Publications

Peer-reviewed Publications:

Hallbert T.B. and Keeley, E.R. 2024. Allometric shifts in foraging site selection and area increase energy intake for Cutthroat Trout but are constrained by functional limits to prey capture. Transactions of the American Fisheries Society, in press.

Hargrove, J.S., Campbell, M.R., Gunnell, K., High, B., Johnson, C.,  Kennedy, P.A., Loxterman, J.L., Ptacek, M.B., Seiler, S.M. and Keeley E.R. 2024. Interspecific hybridization in a large-river population of Yellowstone Cutthroat Trout: a 20-year programmatic evaluation. North American Journal of Fisheries Management. in press.

Hallbert, T.B. and Keeley, E.R. 2023. Instream complexity increases habitat quality and growth for cutthroat trout in headwater streams. Canadian Journal of Fisheries and Aquatic Sciences 80: 992-1005. 

Dunham, J.B., Taylor, E.B., and Keeley, E.R., 2022, Evolutionary and ecological connectivity in westslope cutthroat trout (Oncorhynchus clarkii lewisi) and mountain whitefish (Prosopium williamsoni) in relation to the potential influences of Boundary Dam, Washington, Idaho, and parts of British Columbia: U.S. Geological Survey Open-File Report 2022–1084, 22 p., https://doi.org/10.3133/ofr20221084.

Owens, H.W. and Keeley, E.R.  2022. Quantifying the foodscape for stream-dwelling cutthroat trout reveals spatial and temporal ranges of resource exploitation and energy intake. Transactions of the American Fisheries Society. 

Keeley, E.R., Loxterman, J.L., Matsaw, S.L., Njorge, Z.M., Seiler, M.B., and S.M. Seiler. 2021. Morphological and genetic concordance of cutthroat trout diversification from western North America.  Canadian Journal of Zoology 99:235-248. 

Carmichael, R.A., Tonina, D., Keeley, E.R., Benjankar, R.M. and K. E. See. 2020. Some like it slow: a bioenergetic evaluation of habitat quality for juvenile Chinook salmon in the Lemhi River, Idaho. Canadian Journal of Fisheries and Aquatic Sciences. 77: 1221-1232.

Keeley, E.R. 2019. Origins, species diversity, and ecological diversification in trout and char. Pages 15-39 In: J. L. Kershner, J. E. Williams, R. E. Gresswell, and J. Lobón-Cerviá, editors. Trout and char of the world. American Fisheries Society, Bethesda, MD.

Eaton, K.R., Loxterman, J.L., and Keeley, E.R. 2018. Connections and containers: using genetic data to understand how watershed evolution and human activities influence cutthroat trout biogeography. Plos One, http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0202043

Smith, G.R., Link, P.K., Thackray, G.D., and E.R. Keeley. 2018. Biogeography of the native fish fauna of Idaho. Pages 45-58 in: J.W. Sigler and D.W. Zaroban, Fishes of Idaho: a natural history survey. Caxton Press, Caldwell, Idaho. 

Link, P.K. and E.R. Keeley. 2018. Geologic constraints on Snake-Columbia River drainage changes and the associated evolutionary diversification of Cutthroat Trout over the last 17 million years. In: P. Trotter, P. Bisson, and L. Schultz; Cutthroat Trout: evolutionary biology and taxonomy. American Fisheries Society, Bethesda MD.

Young, M.K., McKelvey, K.S., Jennings, T., Carter, K., Cronn, R., Keeley, E.R., Loxterman, J.L., Pilgrim, K., M. K. Schwartz. 2018. The phylogeography of westslope cutthroat trout. In: P. Trotter, P. Bisson, and L. Schultz; Cutthroat Trout: evolutionary biology and taxonomy. American Fisheries Society, Bethesda MD.

Campbell, M.R., Keeley, E.R., Kozfkay, C.C., Loxterman, J.L., Evans, P.R., and D.K. Shiozawa. 2018. Describing and preserving the diversity of cutthroat trout in the Yellowstone River, Snake River and Bonneville Basin.  In: P. Trotter, P. Bisson, and L. Schultz; Cutthroat Trout: evolutionary biology and taxonomy. American Fisheries Society, Bethesda MD.

Williams, T.H., Griswold, K.E., Keeley, E.R., Currens, K.P., and G.H. Reeves. 2018. The coastal lineage – diversity of coastal cutthroat trout across their distributional range. In: P. Trotter, P. Bisson, and L. Schultz; Cutthroat Trout: evolutionary biology and taxonomy. American Fisheries Society, Bethesda MD.

Penaluna, B. E., A. Abadía-Cardoso, J. B. Dunham, F. J. García-Dé León, R. E. Gresswell, A. R. Luna, E. B. Taylor, B. B. Shepard, R. Al-Chokhachy, C. C. Muhlfeld, K. R. Bestgen, K. Rogers, M. A. Escalante, E. R. Keeley, G. M. Temple, J. E. Williams, K. R. Matthews, R. Pierce, R. L. Mayden, R. P. Kovach, J. C. Garza, and K. D. Fausch. 2016. Conservation of Native Pacific Trout Diversity in Western North America. Fisheries 41:286–300.

Keeley, E.R., Campbell, S.O., and Kohler, A.E. 2016. Bioenergetic calculations evaluate changes to habitat quality for salmonid fishes in streams treated with salmon carcass analog. Canadian Journal of Fisheries and Aquatic Sciences, 73:819-831. download pdf 

Loxterman, J.L., Keeley, E.R. and Njoroge, Z.M. 2014. Evaluating the influence of stocking history and barriers to movement on the spatial extent of hybridization between westslope cutthroat trout and rainbow trout, Canadian Journal of Fisheries and Aquatic Sciences 71:1050-1058.(download pdf version)

Blakney, J.R. Loxterman, J.L. and Keeley, E.R.,2014. Range-wide comparisons of northern leatherside chub reveal historical and contemporary patterns of genetic variation. Conservation Genetics 15:757-770. (download pdf version)

Leeseberg, C.A. and Keeley, E.R. 2014. Prey size, prey abundance, and temperature as correlates of growth in stream populations of cutthroat trout. Environmental Biology of Fishes 97:599-614.

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Hillyard, R.W. and Keeley E.R. 2012. Temperature-related changes in habitat quality and use by Bonneville cutthroat trout in regulated and un-regulated river segments. Transactions of the American Fisheries Society 141:1649-1663.(download pdf version)

Loxterman, J.L. and Keeley, E.R. 2012. Watershed boundaries and geographic isolation: patterns of diversification in cutthroat trout from western North America. BMC Evolutionary Biology 12:38. http://www.biomedcentral.com/1471-2148/12/38 

Schoby, G.S. and Keeley, E.R. 2011. Home range size and foraging ecology of bull trout and westslope cutthroat trout in the upper Salmon River basin, Idaho. Transactions of the American Fisheries Society 140:636-645.(download pdf version)

Taylor, E.B., Tamkee, P., Keeley, E.R. and Parkinson, E. 2011. Conservation prioritization in widespread species: the use of genetic and morphological data to assess population distinctiveness in rainbow trout (Oncorhynchus mykiss) from British Columbia, Canada. Evolutionary Applications 4:100-115.(download pdf version)

Jenkins, A.R. and Keeley, E.R. 2010. Bioenergetic assessment of habitat quality for stream-dwelling cutthroat trout (Oncorhynchus clarkii bouvieri) with implications for climate change and nutrient supplementation. Canadian Journal of Fisheries and Aquatic Sciences 67: 371-385.(download pdf version)

Seiler, S.M., Gunnell, K., Ptacek, M.B. and Keeley, E.R. 2009. Morphological patterns of hybridization between Yellowstone cutthroat trout (Oncorhynchus clarkii bouvieri) and introduced rainbow trout (O. mykiss) in the South Fork of the Snake River watershed, Idaho and Wyoming. North American Journal of Fisheries Management 29: 1529-1539.(download pdf version)

Lamansky, J.A., Keeley, E.R., Young, M.K., and Meyer, K.A. 2009. The use of hoop nets seeded with mature brook trout to capture conspecifics. North American Journal of Fisheries Management 29:10-17. (download pdf version)

Seiler, S.M. and Keeley, E.R. 2009. Competition between native and introduced salmonid fishes: cutthroat trout have lower growth rate in the presence of cutthroat–rainbow trout hybrids. Canadian Journal of Fisheries and Aquatic Sciences 66: 133-141. (download pdf version)

Seiler, M.B. and Keeley, E.R. 2009. Intraspecific taxonomy and ecology characterize morphological divergence among cutthroat trout (Oncorhynchus clarkii ssp. Richardson) populations. Biological Journal of the Linnean Society 96: 266-291.(download pdf version)

Gunnell, K.,, Tada, M.K., Hawthorne, F.A., Keeley, E.R. and Ptacek, M.B. 2008. Geographic patterns of introgressive hybridization between native Yellowstone cutthroat trout (Oncorhynchus clarkii bouvieri) and introduced rainbow trout (O. mykiss) in the South Fork of the Snake River watershed, Idaho. Conservation Genetics 9: 49-64 (download pdf version)

Seiler, S.M. and Keeley, E.R. 2007. A comparison of aggressive and foraging behaviour between juvenile cutthroat trout, rainbow trout, and F1 hybrids. Animal Behaviour 74: 1805-1812. (download pdf version)

Seiler, S.M. and Keeley, E.R. 2007. Morphological and swimming stamina differences between Yellowstone cutthroat trout (Oncorhynchus clarkii bouvieri), rainbow trout (Oncorhynchus mykiss), and their hybrids. Canadian Journal of Fisheries and Aquatic Sciences 64: 127-135. (download pdf version)

Keeley, E.R., Parkinson, E.A., and Taylor, E.B. 2007. The origins of ecotypic variation of rainbow trout (Oncorhynchus mykiss): an experimental assessment of environmental versus genetically based differences in morphology. Journal of Evolutionary Biology 20: 725-736. (download pdf version)

Keeley, E.R., Parkinson, E.A., and Taylor, E.B. 2005. Ecotypic differentiation of native rainbow trout (Oncorhynchus mykiss) populations from British Columbia. Canadian Journal of Fisheries and Aquatic Sciences 62: 1523-1539. (download pdf version)

Rodnick, K.J., Gamperl, A.K., Lizars, K.R., Bennett, M.T., Rausch, R.N., and Keeley, E.R. 2004. Thermal tolerance and metabolic physiology among redband trout populations in southeastern Oregon. Journal of Fish Biology 64: 310-335. (download pdf version)

Imre, I, Grant, J.W.A., and Keeley, E.R. 2004. The effect of food abundance on territory size and population density of juvenile steelhead trout (Oncorhynchus mykiss). Oecologia 138: 371-378. (download pdf version)

Keeley, E.R. 2003. An experimental analysis of self-thinning in juvenile steelhead trout. Oikos 102: 543-550. (download pdf version)

Gamperl, A.K., Rodnick, K.J., Faust, H.A., Venn, E.C., Bennett, M.T., Crawshaw, L.I., Keeley, E.R., Powell, M.S., Li, H.W., 2002. Metabolism, swimming performance, and tissue biochemistry of high desert redband trout (Oncorhynchus mykiss ssp.): evidence for phenotypic differences in physiological function. Physiological and Biochemical Zoology 75: 413-431. (download pdf version)

Imre, I, Grant, J.W.A., and Keeley, E.R. 2002. The effect of visual isolation on territory size and population density of juvenile rainbow trout (Oncorhynchus mykiss). Canadian Journal of Fisheries and Aquatic Sciences 59: 303-309. (download pdf version)

Keeley, E.R. and Grant, J.W.A. 2001. Prey size of salmonid fishes in streams, lakes, and oceans. Canadian Journal of Fisheries and Aquatic Sciences 58: 1122-1132. (download pdf version)      Erratum

Keeley, E.R. 2001. Demographic responses to food and space competition by juvenile steelhead trout. Ecology 82:1247-1259. (downloadpdf )

Keeley, E.R. 2000. An experimental analysis of territory size in juvenile steelhead trout. Animal Behaviour 59: 477-490. (download pdf version)

Grant, J.W.A., Steingrímsson, S.Ó., Keeley, E.R. and Cunjak, R.A. 1998. Implications of territory size for the measurement and prediction of salmonid abundance in streams. Canadian Journal of Fisheries and Aquatic Sciences 55(supplement 1): 181-190. (download pdf version)

Keeley, E.R. and McPhail, J.D. 1998. Food abundance, intruder pressure, and body size as determinants of territory size in juvenile steelhead trout (Oncorhynchus mykiss). Behaviour 135: 65-82.

Keeley, E.R. and Grant, J.W.A. 1997. Allometry of diet selectivity in juvenile Atlantic salmon (Salmo salar). Canadian Journal of Fisheries and Aquatic Sciences 54: 1894-1902. (download pdf version)

Keeley, E.R. 1996. A perspective on brown trout ecology. Environmental Biology of Fishes 45: 325-327.

Keeley, E.R. and Grant, J.W.A. 1995. Allometric and environmental correlates of territory size in juvenile Atlantic salmon (Salmo salar). Canadian Journal of Fisheries and Aquatic Sciences 52: 186-196. (download pdf version)

Keeley, E.R. and Grant, J.W.A. 1993. Visual information, resource value and sequential assessment in convict cichlid (Cichlasoma nigrofasciatum) contests. Behavioral Ecology 4: 345-349.

Keeley, E.R. and Grant, J.W.A. 1993. Asymmetries in the expected value of food do not predict the outcome of contests between convict cichlids. Animal Behaviour 45: 1035-1037.

Technical Publications:


Keeley ER, Ringelman AM, Breech TM, Wooding AP. 2022. Redband Trout Stream Temperature Data for Big Jacks, Duncan, Little Jacks, Upper Mann and Keithly Creeks 2019-2022. [Dataset]. University of Idaho. https://doi.org/10.7923/77j4-7054


Ringelman, A. M., & Keeley, E. R. (2022). Redband Trout Stream Habitat Data for Big Jacks, Duncan, Little Jacks, Upper Mann and Keithly Creeks Idaho USA [Data set]. University of Idaho. https://doi.org/10.7923/8JB6-0123

Keeley, E.R., Blakney, J.R. and Loxterman, J.L. 2012. Distribution, abundance, and genetic population structure of northern leatherside chub in the Snake River basin of Idaho. Final report to the US Fish and Wildlife Service, agreement no. 143309G735.

Parkinson, E.A., Keeley, E.R., Taylor, E.B., Pollard, S. and Tautz, A.F. 2005. A population database for defining conservation units in steelhead trout. Province of British Columbia, Fisheries Management Report No. 119. 74pp.

Parkinson, E.A., E.B. Taylor, E.B., and Keeley, E.R. 2000. Conserving Genetic Diversity in Rainbow Trout. In: L. M. Darling, editor. Proceedings of a Conference on the Biology and Management of Species and Habitats at Risk, Kamloops, B.C., 15 - 19 Feb.,1999. Volume Two. B.C. Ministry of Environment, Lands and Parks, Victoria, B.C. and University College of the Cariboo, Kamloops, B.C. 520pp. (download pdf version)

Keeley, E.R. 2000. A rationale for protecting Idaho's native fish diversity. Rendezvous 35: 45-59.

O’Brien, D.S., and Keeley, E. R. 1997. Post-spawning movements of steelhead trout (Oncorhynchus mykiss) in the Skeena watershed in 1995 and 1996. B. C. Ministry of Environment, Lands and Parks; Skeena Fisheries Report SK-97: 30 pages.

Koning, C.W., and Keeley, E.R. 1997. Salmonid biostandards for estimating production benefits of fish habitat rehabilitation techniques. In: P.A. Slaney and D. Zaldokas (eds.) Fish Habitat Rehabilitation Procedures. Province of B.C. Ministry of Environment, Lands and Parks, and B. C. Ministry of Forests; Watershed Restoration Technical Circular 9: 21 pages.

Keeley, E.R., Slaney, P.A. and Zaldokas, D. 1996. Estimates of fish production benefits from stream restoration initiatives. Province of B.C. Ministry of Environment, Lands and Parks; Watershed Restoration Project Report 3: 21 pages. (download pdf version)

Keeley, E.R., and Slaney, P.A. 1996. Quantitative measures of rearing and spawning habitat characteristics for stream-dwelling salmonids: implications for habitat restoration. Province of B.C. Ministry of Environment, Lands and Parks; Watershed Restoration Project Report 2: 31 pages. (download pdf version)

Keeley, E.R. and Walters, C.J. 1994. The British Columbia watershed restoration program: summary of the experimental design, monitoring and techniques workshop. Province of B.C. Ministry of Environment, Lands and Parks; Watershed Restoration Management Report 1: 34 pages. (download pdf version)