Publications

Peer-reviewed articles are listed below in reverse chronological order, with the oldest article numbered 1. Article titles link to publisher or journal records. Brief publication summaries are available on Kudos, while citation counts and additional indexing can be found on Google Scholar. Please contact the lab if you need a copy of a paper. Copyright remains with the respective publishers. Additional research outputs, including reports, theses, talks, conference presentations, posters, and research resources, can be found on ResearchGate.

2026 (3 articles)

50. Alhajeri BH. (2026). Attendance predicts performance in undergraduate biology: Moderation by instructional and student factors. Kuwait Journal of Science, 53(4), 100650. https://doi.org/10.1016/j.kjs.2026.100650

49. Alhajeri BH. (2026). Accelerated cranial divergence in isolated Ammospermophilus lineages: integrating phylogenetic, allometric, and environmental drivers of morphological evolution. Hystrix, the Italian Journal of Mammalogy, 37(1). https://doi.org/10.4404/hystrix-00800-2025

48. Alhajeri BH. (2026). Allometry and ecology shape cranial variation in phyllotine gerbil mice (Eligmodontia). Zoologischer Anzeiger, 321, 162–178. https://doi.org/10.1016/j.jcz.2026.01.001

2025 (9 articles)

47. Alhajeri BH. (2025). Ecological drivers of cranial evolution in bristly ground squirrels (tribe Xerini). Organisms Diversity & Evolution, 25(3–4), 437–456. https://doi.org/10.1007/s13127-025-00683-0

46. Alhajeri BH. (2025). Cranial shape variation in kangaroo mice (Microdipodops): Taxonomic insights and ecological adaptations. Animal Biology, 75(4), 383–416. https://doi.org/10.1163/15707563-bja10170

45. Alhajeri BH. (2025). Taterillus cranial shape variation is linked to allometry, geography, and climate (Gerbillinae, Rodentia). Mammal Research, 70(4), 577–593. https://doi.org/10.1007/s13364-025-00817-x

44. Alhajeri BH. (2025). Interspecific cranial variation in coarse-haired pocket mice (Chaetodipus): A geometric morphometric analysis within a phylogenetic framework. Mammalian Biology, 105(4), 449–466. https://doi.org/10.1007/s42991-025-00490-2

43. Fourcade Y, Alhajeri BH. (2025). Global phylogenetic and functional structure of rodent assemblages. Ecography, 2025(8), e07534. https://doi.org/10.1002/ecog.07534

42. Alhajeri BH, Alaqeely R, Alhaddad H. (2025). Geometric morphometrics of silky pocket mice (Perognathus: Perognathinae: Rodentia) crania reveals new insights into their variation, evolution, and taxonomy. Zoological Journal of the Linnean Society, 203(1), zlae029. https://doi.org/10.1093/zoolinnean/zlae029

41. Alhajeri BH, Steppan SJ. (2025). Cranial variation across spiny pocket mice (Heteromys, Liomys) in new phylogenetic and taxonomic perspectives. Zoology, 168, 126238. https://doi.org/10.1016/j.zool.2025.126238

40. Akbar KM, Alhajeri BH, Alhaddad H. (2025). Sequencing KRT71 as a candidate gene for hair shape variation in dromedary camels. Kuwait Journal of Science, 52(2), 100364. https://doi.org/10.1016/j.kjs.2025.100364

39. Alhajeri BH. (2025). Cranial variation in species and subspecies of kangaroo rats (Dipodomys, Dipodomyinae, Rodentia) according to geometric morphometrics. Integrative Zoology, 20(1), 108–134. https://doi.org/10.1111/1749-4877.12824

2024 (3 articles)

38. Akbar KM, Alhajeri BH, Alhaddad H. (2024). Fiber characteristics of the dromedary camel in the Arabian Peninsula. Small Ruminant Research, 235, 107276. https://doi.org/10.1016/j.smallrumres.2024.107276

37. Alhaddad H, Alnughaimish A, Alhajeri D, Alhajeri BH. (2024). Towards phenotyping adaptive traits in camels: A study of the influence of hypotonic saline solutions on blood cell area. PLOS ONE, 19(3), e0298336. https://doi.org/10.1371/journal.pone.0298336

36. Alhaddad H, Powell BB, Del Pinto L, Sutter N, Brooks SA, Alhajeri BH. (2024). Geometric morphometrics of face profile across horse breeds and within Arabian horses. Journal of Equine Veterinary Science, 132, 104980. https://doi.org/10.1016/j.jevs.2023.104980

2023 (3 articles)

35. Alhajeri BH, Hasan Z, Alhaddad H. (2023). Cranial differences in three-toed jerboas (Dipodinae, Dipodidae, Rodentia) according to recent taxonomic revisions. Current Zoology, 69(4), 475–490. https://doi.org/10.1093/cz/zoac057

34. Alhajeri BH. (2023). Cranial geometric morphometrics of jumping mice (Genera: Eozapus, Napaeozapus, and Zapus; Zapodinae, Rodentia): Implications for subspecies conservation. Journal of Mammalian Evolution, 30(3), 713–734. https://doi.org/10.1007/s10914-023-09666-4

33. Fourcade Y, Alhajeri BH. (2023). Environmental correlates of body size influence range size and extinction risk: A global study in rodents. Global Ecology and Biogeography, 32(2), 206–217. https://doi.org/10.1111/geb.13622

2022 (5 articles)

32. Dashti Z, Alhaddad H, Alhajeri BH. (2022). Skull variation in populations of the Indian gerbil Tatera indica (Gerbillinae, Rodentia) sampled across its broad geographic range. Vertebrate Zoology, 72, 1077–1098. https://doi.org/10.3897/vz.72.e90474

31. Dashti Z, Alhaddad H, Alhajeri BH. (2022). A geometric morphometric analysis of geographic variation in the Cape short-eared gerbil, Desmodillus auricularis (Rodentia: Gerbillinae). Mammalia, 86(6), 615–631. https://doi.org/10.1515/mammalia-2022-0019

30. Steppan SJ, Meyer AA, Barrow LN, Alhajeri BH, Al-Zaidan ASY, Gignac PM, Erickson GM. (2022). Phylogenetics and the evolution of terrestriality in mudskippers (Gobiidae: Oxudercinae). Molecular Phylogenetics and Evolution, 169, 107416. https://doi.org/10.1016/j.ympev.2022.107416

29. Alhajeri BH. (2022). Desmodilliscus braueri crania compared to Pachyuromys duprasi (Desmodilliscini, Gerbillinae, Rodentia). Mammalia, 86(1), 77–87. https://doi.org/10.1515/mammalia-2021-0036

28. Alhajeri BH. (2022). Geometric differences between the crania of Australian hopping mice (Notomys, Murinae, Rodentia). Australian Mammalogy, 44(1), 24–38. https://doi.org/10.1071/AM20067

2021 (7 articles)

27. Maraqa T, Alhajeri BH, Alhaddad H. (2021). FGF5 missense mutation is associated with dromedary hair length variation. Animal Genetics, 52(6), 848–856. https://doi.org/10.1111/age.13132

26. Alaqeely R, Alhajeri BH, Almathen F, Alhaddad H. (2021). Mitochondrial sequence variation, haplotype diversity, and relationships among dromedary camel-types. Frontiers in Genetics, 12, 723964. https://doi.org/10.3389/fgene.2021.723964

25. Alaskar H, Alaqeely R, Alhajeri BH, Alhaddad H. (2021). The enigma of camel-types: localities, utilities, names, and breed statuses. Journal of Camelid Science, 14(1), 22–34. No DOI assigned.

24. Alhajeri BH, Alhaddad H, Alaqeely R, Alaskar H, Dashti Z, Maraqa T. (2021). Camel breed morphometrics: Current methods and possibilities. Transactions of the Royal Society of South Australia, 145(1), 90–111. https://doi.org/10.1080/03721426.2021.1889347

23. Alhajeri BH. (2021). A geometric morphometric analysis of geographic mandibular variation in the dwarf gerbil Gerbillus nanus (Gerbillinae, Rodentia). Journal of Mammalian Evolution, 28(2), 469–480. https://doi.org/10.1007/s10914-020-09530-9

22. Alhajeri BH. (2021). A morphometric comparison of the cranial shapes of Asian dwarf hamsters (Phodopus, Cricetinae, Rodentia). Zoologischer Anzeiger, 292, 184–196. https://doi.org/10.1016/j.jcz.2021.04.001

21. Alhajeri BH. (2021). Cranial variation in allactagine jerboas (Allactaginae, Dipodidae, Rodentia): A geometric morphometric study. Zoological Research, 42(2), 182–194. https://doi.org/10.24272/j.issn.2095-8137.2020.302

2020 (3 articles)

20. Alhajeri BH, Fourcade Y, Upham NS, Alhaddad H. (2020). A global test of Allen’s rule in rodents. Global Ecology and Biogeography, 29(12), 2248–2260. https://doi.org/10.1111/geb.13198

19. AlAskar H, Alhajeri BH, Almathen F, Alhaddad H. (2020). Genetic diversity and population structure of dromedary camel-types. Journal of Heredity, 111(4), 405–413. https://doi.org/10.1093/jhered/esaa016

18. Alhajeri BH, Porto LMV, Maestri R. (2020). Habitat productivity is a poor predictor of body size in rodents. Current Zoology, 66(2), 135–143. https://doi.org/10.1093/cz/zoz037

2019 (5 articles)

17. Alhajeri BH, Alaqeely R, Alhaddad H. (2019). Classifying camel breeds using geometric morphometrics: A case study in Kuwait. Livestock Science, 230, 103824. https://doi.org/10.1016/j.livsci.2019.103824

16. Alhajeri BH, Fourcade Y. (2019). High correlation between species-level environmental data estimates extracted from IUCN expert range maps and from GBIF occurrence data. Journal of Biogeography, 46(7), 1329–1341. https://doi.org/10.1111/jbi.13619

15. Alhajeri BH. (2019). Cranial variation in geographically widespread dwarf gerbil Gerbillus nanus (Gerbillinae, Rodentia) populations: Isolation by distance versus adaptation to local environments. Journal of Zoological Systematics and Evolutionary Research, 57(1), 191–203. https://doi.org/10.1111/jzs.12247

14. Alhaddad H, Alhajeri BH. (2019). Cdrom Archive: A gateway to study camel phenotypes. Frontiers in Genetics, 10, 48. https://doi.org/10.3389/fgene.2019.00048

13. Alhaddad H, Maraqa T, Alabdulghafour S, Alaskar H, Alaqeely R, Almathen F, Alhajeri BH. (2019). Quality and quantity of dromedary camel DNA sampled from whole-blood, saliva, and tail-hair. PLOS ONE, 14(1), e0211743. https://doi.org/10.1371/journal.pone.0211743

2018 (7 articles)

12. Alhaddad H, Alhajeri BH. (2018). SamplEase: A simple application for collection and organization of biological specimen data in the field. Ecology and Evolution, 8(20), 10266–10271. https://doi.org/10.1002/ece3.4503

11. Tabatabaei Yazdi F, Alhajeri BH. (2018). Sexual dimorphism, allometry, and interspecific variation in the cranial morphology of seven Meriones species (Gerbillinae, Rodentia). Hystrix, the Italian Journal of Mammalogy, 29(2), 162–167. https://doi.org/10.4404/hystrix-00018-2017

10. Alhajeri BH, Steppan SJ. (2018). Disparity and evolutionary rate do not explain diversity patterns in muroid rodents (Rodentia: Muroidea). Evolutionary Biology, 45(3), 324–344. https://doi.org/10.1007/s11692-018-9453-z

9. Alhajeri BH, Steppan SJ. (2018). Ecological and ecomorphological specialization are not associated with diversification rates in muroid rodents (Rodentia: Muroidea). Evolutionary Biology, 45(3), 268–286. https://doi.org/10.1007/s11692-018-9449-8

8. Alhajeri BH, Steppan SJ. (2018). A phylogenetic test of adaptation to deserts and aridity in skull and dental morphology across rodents. Journal of Mammalogy, 99(5), 1197–1216. https://doi.org/10.1093/jmammal/gyy099

7. Alhajeri BH, Steppan SJ. (2018). Community structure in ecological assemblages of desert rodents. Biological Journal of the Linnean Society, 124(3), 308–318. https://doi.org/10.1093/biolinnean/bly068

6. Alhajeri BH. (2018). Craniomandibular variation in the taxonomically problematic gerbil genus Gerbillus (Gerbillinae, Rodentia): Assessing the influence of climate, geography, phylogeny, and size. Journal of Mammalian Evolution, 25(2), 261–276. https://doi.org/10.1007/s10914-016-9377-2

2016 (4 articles)

5. Martin SA, Alhajeri BH, Steppan SJ. (2016). Dietary adaptations in the teeth of murine rodents (Muridae): a test of biomechanical predictions. Biological Journal of the Linnean Society, 119(4), 766–784. https://doi.org/10.1111/bij.12822

4. Alhajeri BH. (2016). A phylogenetic test of the relationship between saltation and habitat openness in gerbils (Gerbillinae, Rodentia). Mammal Research, 61(3), 231–241. https://doi.org/10.1007/s13364-016-0264-2

3. Alhajeri BH, Steppan SJ. (2016). Association between climate and body size in rodents: A phylogenetic test of Bergmann’s rule. Mammalian Biology, 81(2), 219–225. https://doi.org/10.1016/j.mambio.2015.12.001

2. Alhajeri BH, Schenk JJ, Steppan SJ. (2016). Ecomorphological diversification following continental colonization in muroid rodents (Rodentia: Muroidea). Biological Journal of the Linnean Society, 117(3), 463–481. https://doi.org/10.1111/bij.12695

2015 (1 article)

1. Alhajeri BH, Hunt OJ, Steppan SJ. (2015). Molecular systematics of gerbils and deomyines (Rodentia: Gerbillinae, Deomyinae) and a test of desert adaptation in the tympanic bulla. Journal of Zoological Systematics and Evolutionary Research, 53(4), 312–330. https://doi.org/10.1111/jzs.12102