajeri BH, Porto LMV,  Maestri R. Habitat productivity is a poor predictor of body size in rodents. Current Zoology. 66: 135–143. DOI: https://doi.org/10.1093/cz/zoz037. PDF.

2019

• Alhajeri BH, Alaqeely R, Alhaddad H. Classifying camel breeds using geometric morphometrics: A case study in Kuwait. Livestock Science. 230: 103824. DOI: https://doi.org/10.1016/j.livsci.2019.103824. Postprint.

• Alhajeri BH, Fourcade Y. High correlation between species-level environmental data estimates extracted from IUCN expert range maps and from GBIF occurrence data. Journal of Biogeography. 46: 1329-1341. DOI: https://doi.org/10.1111/jbi.13619. Preprint.

• Alhajeri BH. Cranial variation in geographically widespread dwarf gerbil Gerbillus nanus (Gerbillinae, Rodentia) populations: Isolation by distance versus adaptation to local environments. Journal of Zoological Systematics and Evolutionary Research. 57: 191–203. DOI: https://doi.org/10.1111/jzs.12247. Postprint.

• Alhaddad H, Alhajeri BH. Cdrom Archive: A Gateway to Study Camel Phenotypes. Frontiers in Genetics. 10: 48. DOI: https://doi.org/10.3389/fgene.2019.00048. PDF.

• Alhaddad H, Maraqa T, Alabdulghafour S, Alaskar H, Alaqeely R, Almathen F, Alhajeri BH. Quality and quantity of dromedary camel DNA sampled from whole-blood, saliva, and tail-hair. PLOS ONE. 14: e0211743. DOI: https://doi.org/10.1371/journal.pone.0211743. PDF.

2018

• Alhaddad H, Alhajeri BH. SamplEase: a simple application for collection and organization of biological specimen data in the field. Ecology and Evolution. 8: 10266–10271 . DOI: https://doi.org/10.1002/ece3.4503. PDF.

• Tabatabaei Yazdi F, Alhajeri BH. Sexual dimorphism, allometry, and interspecific variation in the cranial morphology of seven Meriones species (Gerbillinae, Rodentia). Hystrix, the Italian Journal of Mammalogy. 29: 162–167. DOI: https://doi.org/10.4404/HYSTRIX-00018-2017. PDF.

• Alhajeri BH, Steppan SJ. Disparity and Evolutionary Rate Do Not Explain Diversity Patterns in Muroid Rodents (Rodentia: Muroidea). Evolutionary Biology. 45: 324–344. DOI: https://doi.org/10.1007/s11692-018-9453-z. PDF.

• Alhajeri BH, Steppan SJ. Ecological and Ecomorphological Specialization Are Not Associated with Diversification Rates in Muroid Rodents (Rodentia: Muroidea). Evolutionary Biology. 45: 268–286.  DOI: https://doi.org/10.1007/s11692-018-9449-8. PDF.

• Alhajeri BH, Steppan SJ. A phylogenetic test of adaptation to deserts and aridity in skull and dental morphology across rodents. Journal of Mammalogy. 99: 1197–1216. DOI: https://doi.org/10.1093/jmammal/gyy099. PDF.

• Alhajeri BH, Steppan SJ. Community structure in ecological assemblages of desert rodents. Biological Journal of the Linnean Society. 124: 308–318. DOI: https://doi.org/10.1093/biolinnean/bly068. PDF.

• Alhajeri BH. Craniomandibular Variation in the Taxonomically Problematic Gerbil Genus Gerbillus (Gerbillinae, Rodentia): Assessing the Influence of Climate, Geography, Phylogeny, and Size. Journal of Mammalian Evolution. 25: 261–276. DOI: https://doi.org/10.1007/s10914-016-9377-2. Postprint.

2016

• Martin SA, Alhajeri BH, Steppan SJ. Dietary adaptations in the teeth of murine rodents (Muridae): A test of biomechanical predictions. Biological Journal of the Linnean Society. 119: 66–784. DOI: https://doi.org/10.1111/bij.12822. PDF.

• Alhajeri BH. A phylogenetic test of the relationship between saltation and habitat openness in gerbils (Gerbillinae, Rodentia). Mammal Research. 61: 231–241. DOI: https://doi.org/10.1007/s13364-016-0264-2. PDF.

• Alhajeri BH, Steppan SJ. Association between climate and body size in rodents: A phylogenetic test of Bergmann’s rule. Mammalian Biology. 81: 219–225. DOI: https://doi.org/10.1016/j.mambio.2015.12.001. Postprint.

• Alhajeri BH, Schenk JJ, Steppan SJ. Ecomorphological diversification following continental colonization in muroid rodents (Rodentia: Muroidea). Biological Journal of the Linnean Society. 117: 463–481. DOI: https://doi.org/10.1111/bij.12695. PDF.

2015

• Alhajeri BH, Hunt OJ, Steppan SJ. Molecular systematics of gerbils and deomyines (Rodentia: Gerbillinae, Deomyinae) and a test of desert adaptation in the tympanic bulla. Journal of Zoological Systematics and Evolutionary Research. 53: 312–330. DOI: https://doi.org/10.1111/jzs.12102 . Postprint.