Central Appalachian Mts

I started research in Papeopalynology as an undergraduate with Alfred Traverse at Penn State University - pictured here with Bill Chaloner circa 1980. In 1979 we published the first taxonomic paper on spore-like microfossils that later become known as cryptospores. This included the tetrad genus, Tetrahedraletes and the paired spores, or dyads, known as Dyadospora.

Spores and Cryptospores

Nematothallus

Pleuralata spinosa McCoy, Strother & Briggs